Khebbashia is a clade of herbivorous
sauropod
Sauropoda (), whose members are known as sauropods (; from '' sauro-'' + '' -pod'', 'lizard-footed'), is a clade of saurischian ('lizard-hipped') dinosaurs. Sauropods had very long necks, long tails, small heads (relative to the rest of their ...
dinosaurs
Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...
belonging to the
Rebbachisauridae
Rebbachisauridae is a family of sauropod dinosaurs known from fragmentary fossil remains from the Cretaceous of South America, Africa, North America, Europe and possibly Central Asia.
Taxonomy
In 1990 sauropod specialist Jack McIntosh included ...
.
[ P. D. Mannion, P. Upchurch, D. Schwarz and O. Wings. (2019). Taxonomic affinities of the putative ]titanosaur
Titanosaurs (or titanosaurians; members of the group Titanosauria) were a diverse group of sauropod dinosaurs, including genera from all seven continents. The titanosaurs were the last surviving group of long-necked sauropods, with taxa still th ...
s from the Late Jurassic
The Late Jurassic is the third epoch of the Jurassic Period, and it spans the geologic time from 163.5 ± 1.0 to 145.0 ± 0.8 million years ago (Ma), which is preserved in Upper Jurassic strata.Owen 1987.
In European lithostratigraphy, the ...
Tendaguru Formation
The Tendaguru Formation, or Tendaguru Beds are a highly fossiliferous formation and Lagerstätte located in the Lindi Region of southeastern Tanzania. The formation represents the oldest sedimentary unit of the Mandawa Basin, overlying Neoprote ...
of Tanzania
Tanzania (; ), officially the United Republic of Tanzania ( sw, Jamhuri ya Muungano wa Tanzania), is a country in East Africa within the African Great Lakes region. It borders Uganda to the north; Kenya to the northeast; Comoro Islands ...
: phylogenetic and biogeographic implications for eusauropod
Eusauropoda (meaning "true sauropods") is a derived clade of sauropod dinosaurs. Eusauropods represent the node-based group that includes all descendant sauropods starting with the basal eusauropods of '' Shunosaurus'', and possibly '' Barapasaur ...
dinosaur evolution. ''Zoological Journal of the Linnean Society
The ''Zoological Journal of the Linnean Society'' is a monthly peer-reviewed scientific journal covering zoology published by Oxford University Press on behalf of the Linnean Society. The editor-in-chief is Maarten Christenhusz (Linnean Society). ...
'' 185(3):784-909 Members of Khebbashia were medium-sized sauropods from the early Cretaceous period of South America, Africa and Europe.
The name "Khebbashia" is derived from "Khebbash" or "Khebbache", a Moroccan tribe that inhabited the region where the first rebbachisaurid specimen was found in North Africa.
Khebbashia is defined as the least inclusive clade including ''
Limaysaurus
''Limaysaurus'' ("Limay lizard") is a genus represented by a single species of rebbachisaurid sauropod dinosaurs, which lived during the mid-Cretaceous period, about 99.6 to 97 million years ago, in the Cenomanian, in what is now South America ( ...
tessonei'', ''
Nigersaurus
''Nigersaurus'' is a genus of rebbachisaurid sauropod dinosaur that lived during the middle Cretaceous period, about 115 to 105 million years ago. It was discovered in the Elrhaz Formation in an area called Gadoufaoua, in Niger. Fossils of t ...
taqueti'', and ''
Rebbachisaurus
''Rebbachisaurus'' (meaning " Aït Rebbach lizard") is a genus of sauropod dinosaur of the superfamily Diplodocoidea, that lived during the Late Cretaceous period in Africa and possibly also South America about 99-97 million years ago. Remains at ...
garasbae''. It therefore includes the rebbachisaurid subfamilies
Rebbachisaurinae
Rebbachisaurinae is a subfamily within the family Rebbachisauridae, defined to include '' Rebbachisaurus garasbae'' and exclude '' Limaysaurus tessonei''. It was first proposed as a rank by Jose Bonaparte in 1995, to include ''Rebbachisaurus''. ...
and
Limaysaurinae, to the exclusion of more basal forms.
References
Rebbachisaurids
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