''Kermitops'' (meaning " Kermit face") is an

extinct Extinction is the termination of an organism by the death of its Endling, last member. A taxon may become Functional extinction, functionally extinct before the death of its last member if it loses the capacity to Reproduction, reproduce and ...

genus of

amphibamiform Amphibamiformes is an unranked clade with Dissorophoidea created by Schoch (2018). It encompasses all of the taxa traditionally considered to be "Amphibamidae, amphibamids" (subsequently restricted to ''Doleserpeton, Doleserpeton annectens'' and ...

temnospondyl Temnospondyli (from Greek language, Greek τέμνειν, ''temnein'' 'to cut' and σπόνδυλος, ''spondylos'' 'vertebra') or temnospondyls is a diverse ancient order (biology), order of small to giant tetrapods—often considered Labyrinth ...

from the

Early Permian 01 or 01 may refer to: * The year 2001, or any year ending with 01 * The month of January * 1 (number) Music * '01 (Richard Müller album), ''01'' (Richard Müller album), 2001 * 01 (Urban Zakapa album), ''01'' (Urban Zakapa album), 2011 * ''01011 ...

Clear Fork Formation of

Texas Texas ( , ; or ) is the most populous U.S. state, state in the South Central United States, South Central region of the United States. It borders Louisiana to the east, Arkansas to the northeast, Oklahoma to the north, New Mexico to the we ...

, United States. The genus contains a single species, ''K. gratus'', known from a partial cranium.

Discovery and naming

The ''Kermitops''

holotype A holotype (Latin: ''holotypus'') is a single physical example (or illustration) of an organism used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of s ...

specimen,

USNM The National Museum of Natural History (NMNH) is a natural history museum administered by the Smithsonian Institution, located on the National Mall in Washington, D.C., United States. It has free admission and is open 364 days a year. With 4.4 ...

407585, was discovered by Nicholas Hotton III and field scientists from the Smithsonian Museum of Natural History in April 1984. The type locality is represented by the lower sediments of the Clear Fork Formation near Lake Kemp in

Wilbarger County Wilbarger County ( ) is a county located in the North Texas region of the U.S. state of Texas. As of the 2020 census, the population was 12,887. The county seat is Vernon. The county was created in 1858 and later organized in 1881. Wilbarg ...

, United States. The specimen consists of most of the

skull roof The skull roof or the roofing bones of the skull are a set of bones covering the brain, eyes and nostrils in bony fishes, including land-living vertebrates. The bones are derived from dermal bone and are part of the dermatocranium. In com ...

, a partial

braincase In human anatomy, the neurocranium, also known as the braincase, brainpan, brain-pan, or brainbox, is the upper and back part of the skull, which forms a protective case around the brain. In the human skull, the neurocranium includes the calv ...

, and the

mandibles In jawed vertebrates, the mandible (from the Latin ''mandibula'', 'for chewing'), lower jaw, or jawbone is a bone that makes up the lowerand typically more mobilecomponent of the mouth (the upper jaw being known as the maxilla). The jawbone i ...

. Most of the

palate The palate () is the roof of the mouth in humans and other mammals. It separates the oral cavity from the nasal cavity. A similar structure is found in crocodilians, but in most other tetrapods, the oral and nasal cavities are not truly sep ...

is not preserved, except for some incomplete vomerine teeth. The marginal teeth are not readily observable. In 2024, So, Pardo & Mann described ''Kermitops gratus'' as a new genus and species of amphibamiforms based on these fossil remains. The generic name, "''Kermitops''", combines a reference to

Kermit the Frog Kermit the Frog is a Muppet character created in 1955 and originally performed by Jim Henson. An anthropomorphic green frog, Kermit is the pragmatic everyman protagonist of numerous Muppet productions, most notably as the showrunner and host o ...

—the famous amphibian character from

the Muppets The Muppets are an American ensemble cast of puppet characters known for an surreal humor, absurdist, slapstick, burlesque, and self-referential humor, self-referential style of Musical theatre, musical Variety show, variety-sketch comedy. Cre ...

—with the

Greek Greek may refer to: Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group *Greek language, a branch of the Indo-European language family **Proto-Greek language, the assumed last common ancestor of all kno ...

suffix "-ops", meaning face. The

specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...

, "''gratus''", means "gratitude" in

Latin Latin ( or ) is a classical language belonging to the Italic languages, Italic branch of the Indo-European languages. Latin was originally spoken by the Latins (Italic tribe), Latins in Latium (now known as Lazio), the lower Tiber area aroun ...

, honoring Nicholas Hotton III and the others who aided in collecting the holotype specimen.

Classification

In their 2024 description, So, Pardo & Mann performed multiple

phylogenetic analyses In biology, phylogenetics () is the study of the evolutionary history of life using observable characteristics of organisms (or genes), which is known as Computational phylogenetics, phylogenetic inference. It infers the relationship among organ ...

; the fossil skull preserves an ossified element that may represent the

basioccipital The basilar part of the occipital bone (also basioccipital) extends forward and upward from the foramen magnum, and presents in front an area more or less quadrilateral in outline. In the young skull, this area is rough and uneven, and is joined ...

, and the coding of this character as "present" or "absent" altered their results. In their Bayesian inference analyses, ''Kermitops'' was consistently recovered as the

sister taxon In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an evolutionary tree. Definition The expression is most easily illustrated by a cladogram: Taxon A and ...

to ''

Plemmyradytes ''Plemmyradytes'' is an extinct genus of dissorophoid temnospondyl from the early Permian (early Asselian stage). It is an amphibamiform from the Eskridge Formation exposures of Nebraska. The type species is ''Plemmyradytes shintoni''. The gen ...

''. The

cladogram A cladogram (from Greek language, Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an Phylogenetic tree, evolutionary tree because it does not s ...

with the basioccipital scored as "present" is displayed below:

References

{{The Muppets Temnospondyli Permian temnospondyls of North America Permian geology of Texas Prehistoric amphibian genera Fossil taxa described in 2024 Amphibians described in 2024