''Jushatyria'' is an

extinct Extinction is the termination of an organism by the death of its Endling, last member. A taxon may become Functional extinction, functionally extinct before the death of its last member if it loses the capacity to Reproduction, reproduce and ...

genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...

archosaur Archosauria () or archosaurs () is a clade of diapsid sauropsid tetrapods, with birds and crocodilians being the only extant taxon, extant representatives. Although broadly classified as reptiles, which traditionally exclude birds, the cladistics ...

Fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...

s have been found in the Koltaevo III Locality, district of

Kumertau Kumertau (; , ''Kümertaw'') is a town in the Republic of Bashkortostan, Russia, located from Ufa and from Sterlitamak. Population: Administrative and municipal status Within the framework of administrative divisions, it is, together with ...

near the

Ural Mountains The Ural Mountains ( ),; , ; , or simply the Urals, are a mountain range in Eurasia that runs north–south mostly through Russia, from the coast of the Arctic Ocean to the river Ural (river), Ural and northwestern Kazakhstan.

European Russia European Russia is the western and most populated part of the Russia, Russian Federation. It is geographically situated in Europe, as opposed to the country's sparsely populated and vastly larger eastern part, Siberia, which is situated in Asia ...

from the Bukobay Gorizont. The locality dates back to the

Ladinian The Ladinian is a stage and age in the Middle Triassic series or epoch. It spans the time between Ma and ~237 Ma (million years ago). The Ladinian was preceded by the Anisian and succeeded by the Carnian (part of the Upper or Late Triassic ...

stage of the

Middle Triassic In the geologic timescale, the Middle Triassic is the second of three epoch (geology), epochs of the Triassic period (geology), period or the middle of three series (stratigraphy), series in which the Triassic system (stratigraphy), system is di ...

. Additional material has been described from a locality on the banks of the Berdyanka River that was previously assigned to a rauisuchid-like archosaur. However, this material differed from the original specimens because it lacked slit-like antorbital openings accompanying the antorbital fossa. Nesbitt (2009) and Gower and Sennikov (2000) suggested that all material currently referred to ''Jushatyria'' most likely does not represent a single taxon.Gower, D. J. and Sennikov, A. G. (2000). Early Archosaurs from Russia ''In:'' Benton, M. J., Kurochkin, E. N., Shishkin, M. A. and Unwin, D. M., eds., ''The Age of Dinosaurs in Russia and Mongolia''. Cambridge, Cambridge University Press; pp. 140–159. Thus, ''Jushatyria'' is known only from its

holotype A holotype (Latin: ''holotypus'') is a single physical example (or illustration) of an organism used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of s ...

PIN 2867/5, an incomplete left

maxilla In vertebrates, the maxilla (: maxillae ) is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. The two maxil ...

. As the maxilla is damaged, many "

rauisuchia "Rauisuchia" is a paraphyletic group of mostly large and carnivorous Triassic archosaurs. Rauisuchians are a category of archosaurs within a larger group called Pseudosuchia, which encompasses all archosaurs more closely related to crocodilians ...

n" characters could not be verified. ''Jushatyria'' was reassigned as an indeterminate archosaur on the basis of the presence of an antorbital fossa on the lateral surface of the maxilla.

References

External links

''Jushatyria''
in the

Paleobiology Database The Paleobiology Database (PBDB) is an online resource for information on the distribution and classification of fossil animals, plants, and microorganisms. History The Paleobiology Database originated in the NCEAS-funded Phanerozoic Marine Pale ...

Rauisuchids Extinct animals of Europe Prehistoric pseudosuchian genera {{triassic-reptile-stub