John S. Clark
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John S. Clark (21 March 1885 – 1 June 1956) was a
Scottish Scottish usually refers to something of, from, or related to Scotland, including: *Scottish Gaelic, a Celtic Goidelic language of the Indo-European language family native to Scotland *Scottish English *Scottish national identity, the Scottish ide ...
-born Australian
entomologist Entomology (from Ancient Greek ἔντομον (''éntomon''), meaning "insect", and -logy from λόγος (''lógos''), meaning "study") is the branch of zoology that focuses on insects. Those who study entomology are known as entomologists. In ...
and
myrmecologist Myrmecology (; from Greek: μύρμηξ, ''myrmex'', "ant" and λόγος, ''logos'', "study") is a branch of entomology focusing on the study of ants. Ants continue to be a model of choice for the study of questions on the evolution of social ...
known for his study of Australian
ant Ants are Eusociality, eusocial insects of the Family (biology), family Formicidae and, along with the related wasps and bees, belong to the Taxonomy (biology), order Hymenoptera. Ants evolved from Vespoidea, vespoid wasp ancestors in the Cre ...
s. Born in
Glasgow Glasgow is the Cities of Scotland, most populous city in Scotland, located on the banks of the River Clyde in Strathclyde, west central Scotland. It is the List of cities in the United Kingdom, third-most-populous city in the United Kingdom ...
, he developed an interest in entomology at a young age. Clark first arrived in Australia in 1905 and originally worked for the state railways in
Queensland Queensland ( , commonly abbreviated as Qld) is a States and territories of Australia, state in northeastern Australia, and is the second-largest and third-most populous state in Australia. It is bordered by the Northern Territory, South Austr ...
. He developed an interest in ants shortly afterwards, collecting his first specimens in North Queensland. He married his first wife, Maggie Forbes in 1908, who bore four children, and died in 1935. He married his second wife, Phyllis Marjorie Claringbulls in 1939 and had two daughters with her. On her suicide in 1943, Clark sent his daughters to an orphanage. In 1919, Clark worked as the assistant to the entomologist on probation in the Western Australian Department of Agriculture, but in 1920, he took on this position full-time. He started to publish his first articles about pest insects and ants from 1921; in 1926 he became an entomologist at the
National Museum A national museum can be a museum maintained and funded by a national government. In many countries it denotes a museum run by the central government, while other museums are run by regional or local governments. In the United States, most nati ...
in
Melbourne Melbourne ( , ; Boonwurrung language, Boonwurrung/ or ) is the List of Australian capital cities, capital and List of cities in Australia by population, most populous city of the States and territories of Australia, Australian state of Victori ...
, remaining there for 20 years. Clark continued to publish more articles until he resigned from the National Museum of
Victoria Victoria most commonly refers to: * Queen Victoria (1819–1901), Queen of the United Kingdom and Empress of India * Victoria (state), a state of Australia * Victoria, British Columbia, Canada, a provincial capital * Victoria, Seychelles, the capi ...
in 1944. Living in poverty, Clark lived a reclusive life, publishing his last book in 1951. He died on 1 June 1956 at the age of 71. One of Clark's most notable achievements was describing '' Nothomyrmecia macrops'', the most primitive living ant. Several ants have been named after him in recognition of his contributions.


Early life

John S. Clark was born on 21 March 1885 in
Glasgow Glasgow is the Cities of Scotland, most populous city in Scotland, located on the banks of the River Clyde in Strathclyde, west central Scotland. It is the List of cities in the United Kingdom, third-most-populous city in the United Kingdom ...
, Scotland, to parents James Souttar Clark, a coach-painter, and Maggie Clark (née Scott). At an early age, Clark had an interest in entomology throughout his years in Glasgow and adult life in Australia. Clark migrated to Australia in 1905 with little formal education but found himself working for the state railways in
Queensland Queensland ( , commonly abbreviated as Qld) is a States and territories of Australia, state in northeastern Australia, and is the second-largest and third-most populous state in Australia. It is bordered by the Northern Territory, South Austr ...
. There, he found an interest in ants and made his first collection of specimens in North Queensland. In May 1908, Clark married his first wife, Maggie Forbes, at the
Cairns Cairns (; ) is a city in the Cairns Region, Queensland, Australia, on the tropical north east coast of Far North Queensland. In the , Cairns had a population of 153,181 people. The city was founded in 1876 and named after William Cairns, Sir W ...
Presbyterian Church. After their marriage, they moved to
Geraldton Geraldton (Wajarri language, Wajarri: ''Jambinu'', Wilunyu language, Wilunyu: ''Jambinbirri'') is a coastal city in the Mid West (Western Australia), Mid West region of Western Australia, north of the state capital, Perth. As of the , Geraldt ...
in Western Australia, where Clark worked for the railways as a wheelwright. He had one son and three daughters with Forbes; Forbes later died from heart disease in 1935.


