Labeobarbus Aspius
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''Labeobarbus'' is a mid-sized
ray-finned fish Actinopterygii (; ), members of which are known as ray-finned fish or actinopterygians, is a class of bony fish that comprise over 50% of living vertebrate species. They are so called because of their lightly built fins made of webbings of sk ...
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
in the
family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
Cyprinidae Cyprinidae is a family of freshwater fish commonly called the carp or minnow family, including the carps, the true minnows, and their relatives the barbs and barbels, among others. Cyprinidae is the largest and most diverse fish family, and t ...
. Its
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
are widely distributed throughout eastern
Africa Africa is the world's second-largest and second-most populous continent after Asia. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 20% of Earth's land area and 6% of its total surfac ...
and especially
southern Africa Southern Africa is the southernmost region of Africa. No definition is agreed upon, but some groupings include the United Nations geoscheme for Africa, United Nations geoscheme, the intergovernmental Southern African Development Community, and ...
, but also in
Lake Tana Lake Tana (; previously transcribed Tsana) is the largest lake in Ethiopia and a source of the Blue Nile. Located in Amhara Region in the north-western Ethiopian Highlands, the lake is approximately long and wide, with a maximum depth of , and ...
in
Ethiopia Ethiopia, officially the Federal Democratic Republic of Ethiopia, is a landlocked country located in the Horn of Africa region of East Africa. It shares borders with Eritrea to the north, Djibouti to the northeast, Somalia to the east, Ken ...
. A
common name In biology, a common name of a taxon or organism (also known as a vernacular name, English name, colloquial name, country name, popular name, or farmer's name) is a name that is based on the normal language of everyday life; and is often con ...
, in particular for the southern species, is yellowfish. The
scientific name In Taxonomy (biology), taxonomy, binomial nomenclature ("two-term naming system"), also called binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, both of which use Latin gramm ...
refers to the fact that these large
barb Barb or the BARBs or ''variation'' may refer to: People * Barb (name), a list of people and fictional characters with the given name or surname * Barb, a term used by fans of Nicki Minaj to refer to themselves * The Barbs, a band Places * Barb, ...
s recall the fairly closely related "
carp The term carp (: carp) is a generic common name for numerous species of freshwater fish from the family (biology), family Cyprinidae, a very large clade of ray-finned fish mostly native to Eurasia. While carp are prized game fish, quarries and a ...
s" in the genus ''
Labeo ''Labeo'' is a genus of carps in the family Cyprinidae. They are found in freshwater habitats in the tropics and subtropics of Africa and Asia. It contains the typical labeos in the subfamily Labeoninae, which may not be a valid group, howeve ...
'' in size and shape. As far as can be told, all ''Labeobarbus'' species are
hexaploid Polyploidy is a condition in which the cells of an organism have more than two paired sets of ( homologous) chromosomes. Most species whose cells have nuclei (eukaryotes) are diploid, meaning they have two complete sets of chromosomes, one fro ...
.de Graaf ''et al.'' (2007), IUCN (2009) One species, '' L. microbarbis'' from
Rwanda Rwanda, officially the Republic of Rwanda, is a landlocked country in the Great Rift Valley of East Africa, where the African Great Lakes region and Southeast Africa converge. Located a few degrees south of the Equator, Rwanda is bordered by ...
, is known to have gone
extinct Extinction is the termination of an organism by the death of its Endling, last member. A taxon may become Functional extinction, functionally extinct before the death of its last member if it loses the capacity to Reproduction, reproduce and ...
in recent times.


Systematics

Like many other "barbs", it was long included in ''
Barbus ''Barbus'' is a genus of ray-finned fish in the family Cyprinidae. The type species of ''Barbus'' is the common barbel, first described as ''Cyprinus barbus'' and now named ''Barbus barbus''. ''Barbus'' is the namesake genus of the subfamily Ba ...
