Kerangas Forest
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The Sundaland heath forests, also known as ''Kerangas'' forest, is a type of tropical moist forest found on the island of
Borneo Borneo () is the List of islands by area, third-largest island in the world, with an area of , and population of 23,053,723 (2020 national censuses). Situated at the geographic centre of Maritime Southeast Asia, it is one of the Greater Sunda ...
, which is divided between
Brunei Brunei, officially Brunei Darussalam, is a country in Southeast Asia, situated on the northern coast of the island of Borneo. Apart from its coastline on the South China Sea, it is completely surrounded by the Malaysian state of Sarawak, with ...
,
Indonesia Indonesia, officially the Republic of Indonesia, is a country in Southeast Asia and Oceania, between the Indian Ocean, Indian and Pacific Ocean, Pacific oceans. Comprising over List of islands of Indonesia, 17,000 islands, including Sumatra, ...
, and
Malaysia Malaysia is a country in Southeast Asia. Featuring the Tanjung Piai, southernmost point of continental Eurasia, it is a federation, federal constitutional monarchy consisting of States and federal territories of Malaysia, 13 states and thre ...
, as well as on the Indonesian islands of Belitung and Bangka, which lie to the west of Borneo.


Setting

The word ''Kerangas'', which means "land which cannot grow rice", comes from the Iban language. Heath forests occur on acidic sandy soils that are the result of the area's siliceous parent rocks. Permanently waterlogged heath forests are known as ''kerapah'' forests.Wikramanayake, Eric; Eric Dinerstein; Colby J. Loucks; et al. (2002). ''Terrestrial Ecoregions of the Indo-Pacific: a Conservation Assessment.'' Island Press;
Washington, DC Washington, D.C., formally the District of Columbia and commonly known as Washington or D.C., is the capital city and Federal district of the United States, federal district of the United States. The city is on the Potomac River, across from ...
.
Open-canopied woodlands are known as ''padang''.Hazimah Din, Faizah Metali, Rahayu Sukmaria Sukri, "Tree Diversity and Community Composition of the Tutong White Sands, Brunei Darussalam: A Rare Tropical Heath Forest Ecosystem", ''International Journal of Ecology'', vol. 2015, Article ID 807876, 10 pages, 2015. https://doi.org/10.1155/2015/807876 The sandy soil of the heath forest are often lacking in nutrients; it is generally considered that
nitrogen Nitrogen is a chemical element; it has Symbol (chemistry), symbol N and atomic number 7. Nitrogen is a Nonmetal (chemistry), nonmetal and the lightest member of pnictogen, group 15 of the periodic table, often called the Pnictogen, pnictogens. ...
is the nutrient which is most lacking for plant growth in these forests. This is in contrast to many other lowland rain forests where
phosphorus Phosphorus is a chemical element; it has Chemical symbol, symbol P and atomic number 15. All elemental forms of phosphorus are highly Reactivity (chemistry), reactive and are therefore never found in nature. They can nevertheless be prepared ar ...
is considered to be lacking. A more recent hypothesis, proposed by Proctor (1999), is that these forests are growing on soils which are highly acidic, such that hydrogen ion toxicity prevents the growth of non-adapted species. Moreover, heath forests' low soil pH hampers organic matter decomposition thus further slowing nutrient cycling.