Career

Australian entomologist Leslie John William Newman of the Western Australian Department of Agriculture noticed Clark's enthusiasm for natural history, and on probation, took Clark in as the assistant to the entomologist in 1919. In 1920, Clark's position in the Department of Agriculture was confirmed; due to this, Clark and his family moved to
Perth Perth () is the list of Australian capital cities, capital city of Western Australia. It is the list of cities in Australia by population, fourth-most-populous city in Australia, with a population of over 2.3 million within Greater Perth . The ...
. He began publishing his first papers discussing the history of entomology in Western Australia in 1921, and also basic articles about ants, insect pests in Australian forests and
myrmecophilous thumb , Myrmecophilous aphids being tended by ants Myrmecophily ( , ) consists of positive, mutualistic, interspecies associations between ants and a variety of other organisms, such as plants, other arthropods, and fungi. It may also incl ...
beetles. Three years after his articles were first published, Clark was promoted to assistant-entomologist. Scientists attending the 1926 Australasian Association for the Advancement of Science suggested that Clark should apply for a position as "entomologist" at the
National Museum of Victoria National may refer to: Common uses * Nation or country ** Nationality – a ''national'' is a person who is subject to a nation, regardless of whether the person has full rights as a citizen Places in the United States * National, Maryland, ...
, in which he was appointed in late 1926 and began working there in 1927. After working for the museum, he found museum work "not to his liking" and tried to apply for the post of "economic entomologist" in
New Guinea New Guinea (; Hiri Motu: ''Niu Gini''; , fossilized , also known as Papua or historically ) is the List of islands by area, world's second-largest island, with an area of . Located in Melanesia in the southwestern Pacific Ocean, the island is ...
, which at the time was an Australian administered-territory. However, this was unsuccessful for Clark, who remained at the National Museum of
Victoria Victoria most commonly refers to: * Queen Victoria (1819–1901), Queen of the United Kingdom and Empress of India * Victoria (state), a state of Australia * Victoria, British Columbia, Canada, a provincial capital * Victoria, Seychelles, the capi ...
for nearly 20 years. In 1933, Clark sold 8,000 ant specimens he collected to the museum for about £200. Following the death of his first wife in 1935, Clark moved from Hawthorn to Ferntree Gully in Melbourne. A couple of years later in 1939, he married his second wife, Phyllis Marjorie Claringbull, at the Office of the Government Statist, bearing two more daughters. Claringbull committed suicide in 1943, three months after she gave birth to her second daughter; Clark later sent them to an orphanage as he was unable to look after them. Clark's intemperate attitude to his peers and superiors and lack of qualifications bedevilled his career, although he was welcoming to amateurs. In 1944, Clark resigned from the National Museum of Victoria after Australian botanist Richard Pescott became the director of the museum. As a result, he moved to Mooroolbark and lived in poverty, losing all of his entitlements. Due to his reputable view as an authority on ants, Clark worked on a monograph of ants in Australia, supported by the
Commonwealth Scientific and Industrial Research Organisation The Commonwealth Scientific and Industrial Research Organisation (CSIRO) is an Australian Government agency that is responsible for scientific research and its commercial and industrial applications. CSIRO works with leading organisations arou ...
through grants. His first volume was published in 1951, which covered the Australian bulldog ants (subfamily
Myrmeciinae Myrmeciinae is a subfamily of the Formicidae, ants once found worldwide but now restricted to Australia and New Caledonia. This subfamily is one of several ant subfamilies which possess gamergates, female worker ants which are able to mate an ...
). However, the first volume received poor reviews, and no further volume was released. In fact, it is unknown whether or not any further volume will be released.


Research

Aside from some papers published discussing myrmecophilous beetles and insect pests in forests, Clark predominately researched and published articles about the taxonomy of Australian ants. All of his articles and monographs were published in Australia. Throughout his career, Clark described around 200 different species of ants, but half of these may be
synonyms A synonym is a word, morpheme, or phrase that means precisely or nearly the same as another word, morpheme, or phrase in a given language. For example, in the English language, the words ''begin'', ''start'', ''commence'', and ''initiate'' are a ...
. He worked and researched with all of the main groups of Australian ants, but his research and revisionary efforts were extensive with members of the former subfamily Cerapachyinae (now
Dorylinae Dorylinae is an ant subfamily, with distributions in both the Old World and New World. Brady ''et al.'' (2014) Synonym (taxonomy), synonymized the previous dorylomorph subfamilies (Aenictinae, Aenictogitoninae, Cerapachyinae, Ecitoninae, and Lep ...
), especially those of '' Phyracaces'', the Myrmeciinae, '' Rhytidoponera'' and members of the tribe
Dolichoderini ''Dolichoderus'' is a genus of ants found worldwide. Taxonomy The ants of the Neotropical genus ''Monacis'' were revised in 1959 by Kempf. However, Brown in 1973 and G. C. Wheeler and J. Wheeler in 1973 and 1976 considered both ''Monacis'' and ' ...
. Clark is most notable for describing the dinosaur ant ('' Nothomyrmecia macrops''), which is the most primitive living ant in the world and second most primitive when the fossil record is included. The first collection of ''Nothomyrmecia'' was made in December 1931 by Amy Crocker who collected specimens of two worker ants, reportedly near the Russell Range from Israelite Bay in Western Australia. Crocker sent the two specimens to Clark at the museum for study; in 1934, Clark published a formal description of ''Nothomyrmecia'' as a completely new genus of the Myrmeciinae. He did so because the two specimens (which then became the
syntype In biological nomenclature, a syntype is any one of two or more biological types that is listed in a description of a taxon where no holotype was designated. Precise definitions of this and related terms for types have been established as part o ...
s) bore no resemblance to any ant species he knew of, although they did share similar morphological characteristics with the extinct genus '' Prionomyrmex''. This unusual ant remained unknown to scientists, causing intense scientific interest in the early 1950s. Over three decades, however, teams of Australian and American collectors failed to re-find it after they initiated a series of searches. Then, in 1977, entomologist Robert Taylor and his party of entomologists from Canberra found a solitary worker ant at Poochera, southeast of Ceduna, some from the reported site of the 1931 discovery. After 46 years of searching for it, entomologists have dubbed the ant the "
Holy Grail The Holy Grail (, , , ) is a treasure that serves as an important motif in Arthurian literature. Various traditions describe the Holy Grail as a cup, dish, or stone with miraculous healing powers, sometimes providing eternal youth or sustenanc ...
" of
myrmecology Myrmecology (; from Greek: μύρμηξ, ''myrmex'', "ant" and λόγος, ''logos'', "study") is a branch of entomology focusing on the study of ants. Ants continue to be a model of choice for the study of questions on the evolution of social ...
. Such discovery of ''Nothomyrmecia'' and its general nature marks it as one of Clark's most notable achievements.