''. It appears to be a fairly close relative of the typical
barbel Barbel may refer to: *Barbel (anatomy), a whisker-like organ near the mouth found in some fish (notably catfish, loaches and cyprinids) and turtles *Barbel (fish), a common name for certain species of fish **''Barbus barbus'', a species of cyprini ...
s and relatives – the genus ''Barbus'' proper – but closer still to the large
Near East The Near East () is a transcontinental region around the Eastern Mediterranean encompassing the historical Fertile Crescent, the Levant, Anatolia, Egypt, Mesopotamia, and coastal areas of the Arabian Peninsula. The term was invented in the 20th ...
ern species nowadays separated in ''
Carasobarbus ''Carasobarbus'', the himris, is a small genus of ray-finned fishes in the family Cyprinidae. Its species are found in rivers, streams, lakes and ponds in Western Asia and Northwest Africa. ''C. canis'' can reach in total length, but most other ...
''. ''Barbus'' has been split to account for the improved
phylogenetic In biology, phylogenetics () is the study of the evolutionary history of life using observable characteristics of organisms (or genes), which is known as phylogenetic inference. It infers the relationship among organisms based on empirical dat ...
knowledge which indicated it was highly
paraphyletic Paraphyly is a taxonomic term describing a grouping that consists of the grouping's last common ancestor and some but not all of its descendant lineages. The grouping is said to be paraphyletic ''with respect to'' the excluded subgroups. In co ...
in its wide circumscription; it may be that ''Carasobarbus'' and some other closely related "barbs" (e.g. ''
Labeobarbus reinii The giant Atlas barbel (''Atlantor reinii'') was a ray-finned fish species in the family Cyprinidae. It was the only species in the genus ''Atlantor'' but it is now thought to be extinct. It was previously considered to be fairly close to the c ...
'') are now included in ''Labeobarbus'' to avoid a profusion of very small genera.de Graaf ''et al.'' (2007) The
taxonomy image:Hierarchical clustering diagram.png, 280px, Generalized scheme of taxonomy Taxonomy is a practice and science concerned with classification or categorization. Typically, there are two parts to it: the development of an underlying scheme o ...
of many species in the " wastebin genus" ''Barbus'' has recently been re-evaluated. Though
hybrid Hybrid may refer to: Science * Hybrid (biology), an offspring resulting from cross-breeding ** Hybrid grape, grape varieties produced by cross-breeding two ''Vitis'' species ** Hybridity, the property of a hybrid plant which is a union of two diff ...
introgression may confound studies based in
mtDNA Mitochondrial DNA (mtDNA and mDNA) is the DNA located in the mitochondria organelles in a eukaryotic cell that converts chemical energy from food into adenosine triphosphate (ATP). Mitochondrial DNA is a small portion of the DNA contained in ...
data alone, a number of these species appear to be so closely related to ''Labeobarbus'' as to warrant inclusion in the present genus outright, irrespective of whether ''Carasobarbus'' is considered distinct or not. These include '' L. ethiopicus''.


Species

''Labeobarbus'' at present contains the following
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
: * ''
Labeobarbus acuticeps ''Labeobarbus acuticeps'' is a species of ray-finned fish in the family Cyprinidae. It is found in Burundi, Rwanda, and Tanzania. Its natural habitats are rivers, freshwater lakes, freshwater marshes, and inland deltas. It is threatened by habit ...
'' ( Matthes, 1959) * '' Labeobarbus acutirostris'' ( Bini, 1940) * ''
Labeobarbus aeneus The smallmouth yellowfish (''Labeobarbus aeneus'') is a species of ray-finned fish in the genus ''Labeobarbus''. It has become an invasive species in rivers of the Eastern Cape, South Africa, such as the Mbhashe River. Introduction The Orange-Va ...
'' ( Burchell, 1822) (Smallmouth yellowfish) * '' Labeobarbus altianalis'' ( Boulenger, 1900) (Ripon barbel) * ''
Labeobarbus altipinnis ''Labeobarbus altipinnis'' is a species of ray-finned fish in the family Cyprinidae. It is endemic to the Lufira River The Lufira River is a tributary of the Lualaba River in the Democratic Republic of the Congo (DRC). The Lufira rises in the ...
'' (
Banister A handrail is a rail that is designed to be grasped by the hand so as to provide safety or support. In Britain, handrails are referred to as banisters. Handrails are commonly used while ascending or descending stairways and escalators in order t ...
