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Labrisomids are small blennioids (blennies), percomorph marine
fish Fish are aquatic, craniate, gill-bearing animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish as well as various extinct related groups. Approximately 95% ...
belonging to the family Labrisomidae. Found mostly in the tropical
Atlantic The Atlantic Ocean is the second-largest of the world's five oceans, with an area of about . It covers approximately 20% of Earth's surface and about 29% of its water surface area. It is known to separate the "Old World" of Africa, Europe an ...
and
Pacific Ocean The Pacific Ocean is the largest and deepest of Earth's five oceanic divisions. It extends from the Arctic Ocean in the north to the Southern Ocean (or, depending on definition, to Antarctica) in the south, and is bounded by the contine ...
, the family contains about 110 species in 15 genera. Stockier than the average blenny, labrisomids are
elongated Elongation may refer to: * Elongation (astronomy) * Elongation (geometry) * Elongation (plasma physics) * Part of transcription of DNA into RNA of all types, including mRNA, tRNA, rRNA, etc. * Part of translation (biology) of mRNA into proteins ...
nonetheless; their dorsal fin spines outnumber soft rays (which may be absent altogether), and their
pelvic fin Pelvic fins or ventral fins are paired fins located on the ventral surface of fish. The paired pelvic fins are homologous to the hindlimbs of tetrapods. Structure and function Structure In actinopterygians, the pelvic fin consists of two e ...
s are long and slender. Like many other blennies, labrisomids have whisker-like structures called
cirri Giovanni Battista Cirri (1 October 1724 – 11 June 1808) was an Italian cellist and composer in the 18th century. Biography Cirri was born in Forlì in the Emilia-Romagna Region of Italy. He had his first musical training with his brother ...
on their heads and napes. Scales may be
cycloid In geometry, a cycloid is the curve traced by a point on a circle as it rolls along a straight line without slipping. A cycloid is a specific form of trochoid and is an example of a roulette, a curve generated by a curve rolling on another ...
or absent in labrisomids; many species are brightly coloured. The hairy blenny (''Labrisomus nuchipinnis'') is the largest species at 23 cm in length; most are far smaller. Generally staying within shallow coastal regions to depths around 10 m, labrisomids are
benthic The benthic zone is the ecological region at the lowest level of a body of water such as an ocean, lake, or stream, including the sediment surface and some sub-surface layers. The name comes from ancient Greek, βένθος (bénthos), meaning "t ...
fish spending most of their time on or near the bottom. Both sandy and rocky substrates are frequented, sometimes at
reef A reef is a ridge or shoal of rock, coral or similar relatively stable material, lying beneath the surface of a natural body of water. Many reefs result from natural, abiotic component, abiotic processes—deposition (geology), deposition of ...
s or amongst beds of
seagrass Seagrasses are the only flowering plants which grow in marine environments. There are about 60 species of fully marine seagrasses which belong to four families ( Posidoniaceae, Zosteraceae, Hydrocharitaceae and Cymodoceaceae), all in the ...
. Labrisomids are shy fish and will retreat into crevices if threatened.
Crustacean Crustaceans (Crustacea, ) form a large, diverse arthropod taxon which includes such animals as decapods, seed shrimp, branchiopods, fish lice, krill, remipedes, isopods, barnacles, copepods, amphipods and mantis shrimp. The crustacean gro ...
s, gastropods,
brittle star Brittle stars, serpent stars, or ophiuroids (; ; referring to the serpent-like arms of the brittle star) are echinoderms in the class Ophiuroidea, closely related to starfish. They crawl across the sea floor using their flexible arms for locom ...
s, and
sea urchin Sea urchins () are spiny, globular echinoderms in the class Echinoidea. About 950 species of sea urchin live on the seabed of every ocean and inhabit every depth zone from the intertidal seashore down to . The spherical, hard shells (tests) ...
s make up much of the labrisomid diet. Two genera of labrisomid are noted for their
ovoviviparity Ovoviviparity, ovovivipary, ovivipary, or aplacental viviparity is a term used as a "bridging" form of reproduction between egg-laying oviparous and live-bearing viviparous reproduction. Ovoviviparous animals possess embryos that develop insi ...
; '' Xenomedea'' and '' Starksia'' both retain eggs within their
oviduct The oviduct in mammals, is the passageway from an ovary. In human females this is more usually known as the Fallopian tube or uterine tube. The eggs travel along the oviduct. These eggs will either be fertilized by spermatozoa to become a zygote, ...
s, where they develop in safety. However, only ''Starksia'' species possess gonopodia (modified anal fins used as a copulatory organ). Some workers have found that the Labrisomidae is
paraphyletic In taxonomy (general), taxonomy, a group is paraphyletic if it consists of the group's most recent common ancestor, last common ancestor and most of its descendants, excluding a few Monophyly, monophyletic subgroups. The group is said to be pa ...
.