Death and recognition

Clark died at his Mooroolbark home on 1 June 1956, at the age of 71. American entomologist William Brown Jr. notes that Clark was living the life of a "
recluse A recluse is a person who lives in voluntary seclusion and solitude. The word is from the Latin , which means 'to open' or 'disclose'. Examples of recluses are Symeon of Trier, who lived within the great Roman gate Porta Nigra with permissio ...
" during his declining years. Clark was buried in
Burwood Cemetery Burwood Cemetery is a cemetery in Burwood, Victoria in Australia. It dates back to 1858, and was originally known as Nunawading General Cemetery. It is known as a resting place of notable figures from Melbourne. The site is operated by Greater Me ...
. At the time of his death, the majority of his ant collection were housed in the National Museum of Victoria, the
Natural History Museum A natural history museum or museum of natural history is a scientific institution with natural history scientific collection, collections that include current and historical records of animals, plants, Fungus, fungi, ecosystems, geology, paleo ...
in
London London is the Capital city, capital and List of urban areas in the United Kingdom, largest city of both England and the United Kingdom, with a population of in . London metropolitan area, Its wider metropolitan area is the largest in Wester ...
and the
Museum of Comparative Zoology The Museum of Comparative Zoology (formally the Agassiz Museum of Comparative Zoology and often abbreviated to MCZ) is a zoology museum located on the grounds of Harvard University in Cambridge, Massachusetts. It is one of three natural-history r ...
in
Cambridge Cambridge ( ) is a List of cities in the United Kingdom, city and non-metropolitan district in the county of Cambridgeshire, England. It is the county town of Cambridgeshire and is located on the River Cam, north of London. As of the 2021 Unit ...
. One of Clark's daughters, Ellen Clark, was a renowned naturalist. She had worked with her father at the museum in 1940 and was the secretary of the virus department of the
Walter and Eliza Hall Institute of Medical Research Walter may refer to: People and fictional characters * Walter (name), including a list of people and fictional and mythical characters with the given name or surname * Little Walter, American blues harmonica player Marion Walter Jacobs (1930–1 ...
. As well as that, she worked for the Argus and Australasian and published papers on influenza virus research and
crustacean Crustaceans (from Latin meaning: "those with shells" or "crusted ones") are invertebrate animals that constitute one group of arthropods that are traditionally a part of the subphylum Crustacea (), a large, diverse group of mainly aquatic arthrop ...
s. A number of ants have been named after Clark. The
holotype A holotype (Latin: ''holotypus'') is a single physical example (or illustration) of an organism used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of s ...
of '' Polyrhachis clarki'' was originally collected by Clark, but it was not described until 2013. There, the author of the article named the ant after him. '' Leptogenys clarki'' is another ant named after Clark for his contributions towards Australian ants, being described by American entomologist
William Morton Wheeler William Morton Wheeler (March 19, 1865 – April 19, 1937) was an American entomologist, myrmecologist and professor at Harvard University. Biography Early life and education William Morton Wheeler was born on March 19, 1865, to parents Juliu ...
in 1933. Ants he collected that bear his name include '' Dolichoderus clarki'', '' Plagiolepis clarki'' and '' Stigmacros clarki''.


Published works

Over the course of his career, Clark published over 35 entomological papers. The following publications are found at the Hymenoptera Online Database: Books * Journals * * * * * * * * * * * * * * * * * * * * * *


Notes


References


External links


John S. Clark at the AntWiki – Bringing Ants to the World
* {{DEFAULTSORT:Clark, John S. 1885 births 1956 deaths Scientists from Glasgow Australian entomologists Scottish entomologists Myrmecologists Scottish emigrants to Australia 20th-century Australian zoologists