&
Poll Poll, polled, or polling may refer to: Forms of voting and counting * Poll, a formal election ** Election verification exit poll, a survey taken to verify election counts ** Polling, voting to make decisions or determine opinions ** Polling pla ...
, 1973)
* '' Labeobarbus ansorgii'' (Boulenger, 1906) * '' Labeobarbus aspius'' (Boulenger, 1912) * '' Labeobarbus axelrodi'' ( Getahun, Stiassny & Teugels, 2004) * '' Labeobarbus batesii'' (Boulenger, 1903) * '' Labeobarbus beso'' ( Rüppell, 1835) (African scraping feeder) * '' Labeobarbus boulengeri'' Vreven, Musschoot, Snoeks & Schliewen, 2016 * '' Labeobarbus brauni'' ( Pellegrin, 1935) * ''
Labeobarbus brevicephalus ''Labeobarbus brevicephalus'' is a species of ray-finned fish in the genus ''Labeobarbus'' which is endemic to Lake Tana in Ethiopia. The ''L. brevicephalus'' is one of 17 Endemism, endemic migratory African barb species that makes a species flo ...
'' ( Nagelkerke & Sibbing, 1997) * '' Labeobarbus brevispinis'' (
Holly ''Ilex'' () or holly is a genus of over 570 species of flowering plants in the family Aquifoliaceae, and the only living genus in that family. ''Ilex'' has the most species of any woody dioecious angiosperm genus. The species are evergreen o ...
, 1927)
* '' Labeobarbus bynni'' (
Fabricius Fabricius (, ) is a surname. Notable people with the surname include: *people from the Ancient Roman Fabricia gens, gens Fabricia: **Gaius Fabricius Luscinus, the first of the Fabricii to move to Rome * Johann Goldsmid (1587–1616), known by his ...
, 1775)
* '' Labeobarbus cardozoi'' (Boulenger, 1912) * '' Labeobarbus caudovittatus'' (Boulenger, 1902) * ''
Labeobarbus clarkeae ''Labeobarbus'' is a mid-sized ray-finned fish genus in the family Cyprinidae. Its species are widely distributed throughout eastern Africa and especially southern Africa, but also in Lake Tana in Ethiopia. A common name, in particular for the s ...
'' (Banister, 1984) * '' Labeobarbus claudinae'' ( De Vos & Thys van den Audenaerde, 1990) * '' Labeobarbus codringtonii'' (Boulenger, 1908) (Upper Zambezi yellowfish) * '' Labeobarbus compiniei'' ( Sauvage,1879) * '' Labeobarbus crassibarbis'' (Nagelkerke & Sibbing, 1997) * '' Labeobarbus dainellii'' (Bini, 1940) * '' Labeobarbus dartevellei'' (
Poll Poll, polled, or polling may refer to: Forms of voting and counting * Poll, a formal election ** Election verification exit poll, a survey taken to verify election counts ** Polling, voting to make decisions or determine opinions ** Polling pla ...
, 1945)
* '' Labeobarbus dimidiatus'' ( Tweddle & P. H. Skelton, 1998) * '' Labeobarbus ensifer'' (Boulenger, 1910) * '' Labeobarbus ensis'' (Boulenger, 1910) * ''
Labeobarbus ethiopicus ''Labeobarbus ethiopicus'' is a species of ray-finned fish, usually placed in the genus ''Labeobarbus''. It is endemic to Lake Ziway, in Ethiopia Ethiopia, officially the Federal Democratic Republic of Ethiopia, is a landlocked country ...
'' ( Zolezzi, 1939) * '' Labeobarbus fimbriatus'' (Holly, 1926) * '' Labeobarbus gananensis'' ( Vinciguerra, 1895) * '' Labeobarbus gestetneri'' (Banister & R. G. Bailey, 1979) * '' Labeobarbus girardi'' (Boulenger, 1910) * '' Labeobarbus gorgorensis'' (Bini, 1940) * '' Labeobarbus gorguari'' ( Rüppell, 1835) * '' Labeobarbus gruveli'' (Pellegrin, 1911) * '' Labeobarbus gulielmi'' (Boulenger, 1910) * ''
Labeobarbus habereri ''Labeobarbus habereri'' is a species of Cyprinid fish endemic to Cameroon in Africa Africa is the world's second-largest and second-most populous continent after Asia. At about 30.3 million km2 (11.7 million square miles) i ...