Flora

The Sundaland heath forests are distinct from the surrounding Borneo lowland rain forests in species composition, structure, texture, and color. The heath forests typically have a low, uniform canopy approximately 20 metres high, with thick underbrush and rich growth of moss and
epiphyte An epiphyte is a plant or plant-like organism that grows on the surface of another plant and derives its moisture and nutrients from the air, rain, water (in marine environments) or from debris accumulating around it. The plants on which epiphyt ...
s. Trees often have closely-spaced trunks. In contrast, ''padang'' woodland is relatively open, with shrubs and trees up 5 metres tall, and a ground layer of sparse grasses and sedges. Leaf sizes are generally smaller than other lowland rain forest trees. As an adaptation to the nutrient-poor conditions, some heath forest species have small, hard, thick sclerophyll leaves which are low in nitrogen. Many tree and plant species in the nutrient-deprived heath forests have developed unconventional ways to get their nutrients. Some tree species ('' Gymnostoma nobile'', for example) utilise rhizobia ( nitrogen fixing
bacteria Bacteria (; : bacterium) are ubiquitous, mostly free-living organisms often consisting of one Cell (biology), biological cell. They constitute a large domain (biology), domain of Prokaryote, prokaryotic microorganisms. Typically a few micr ...
) in their root nodules. Myrmecophytes, including ''
Myrmecodia ''Myrmecodia'' is a genus of Epiphyte, epiphytic plants, present in Indochina, Malesia, Papuasia, and Queensland, Australia. It is one of five ant-plant genera in the family Rubiaceae, the others being ''Anthorrhiza'', ''Hydnophytum'', ''Myrme ...
'' spp. and '' Hydnophytum'' spp., are tree species that develop symbiotic associations with ants to get their nutrients. Other plants, including pitcher plants ''( Nepenthes'' spp.), sundews ''( Drosera'' spp.), and bladderwort ''(Utricularia'' spp.), are carnivorous, trapping and digesting insects. Dipterocarps, including species of '' Shorea'' and '' Hopea'', are common canopy trees on less barren soils, along with palms. Native trees include both heath forest endemics, typical lowland forest species like '' Cocos nucifera'', '' Quercus acutissima'', '' Agathis borneensis, Cotylelobium burckii, Dipterocarpus borneensis,
Madhuca curtisii ''Madhuca curtisii'' is a tree in the family Sapotaceae. Description ''Madhuca curtisii'' grows up to tall, with a trunk diameter of up to . The bark is reddish brown. Inflorescences bear up to 12 flowers, which are fragrant and greenish cream- ...
, Hopea vacciniifolia, Koompassia malaccensis, Shorea ovata, Shorea scabrida, Richetia faguetiana'', and species shared with peat swamp forests like '' Shorea albida, Shorea pachyphylla'', '' Shorea scabrida'', and '' Nypa fruticans''. Heath forests are also characterized by many plants of Australasian origin, including trees of families
Myrtaceae Myrtaceae (), the myrtle family, is a family of dicotyledonous plants placed within the order Myrtales. Myrtle, pōhutukawa, bay rum tree, clove, guava, acca (feijoa), allspice, and eucalyptus are some notable members of this group. All ...
and Casuarinaceae and the southern hemisphere
conifer Conifers () are a group of conifer cone, cone-bearing Spermatophyte, seed plants, a subset of gymnosperms. Scientifically, they make up the phylum, division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a sin ...
s '' Agathis'', ''
Podocarpus ''Podocarpus'' () is a genus of conifers, the most numerous and widely distributed of the podocarp family, the Podocarpaceae. ''Podocarpus'' species are evergreen shrubs or trees, usually from tall, known to reach at times. The cones have ...
'', and '' Dacrydium''.
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Fauna

The heath forests generally have less wildlife and species diversity than the surrounding lowland rain forests, with little or no endemic species.


Protected areas

9.695% of the ecoregion is in protected areas. These include:Sundaland heath forests
DOPA Explorer. Accessed 1 April 2023.
* Sungai Serudong Forest Reserve * Bukit Soeharto Grand Forest Park * Gunung Lalang Grand Forest Park * Gunung Menumbing Grand Forest Park * Gunung Mangkol Grand Forest Park * Kutai National Park * Tanjung Puting National Park * Betung Kerihun National Park * Danau Sentarum National Park * Sebangau National Park * Meratus National Park * Gunung Maras National Park * Dered Krian National Park * Jering Menduyung Nature Recreation Park * Bukit Tangkiling Nature Recreation Park * Bukit Tangkiling Nature Reserve * Padang Luway Nature Reserve * Muara Kaman Sedulang Nature Reserve * Teluk Kelumpang, Selat Laut dan Selat Sebuku Nature Reserve


References


External links

* {{Borneo Borneo Ecoregions of Asia Ecoregions of Indonesia Ecoregions of Malaysia Ecoregions of Malesia * * Indomalayan ecoregions Tropical and subtropical moist broadleaf forests * Sclerophyll forests