Genera

The following genera are currently included in this family: * '' Alloclinus'' C.L. Hubbs, 1927 * '' Auchenionchus''
Gill A gill () is a respiratory organ that many aquatic organisms use to extract dissolved oxygen from water and to excrete carbon dioxide. The gills of some species, such as hermit crabs, have adapted to allow respiration on land provided they ar ...
, 1860
* '' Brockius''
Clark L. Hubbs Clark Hubbs (March 15, 1921 – February 3, 2008) was an American ichthyologist who was professor of zoology at the University of Texas from 1963 until he accepted emeritus status in 1991. He was a leading figure in ichthyology in Texas, teaching m ...
, 1953
* '' Calliclinus'' Gill, 1860 * '' Cottoclinus'' McCosker, Stephens &
Rosenblatt Rosenblatt is a surname of German and Jewish origin, meaning " rose leaf". People with this surname include: *Albert Rosenblatt (born 1936), New York Court of Appeals judge *Dana Rosenblatt, known as "Dangerous" (born 1972), American boxer *Elie R ...
, 2003
* ''
Cryptotrema ''Cryptotrema'' is a genus of labrisomid blennies native to the eastern Pacific Ocean The Pacific Ocean is the largest and deepest of Earth's five oceanic divisions. It extends from the Arctic Ocean in the north to the Southern Ocean ...
''
Gilbert Gilbert may refer to: People and fictional characters *Gilbert (given name), including a list of people and fictional characters *Gilbert (surname), including a list of people Places Australia * Gilbert River (Queensland) * Gilbert River (South A ...
, 1890
* ''
Dialommus ''Dialommus'' is a genus of labrisomid blennies native to the eastern Pacific Ocean. Species There are currently two recognized species in this genus: * '' Dialommus fuscus'' C. H. Gilbert, 1891 (Galápagos four-eyed blenny) * '' Dialommus ma ...
'' Gilbert, 1891 * ''
Exerpes ''Exerpes asper'', the Sargassum blenny, is a species of labrisomid blenny native to the Gulf of California and the Pacific coast of Baja California. According to Fishbase it is currently the only known member of its genus, however, the Catal ...
''
Jordan Jordan ( ar, الأردن; tr. ' ), officially the Hashemite Kingdom of Jordan,; tr. ' is a country in Western Asia. It is situated at the crossroads of Asia, Africa, and Europe, within the Levant region, on the East Bank of the Jordan Ri ...
& Evermann, 1896
* ''
Gobioclinus ''Gobioclinus'' is a genus of labrisomid blennies from the coasts of the western Atlantic and eastern Pacific Oceans off the Americas. Species The following species are classified within the genus ''Gobioclinus'': *'' Gobioclinus bucciferus'' ...
'' Gill, 1860 * ''
Haptoclinus ''Haptoclinus'' is a genus of labrisomid blennies native to the Caribbean Sea. Species There are currently two recognized species in this genus: * '' Haptoclinus apectolophus'' J. E. Böhlke & C. R. Robins, 1974 (Uncombed blenny) * '' Haptocl ...
'' Böhlke & Robertson, 1974 * '' Labrisomus'' Swainson, 1839 * '' Malacoctenus'' Gill, 1860 * ''
Nemaclinus ''Nemaclinus atelestos'', the Threadfin blenny, is a species of labrisomid blenny known from the western Atlantic Ocean. This species is the only known member of its genus. References Labrisomidae Monotypic fish genera Fish described ...
'' Böhlke &
Springer Springer or springers may refer to: Publishers * Springer Science+Business Media, aka Springer International Publishing, a worldwide publishing group founded in 1842 in Germany formerly known as Springer-Verlag. ** Springer Nature, a multinationa ...
, 1975
* ''
Paraclinus ''Paraclinus'' is a genus of labrisomid blennies native to eastern Pacific Ocean and the western Atlantic Ocean. Species There are currently 23 recognized species in this genus: * ''Paraclinus altivelis'' (William Neale Lockington, Lockington, 1 ...
'' Mocquard, 1888 * '' Starksia'' Jordan & Evermann, 1896 * '' Xenomedea'' Rosenblatt &
Taylor Taylor, Taylors or Taylor's may refer to: People * Taylor (surname) ** List of people with surname Taylor * Taylor (given name), including Tayla and Taylah * Taylor sept, a branch of Scottish clan Cameron * Justice Taylor (disambiguation) ...
, 1971