'' (
Steindachner Franz Steindachner (11 November 1834 in Vienna – 10 December 1919 in Vienna) was an Austrian zoologist, ichthyologist, and herpetologist. He published over 200 papers on fishes and over 50 papers on reptiles and amphibians. Steindachner descri ...
, 1912)
* '' Labeobarbus huloti'' (Banister, 1976) * '' Labeobarbus humphri'' (Banister, 1976) * ''
Labeobarbus intermedius ''Labeobarbus intermedius'' is an East African ray-finned fish species in the family Cyprinidae. Like the closely related yellowfish, it is hexaploid. A large species, the maximum recorded standard length is nearly . This species has a subspeci ...
'' (Rüppell, 1835) * '' Labeobarbus iphthimostoma'' (Banister & Poll, 1973) * '' Labeobarbus iturii'' (Holly, 1929) * ''
Labeobarbus jaegeri ''Labeobarbus'' is a mid-sized ray-finned fish genus in the family Cyprinidae. Its species are widely distributed throughout eastern Africa and especially southern Africa, but also in Lake Tana in Ethiopia. A common name, in particular for the s ...
'' (Holly, 1930) * '' Labeobarbus johnstonii'' (Boulenger, 1907) * '' Labeobarbus jubae'' (Banister, 1984) * '' Labeobarbus jubbi'' (Poll, 1967) * '' Labeobarbus kimberleyensis'' ( Gilchrist & W. W. Thompson, 1913) (Largemouth yellowfish) * '' Labeobarbus lagensis'' (Günther, 1868) * '' Labeobarbus latirostris'' ( Keilhack, 1908) * '' Labeobarbus leleupanus'' (Matthes, 1959) * ''
Labeobarbus lobogenysoides ''Labeobarbus'' is a mid-sized ray-finned fish genus in the family Cyprinidae. Its species are widely distributed throughout eastern Africa and especially southern Africa, but also in Lake Tana in Ethiopia. A common name, in particular for the s ...
'' (Pellegrin, 1935) * '' Labeobarbus longidorsalis'' (Pellegrin, 1935) * ''
Labeobarbus longifilis ''Labeobarbus'' is a mid-sized ray-finned fish genus in the family Cyprinidae. Its species are widely distributed throughout eastern Africa and especially southern Africa, but also in Lake Tana in Ethiopia. A common name, in particular for the s ...
'' (Pellegrin, 1935) * '' Labeobarbus longissimus'' (Nagelkerke & Sibbing, 1997) * '' Labeobarbus lucius'' (Boulenger, 1910) * '' Labeobarbus lufupensis'' (Banister & Bailey, 1979) * '' Labeobarbus macroceps'' ( Fowler 1936) * '' Labeobarbus macrolepidotus'' (Pellegrin, 1928) * '' Labeobarbus macrolepis'' (
Pfeffer Pfeffer is a German language, German surname meaning "Black pepper, pepper" and may refer to: * Anna Pfeffer (born 1946), Hungarian Olympic medalist sprint canoer * Anshel Pfeffer, British journalist * Anton Pfeffer (born 1965), Austrian football ...
, 1889)
* '' Labeobarbus macrophtalmus'' (Bini, 1940) * '' Labeobarbus malacanthus'' (
Pappenheim Pappenheim is a town in the Weißenburg-Gunzenhausen district, in Bavaria, Germany. It is situated on the river Altmühl, south of Weißenburg in Bayern. History Historically, Pappenheim was a statelet within the Holy Roman Empire. It was ...
, 1911)
* ''
Labeobarbus marequensis The largescale yellowfish or Lowveld largescale yellowfish (''Labeobarbus marequensis'') is a species of fish in the family ''Cyprinidae''. An African freshwater fish found from the Zambezi south to the Pongola River The Phongolo River is a rive ...
'' (
A. Smith A is the first letter of the Latin and English alphabet. A may also refer to: Science and technology Quantities and units * ''a'', a measure for the attraction between particles in the Van der Waals equation * ''A'' value, a measure of s ...
