Koklass
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The koklass pheasant (''Pucrasia macrolopha'') is a species of
gamebird Galliformes is an order of heavy-bodied ground-feeding birds that includes turkeys, chickens, quail, and other landfowl. Gallinaceous birds, as they are called, are important in their ecosystems as seed dispersers and predators, and are ofte ...
, being closely related to progenitive
grouse Grouse are a group of birds from the order (biology), order Galliformes, in the family (biology), family Phasianidae. Grouse are presently assigned to the Tribe (biology), tribe Tetraonini (formerly the subfamily Tetraoninae and the family Tetr ...
that lived during the
Miocene The Miocene ( ) is the first epoch (geology), geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and mea ...
. They are distantly related to
pheasants Pheasants ( ) are birds of several genera within the family Phasianidae in the order Galliformes. Although they can be found all over the world in introduced (and captive) populations, the pheasant genera's native range is restricted to Euras ...
and are most closely related to
grouse Grouse are a group of birds from the order (biology), order Galliformes, in the family (biology), family Phasianidae. Grouse are presently assigned to the Tribe (biology), tribe Tetraonini (formerly the subfamily Tetraoninae and the family Tetr ...
and
turkeys The turkey is a large bird in the genus ''Meleagris'', native to North America. There are two extant turkey species: the wild turkey (''Meleagris gallopavo'') of eastern and central North America and the ocellated turkey (''Meleagris ocell ...
. Koklass are the only species in the monotypic genus ''Pucrasia''. Both the words ''koklass'' and ''pucrasia'' have been onomatopœically derived from the bird's territorial call. Koklass are boreal adapted species which separate into three distinct species groups. They are one of the few gamebirds that regularly fly uphill and are capable of sustained flights of many miles. They are monogamous with a slight tendency toward social
polyandry Polyandry (; ) is a form of polygamy in which a woman takes two or more husbands at the same time. Polyandry is contrasted with polygyny, involving one male and two or more females. If a marriage involves a plural number of "husbands and wives ...
. Both parents rear the chicks. Koklass are largely vegetarian for much of the year consuming pine nuts, pine shoots, bamboo shoots and seeds. They are highly insectivorous during the warmer months that coincide with nesting and chick-rearing. During this phase of their life cycle they live almost exclusively on ants but also are documented consuming
catkins A catkin or ament is a slim, cylindrical flower cluster (a spike), with inconspicuous or no petals, usually wind-pollinated (anemophilous) but sometimes insect-pollinated (as in ''Salix''). It contains many, usually unisexual flowers, arranged cl ...
,
pollen Pollen is a powdery substance produced by most types of flowers of seed plants for the purpose of sexual reproduction. It consists of pollen grains (highly reduced Gametophyte#Heterospory, microgametophytes), which produce male gametes (sperm ...
and fruit. The koklass pheasant is a polytypic species, with nine recognized
subspecies In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...
: * Kashmir koklass pheasant - ''P. m. biddulphi'' (Marshall G.F.L, 1879) * Western koklass pheasant - ''P. m. castanea'' (Gould, 1854) * Darwin's koklass pheasant - ''P. m. darwini'' ( Swinhoe, 1872) * Joret's koklass pheasant - ''P. m. joretiana'' (Heude, 1883) * Indian koklass pheasant - ''P. m. macrolopha'' (Lesson R.P., 1829) * Meyer's koklass pheasant - ''P. m. meyeri'' (Madarász, 1886) *
Nepal koklass pheasant Nepal, officially the Federal Democratic Republic of Nepal, is a landlocked country in South Asia. It is mainly situated in the Himalayas, but also includes parts of the Indo-Gangetic Plain. It borders the Tibet Autonomous Region of China t ...
- ''P. m. nipalensis'' (Gould, 1854) * Orange-collared koklass pheasant - ''P. m. ruficollis'' (David A. & Oustalet, 1877) * Yellow-necked koklass pheasant - ''P. m. xanthospila'' (Gray & GR, 1886) This entry deals with the subspecies ''P. m. biddulphi'', which ranges from
Kashmir Kashmir ( or ) is the Northwestern Indian subcontinent, northernmost geographical region of the Indian subcontinent. Until the mid-19th century, the term ''Kashmir'' denoted only the Kashmir Valley between the Great Himalayas and the Pir P ...
to
Kullu Kullu () is a municipal council town that serves as the administrative headquarters of the Kullu district of the Indian state of Himachal Pradesh. It is located on the banks of the Beas River in the Kullu Valley about north of the airport ...
in
India India, officially the Republic of India, is a country in South Asia. It is the List of countries and dependencies by area, seventh-largest country by area; the List of countries by population (United Nations), most populous country since ...
. With exception of the subspecies ''P. m. nipalensis'', ''P. m. castanea'' and ''P. m. macrolopha'', which are
endemic Endemism is the state of a species being found only in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also foun ...
to the southern side of northwest and western
Himalaya The Himalayas, or Himalaya ( ), is a mountain range in Asia, separating the plains of the Indian subcontinent from the Tibetan Plateau. The range has some of the Earth's highest peaks, including the highest, Mount Everest. More than 100 pea ...
, the other five are confined to
China China, officially the People's Republic of China (PRC), is a country in East Asia. With population of China, a population exceeding 1.4 billion, it is the list of countries by population (United Nations), second-most populous country after ...
and
Mongolia Mongolia is a landlocked country in East Asia, bordered by Russia to the north and China to the south and southeast. It covers an area of , with a population of 3.5 million, making it the world's List of countries and dependencies by po ...
. The koklass pheasant is a medium-sized elusive bird confined to high altitude
forest A forest is an ecosystem characterized by a dense ecological community, community of trees. Hundreds of definitions of forest are used throughout the world, incorporating factors such as tree density, tree height, land use, legal standing, ...
s from
Afghanistan Afghanistan, officially the Islamic Emirate of Afghanistan, is a landlocked country located at the crossroads of Central Asia and South Asia. It is bordered by Pakistan to the Durand Line, east and south, Iran to the Afghanistan–Iran borde ...
to central
Nepal Nepal, officially the Federal Democratic Republic of Nepal, is a landlocked country in South Asia. It is mainly situated in the Himalayas, but also includes parts of the Indo-Gangetic Plain. It borders the Tibet Autonomous Region of China Ch ...
, and in northeastern
Tibet Tibet (; ''Böd''; ), or Greater Tibet, is a region in the western part of East Asia, covering much of the Tibetan Plateau and spanning about . It is the homeland of the Tibetan people. Also resident on the plateau are other ethnic groups s ...
to northern and eastern
China China, officially the People's Republic of China (PRC), is a country in East Asia. With population of China, a population exceeding 1.4 billion, it is the list of countries by population (United Nations), second-most populous country after ...
. Upper parts of male koklass pheasant are covered with silver-grey
plumage Plumage () is a layer of feathers that covers a bird and the pattern, colour, and arrangement of those feathers. The pattern and colours of plumage differ between species and subspecies and may vary with age classes. Within species, there can b ...
streaked velvety-black down the centre of each feather, and it has the unique feature of a black head, chestnut breast and prominent white patches on the sides of neck. The females differ from males in above characters and instead their upper parts are covered with pale brown plumage. Both sexes, however, have distinct elongated tails tipped with pale feathers. The males are known to weigh about 1135–1415 g and the females, about 1025–1135 g, with the body length varying from 58 to 64 cm and 18–22 cm respectively. Immature and juveniles resemble adult females in plumage pattern. Like the
western tragopan The western tragopan or western horned tragopan (''Tragopan melanocephalus'') is a medium-sized brightly plumed pheasant found along the range of Himalayas from north-eastern districts of Khyber Pakhtunkhwa province in northern Pakistan in the we ...
, it does not extend its range above the tree line. One of the less colourful pheasants, the koklass exhibits moderate
sexual dimorphism Sexual dimorphism is the condition where sexes of the same species exhibit different Morphology (biology), morphological characteristics, including characteristics not directly involved in reproduction. The condition occurs in most dioecy, di ...
. Though they skulk under bushes, which makes direct sighting difficult, they give loud chorus/predawn calls during the breeding season and during autumn, revealing their presence and allowing the populations to be estimated. They remain in pairs or small family groups throughout the year. They nest on the ground and spend the nights roosting on trees, or under rock overhangs.