Timeline

ImageSize = width:1000px height:auto barincrement:15px PlotArea = left:10px bottom:50px top:10px right:10px Period = from:-65.5 till:10 TimeAxis = orientation:horizontal ScaleMajor = unit:year increment:5 start:-65.5 ScaleMinor = unit:year increment:1 start:-65.5 TimeAxis = orientation:hor AlignBars = justify Colors = #legends id:CAR value:claret id:ANK value:rgb(0.4,0.3,0.196) id:HER value:teal id:HAD value:green id:OMN value:blue id:black value:black id:white value:white id:cenozoic value:rgb(0.54,0.54,0.258) id:paleogene value:rgb(0.99,0.6,0.32) id:paleocene value:rgb(0.99,0.65,0.37) id:eocene value:rgb(0.99,0.71,0.42) id:oligocene value:rgb(0.99,0.75,0.48) id:neogene value:rgb(0.999999,0.9,0.1) id:miocene value:rgb(0.999999,0.999999,0) id:pliocene value:rgb(0.97,0.98,0.68) id:quaternary value:rgb(0.98,0.98,0.5) id:pleistocene value:rgb(0.999999,0.95,0.68) id:holocene value:rgb(0.999,0.95,0.88) BarData= bar:eratop bar:space bar:periodtop bar:space bar:NAM1 bar:space bar:period bar:space bar:era PlotData= align:center textcolor:black fontsize:M mark:(line,black) width:25 shift:(7,-4) bar:periodtop from: -65.5 till: -55.8 color:paleocene text:
Paleocene The Paleocene, ( ) or Palaeocene, is a geological epoch that lasted from about 66 to 56 million years ago (mya). It is the first epoch of the Paleogene Period in the modern Cenozoic Era. The name is a combination of the Ancient Greek ''pal ...
from: -55.8 till: -33.9 color:eocene text:
Eocene The Eocene ( ) Epoch is a geological epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period in the modern Cenozoic Era. The name ''Eocene'' comes from the Ancient Greek (''ēṓs'', " ...
from: -33.9 till: -23.03 color:oligocene text:
Oligocene The Oligocene ( ) is a geologic epoch of the Paleogene Period and extends from about 33.9 million to 23 million years before the present ( to ). As with other older geologic periods, the rock beds that define the epoch are well identified but ...
from: -23.03 till: -5.332 color:miocene text:
Miocene The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recent" ...
from: -5.332 till: -2.588 color:pliocene text: Plio. from: -2.588 till: -0.0117 color:pleistocene text: Pleist. from: -0.0117 till: 0 color:holocene text: H. bar:eratop from: -65.5 till: -23.03 color:paleogene text:
Paleogene The Paleogene ( ; also spelled Palaeogene or Palæogene; informally Lower Tertiary or Early Tertiary) is a geologic period and system that spans 43 million years from the end of the Cretaceous Period million years ago (Mya) to the beginning of ...
from: -23.03 till: -2.588 color:neogene text:
Neogene The Neogene ( ), informally Upper Tertiary or Late Tertiary, is a geologic period and system that spans 20.45 million years from the end of the Paleogene Period million years ago (Mya) to the beginning of the present Quaternary Period Mya. ...
from: -2.588 till: 0 color:quaternary text: Q. PlotData= align:left fontsize:M mark:(line,white) width:5 anchor:till align:left color:miocene bar:NAM1 from: -11.608 till: 0 text: Labrisomus PlotData= align:center textcolor:black fontsize:M mark:(line,black) width:25 bar:period from: -65.5 till: -55.8 color:paleocene text:
Paleocene The Paleocene, ( ) or Palaeocene, is a geological epoch that lasted from about 66 to 56 million years ago (mya). It is the first epoch of the Paleogene Period in the modern Cenozoic Era. The name is a combination of the Ancient Greek ''pal ...
from: -55.8 till: -33.9 color:eocene text:
Eocene The Eocene ( ) Epoch is a geological epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period in the modern Cenozoic Era. The name ''Eocene'' comes from the Ancient Greek (''ēṓs'', " ...
from: -33.9 till: -23.03 color:oligocene text:
Oligocene The Oligocene ( ) is a geologic epoch of the Paleogene Period and extends from about 33.9 million to 23 million years before the present ( to ). As with other older geologic periods, the rock beds that define the epoch are well identified but ...
from: -23.03 till: -5.332 color:miocene text:
Miocene The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recent" ...
from: -5.332 till: -2.588 color:pliocene text: Plio. from: -2.588 till: -0.0117 color:pleistocene text: Pleist. from: -0.0117 till: 0 color:holocene text: H. bar:era from: -65.5 till: -23.03 color:paleogene text:
Paleogene The Paleogene ( ; also spelled Palaeogene or Palæogene; informally Lower Tertiary or Early Tertiary) is a geologic period and system that spans 43 million years from the end of the Cretaceous Period million years ago (Mya) to the beginning of ...
from: -23.03 till: -2.588 color:neogene text:
Neogene The Neogene ( ), informally Upper Tertiary or Late Tertiary, is a geologic period and system that spans 20.45 million years from the end of the Paleogene Period million years ago (Mya) to the beginning of the present Quaternary Period Mya. ...
from: -2.588 till: 0 color:quaternary text: Q.


References

{{Taxonbar, from=Q183463 Blenniiformes Taxa named by Clark Hubbs