1841)
(Largescale yellowfish) * '' Labeobarbus mariae'' (Holly, 1926) * '' Labeobarbus matris'' (Holly, 1928). * ''
Labeobarbus mawambiensis ''Labeobarbus mawambiensis'' is a species of ray-finned fish in the genus ''Labeobarbus'' from the central Congo Basin The Congo Basin () is the sedimentary basin of the Congo River. The Congo Basin is located in Central Africa, in a region kn ...
'' (Steindachner, 1911) * '' Labeobarbus mbami'' (Holly, 1927) * '' Labeobarbus megastoma'' (Nagelkerke & Sibbing, 1997) * '' Labeobarbus micronema'' (Boulenger, 1904) * ''
Labeobarbus mirabilis ''Labeobarbus mirabilis'' is a species of ray-finned fish in the genus ''Labeobarbus''. Its type locality is the Ituri River, near Mawambi, Ituri Province, Democratic Republic of the Congo. It has not been found in other locations. This species ...
'' (Pappenheim, 1914) * '' Labeobarbus mungoensis'' ( Trewavas, 1974) * '' Labeobarbus nanningsi'' de Beaufort, 1933 * '' Labeobarbus natalensis'' ( Castelnau, 1861) (Scaly yellowfish) * ''
Labeobarbus nedgia ''Labeobarbus nedgia'' is a species of ray-finned fish in the genus ''Labeobarbus'' which is endemic to Lake Tana Lake Tana (; previously transcribed Tsana) is the largest lake in Ethiopia and a source of the Blue Nile. Located in Amhara Regio ...
'' Rüppell, 1835 * '' Labeobarbus nelspruitensis'' (Gilchrist & W. W. Thompson, 1911) (Incomati chiselmouth) * '' Labeobarbus nthuwa'' Tweddle & Skelton, 2008 * '' Labeobarbus nzadimalawu'' Vreven, Musschoot, Decru, Lunkayilakio, Obiero, Cerwenka & Schliewen, 2018. * '' Labeobarbus nzadinkisi'' Vreven, Musschoot, Decru, Lunkayilakio, Obiero, Cerwenka & Schliewen, 2018. * '' Labeobarbus osseensis'' (Nagelkerke & Sibbing, 2000) * '' Labeobarbus oxyrhynchus'' (Pfeffer, 1889) (Pangani barb) * '' Labeobarbus pagenstecheri'' ( J. G. Fischer, 1884) * '' Labeobarbus parawaldroni'' ( Lévêque, Thys van den Audenaerde &
Traoré Traoré or Traore is a surname of Manding origin (), as written in French orthography, which is common in Mali, Senegal, Burkina Faso, Ivory Coast and Guinea. In anglophone West Africa the name is often spelled Trawally. Notable people with the s ...
, 1987)
* '' Labeobarbus pellegrini'' (
Bertin Bertin (; 615 – ''c''. 709 AD), also known as Saint Bertin the Great, was the Frankish abbot of a monastery in Saint-Omer later named the Abbey of Saint Bertin after him. He is venerated as a saint by the Catholic and Orthodox Churches. The ...
& Estève, 1948)
* '' Labeobarbus petitjeani'' (
Daget Jacques Daget (30 June 1919, Vineuil – 29 June 2009), was a French ichthyologist. He was a professor at the National Museum of Natural History, in Paris. Several marine species have been named after him. Species named after him Species named ...
, 1962)
* '' Labeobarbus platydorsus'' (Nagelkerke & Sibbing, 1997) * '' Labeobarbus platyrhinus'' (Boulenger, 1900) * ''
Labeobarbus platystomus ''Labeobarbus'' is a mid-sized ray-finned fish genus in the family Cyprinidae. Its species are widely distributed throughout eastern Africa and especially southern Africa, but also in Lake Tana in Ethiopia. A common name, in particular for the s ...