Gallery

File:Koklass Pheasant @ Manila.jpg, ''Pucrasia macrolopha macrolopha'' Male at Manila,
Uttarakhand Uttarakhand (, ), also known as Uttaranchal ( ; List of renamed places in India, the official name until 2007), is a States and union territories of India, state in North India, northern India. The state is bordered by Himachal Pradesh to the n ...
File:Koklass Pheasant Manila3.jpg, ''Pucrasia macrolopha macrolopha'' Male at Manila,
Uttarakhand Uttarakhand (, ), also known as Uttaranchal ( ; List of renamed places in India, the official name until 2007), is a States and union territories of India, state in North India, northern India. The state is bordered by Himachal Pradesh to the n ...


References

{{Authority control
koklass pheasant The koklass pheasant (''Pucrasia macrolopha'') is a species of gamebird, being closely related to progenitive grouse that lived during the Miocene. They are distantly related to pheasants and are most closely related to grouse and turkeys. Kok ...
koklass pheasant The koklass pheasant (''Pucrasia macrolopha'') is a species of gamebird, being closely related to progenitive grouse that lived during the Miocene. They are distantly related to pheasants and are most closely related to grouse and turkeys. Kok ...
Birds of Afghanistan Birds of China Birds of the Himalayas Birds of North India Birds of Nepal Birds of Pakistan
koklass pheasant The koklass pheasant (''Pucrasia macrolopha'') is a species of gamebird, being closely related to progenitive grouse that lived during the Miocene. They are distantly related to pheasants and are most closely related to grouse and turkeys. Kok ...
Taxa named by René Lesson