''Karanisia'' is an extinct genus of

strepsirrhine Strepsirrhini or Strepsirhini (; ) is a suborder of primates that includes the lemuriform primates, which consist of the lemurs of Madagascar, galagos ("bushbabies") and pottos from Africa, and the lorises from India and Southeast Asia. Colle ...

primate Primates is an order (biology), order of mammals, which is further divided into the Strepsirrhini, strepsirrhines, which include lemurs, galagos, and Lorisidae, lorisids; and the Haplorhini, haplorhines, which include Tarsiiformes, tarsiers a ...

from middle

Eocene The Eocene ( ) is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (Ma). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene'' comes ...

deposits in

Egypt Egypt ( , ), officially the Arab Republic of Egypt, is a country spanning the Northeast Africa, northeast corner of Africa and Western Asia, southwest corner of Asia via the Sinai Peninsula. It is bordered by the Mediterranean Sea to northe ...

Classification

Two species are known, ''K. clarki'' and ''K. arenula''. Originally considered a

crown A crown is a traditional form of head adornment, or hat, worn by monarchs as a symbol of their power and dignity. A crown is often, by extension, a symbol of the monarch's government or items endorsed by it. The word itself is used, parti ...

lorisid, more comprehensive

phylogenetic analyses In biology, phylogenetics () is the study of the evolutionary history of life using observable characteristics of organisms (or genes), which is known as Computational phylogenetics, phylogenetic inference. It infers the relationship among organ ...

suggest it is a more basal to crown lorisiformes.Gregg F. Gunnell; Doug M. Boyer; Anthony R. Friscia; Steven Heritage; Fredrick Kyalo Manthi; Ellen R. Miller; Hesham M. Sallam; Nancy B. Simmons; Nancy J. Stevens; Erik R. Seiffert (2018). "Fossil lemurs from Egypt and Kenya suggest an African origin for Madagascar's aye-aye". Nature Communications. 9: Article number 3193. doi:10.1038/s41467-018-05648-w. ''K. clarki'' was described in 2003 from isolated teeth and jaw fragments found in Late Middle

(''c.'' 40 million years ago) sediments of the Birket Qarun Formation in the Egyptian

Faiyum Faiyum ( ; , ) is a city in Middle Egypt. Located southwest of Cairo, in the Faiyum Oasis, it is the capital of the modern Faiyum Governorate. It is one of Egypt's oldest cities due to its strategic location. Name and etymology Originally f ...

. The specimens indicate the presence of a

toothcomb A toothcomb (also called a tooth comb or dental comb) is a Dentition, dental structure found in some mammals, comprising a group of front teeth arranged in a manner that facilitates grooming, similar to a hair comb. The toothcomb occurs in Lem ...

, making it the earliest fossil primate to indisputably bear this trait, which is unique to all living strepsirrhines (

lemur Lemurs ( ; from Latin ) are Strepsirrhini, wet-nosed primates of the Superfamily (biology), superfamily Lemuroidea ( ), divided into 8 Family (biology), families and consisting of 15 genera and around 100 existing species. They are Endemism, ...

loris Loris is the common name for the strepsirrhine mammals of the subfamily Lorinae (sometimes spelled Lorisinae) in the family Lorisidae. ''Loris'' is one genus in this subfamily and includes the slender lorises, ''Nycticebus'' is the genus cont ...

es, and

galago Galagos , also known as bush babies or ''nagapies'' (meaning "night monkeys" in Afrikaans), are small nocturnal primates native to continental, sub-Sahara Africa, and make up the family Galagidae (also sometimes called Galagonidae). They are ...

s). In 2010 a second species, ''K. arenula'', was described in the journal ''

Nature Nature is an inherent character or constitution, particularly of the Ecosphere (planetary), ecosphere or the universe as a whole. In this general sense nature refers to the Scientific law, laws, elements and phenomenon, phenomena of the physic ...

'' from Late Middle

rocks in

Libya Libya, officially the State of Libya, is a country in the Maghreb region of North Africa. It borders the Mediterranean Sea to the north, Egypt to Egypt–Libya border, the east, Sudan to Libya–Sudan border, the southeast, Chad to Chad–L ...

References

{{Taxonbar, from=Q3130075, from2=Q20718523 Prehistoric strepsirrhines Eocene primates Prehistoric primate genera Fossil taxa described in 2003 Eocene mammals of Africa