'' (Pappenheim, 1914) (Rwandese carp) * '' Labeobarbus pojeri'' (Poll, 1944) * '' Labeobarbus polylepis'' (Boulenger, 1907) (Smallscale yellowfish) * '' Labeobarbus progenys'' (Boulenger, 1903) * '' Labeobarbus pungweensis'' ( Jubb, 1959) (Pungwe chiselmouth) * '' Labeobarbus rhinoceros'' ( Copley, 1938) (Rhinofish) * '' Labeobarbus rhinophorus'' (Boulenger, 1910) * '' Labeobarbus robertsi'' (Banister, 1984) * '' Labeobarbus rocadasi'' (Boulenger, 1910) * '' Labeobarbus rosae'' (Boulenger, 1910). * '' Labeobarbus roylii'' (Boulenger, 1912) * '' Labeobarbus ruandae'' (Pappenheim, 1914) * '' Labeobarbus ruasae'' (Pappenheim, 1914) * ''
Labeobarbus ruwenzorii ''Labeobarbus ruwenzorii'' is a species of ray-finned fish in the family Cyprinidae. It is found only in Uganda. Its natural habitat is rivers. It reproduces by inserting its seminiferous tubule Seminiferous tubules are located within the te ...
'' (Pellegrin, 1909) * '' Labeobarbus sacratus'' (Daget, 1963) * '' Labeobarbus sandersi'' (Boulenger, 1912) * '' Labeobarbus seeberi'' (Gilchrist & W. W. Thompson, 1913) (Clanwilliam yellowfish) * '' Labeobarbus semireticulatus'' (Pellegrin, 1924) * ''
Labeobarbus somereni ''Labeobarbus somereni'', or Someren's barb, is a species of ray-finned fish in the family Cyprinidae. It is found in Burundi, Rwanda, Tanzania, and Uganda. Its natural habitat is rivers. It is not considered a threatened species by the IUCN. Lo ...
'' (Boulenger, 1911) * '' Labeobarbus stappersii'' (Boulenger, 1915) * '' Labeobarbus steindachneri'' (Boulenger, 1910) * '' Labeobarbus stenostoma'' (Boulenger, 1910) * '' Labeobarbus surkis'' (Rüppell, 1835) * '' Labeobarbus tornieri'' (Steindachner, 1906) * '' Labeobarbus trachypterus'' (Boulenger, 1915) * '' Labeobarbus tropidolepis'' (Boulenger, 1900) * '' Labeobarbus truttiformis'' (Nagelkerke & Sibbing, 1997) * '' Labeobarbus tsanensis'' (Nagelkerke & Sibbing, 1997)) * '' Labeobarbus upembensis'' (Banister & Bailey, 1979) * ''
Labeobarbus urotaenia ''Labeobarbus urotaenia'' is a species of ray-finned fish in the genus ''Labeobarbus'' from the central Congo River system. References

* Labeobarbus, urotaenia Taxa named by George Albert Boulenger Fish described in 1913 Endemic fauna o ...
'' (Boulenger, 1913) * ''
Labeobarbus varicostoma ''Labeobarbus varicostoma'' is a species of ray-finned fish in the genus ''Labeobarbus'' is endemic to the Lucalla River in Angola Angola, officially the Republic of Angola, is a country on the west-Central Africa, central coast of Southe ...
'' (Boulenger, 1910) * '' Labeobarbus versluysii'' (Holly, 1929) * '' Labeobarbus werneri'' (Holly, 1929) * '' Labeobarbus wittei'' (Banister & Poll, 1973) * ''
Labeobarbus wurtzi ''Labeobarbus wurtzi'' is a species of ray-finned fish in the genus ''Labeobarbus ''Labeobarbus'' is a mid-sized ray-finned fish genus in the family Cyprinidae. Its species are widely distributed throughout eastern Africa and especially south ...
'' (Pellegrin, 1908) * '' Labeobarbus xyrocheilus'' (Tweddle & Skelton, 1998)


Footnotes


References

* * (2007): Evolutionary origin of Lake Tana's (Ethiopia) small ''Barbus'' species: indications of rapid ecological divergence and speciation. '' Anim. Biol.'' 57(1): 39–48. (HTML abstract) * (2009):
2009 IUCN Red List of Threatened Species
'. Version 2009.1. Retrieved 2009-SEP-20. {{Taxonbar, from=Q2193181 Torinae Taxa named by Eduard Rüppell Cyprinidae genera