Jenny Clack
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Jennifer Alice Clack, (''née'' Agnew; 3 November 1947 – 26 March 2020) was an English
palaeontologist Paleontology, also spelled as palaeontology or palæontology, is the scientific study of the life of the past, mainly but not exclusively through the study of fossils. Paleontologists use fossils as a means to classify organisms, measure geolo ...
and
evolutionary biologist Evolutionary biology is the subfield of biology that studies the evolutionary processes such as natural selection, common descent, and speciation that produced the diversity of life on Earth. In the 1930s, the discipline of evolutionary biol ...
. She specialised in the early evolution of
tetrapod A tetrapod (; from Ancient Greek :wiktionary:τετρα-#Ancient Greek, τετρα- ''(tetra-)'' 'four' and :wiktionary:πούς#Ancient Greek, πούς ''(poús)'' 'foot') is any four-Limb (anatomy), limbed vertebrate animal of the clade Tetr ...
s, specifically studying the "fish to tetrapod" transition: the origin, evolutionary development and radiation of early tetrapods and their relatives among the
lobe-finned fish Sarcopterygii (; )—sometimes considered synonymous with Crossopterygii ()—is a clade (traditionally a class or subclass) of vertebrate animals which includes a group of bony fish commonly referred to as lobe-finned fish. These vertebrates ar ...
es. She is best known for her book ''Gaining Ground: the Origin and Early Evolution of Tetrapods'', published in 2002 (second edition, 2012) and written with the layperson in mind. Clack was curator at the
Museum of Zoology The Museum of Zoology () of National Taiwan University (NTU) is a museum about zoology at the NTU main campus in Da'an District, Taipei, Taiwan. History The museum was originally established in 1928 during the Japanese rule. In the early days ...
and
Professor Professor (commonly abbreviated as Prof.) is an Academy, academic rank at university, universities and other tertiary education, post-secondary education and research institutions in most countries. Literally, ''professor'' derives from Latin ...
of Vertebrate Palaeontology at the
University of Cambridge The University of Cambridge is a Public university, public collegiate university, collegiate research university in Cambridge, England. Founded in 1209, the University of Cambridge is the List of oldest universities in continuous operation, wo ...
, where she devoted her career to studying the early development of tetrapods, the "four-legged" animals said to have evolved from
Devonian The Devonian ( ) is a period (geology), geologic period and system (stratigraphy), system of the Paleozoic era (geology), era during the Phanerozoic eon (geology), eon, spanning 60.3 million years from the end of the preceding Silurian per ...
lobe-finned fishes and colonised the freshwater swamps of the
Carboniferous The Carboniferous ( ) is a Geologic time scale, geologic period and System (stratigraphy), system of the Paleozoic era (geology), era that spans 60 million years, from the end of the Devonian Period Ma (million years ago) to the beginning of the ...
period.


Early life and education

Clack was born on 3 November 1947, the only child of Ernest and Alice Agnew. She was brought up in
Manchester Manchester () is a city and the metropolitan borough of Greater Manchester, England. It had an estimated population of in . Greater Manchester is the third-most populous metropolitan area in the United Kingdom, with a population of 2.92&nbs ...
, England. She was educated at
Bolton School Bolton School is a private day school in Bolton, Greater Manchester. It comprises a co-educational nursery (ages 0–4), co-educational infant school (ages 3–7), single sex junior schools (ages 7–11) and single sex senior schools including ...
(Girls' Division), a
private school A private school or independent school is a school not administered or funded by the government, unlike a State school, public school. Private schools are schools that are not dependent upon national or local government to finance their fina ...
in
Bolton Bolton ( , locally ) is a town in Greater Manchester in England. In the foothills of the West Pennine Moors, Bolton is between Manchester, Blackburn, Wigan, Bury, Greater Manchester, Bury and Salford. It is surrounded by several towns and vill ...
,
Lancashire Lancashire ( , ; abbreviated ''Lancs'') is a ceremonial county in North West England. It is bordered by Cumbria to the north, North Yorkshire and West Yorkshire to the east, Greater Manchester and Merseyside to the south, and the Irish Sea to ...
. She received a B.Sc. in Zoology from the
University of Newcastle upon Tyne Newcastle University (legally the University of Newcastle upon Tyne) is a public university, public research university based in Newcastle upon Tyne, England. It has overseas campuses in Singapore and Malaysia. The university is a red brick un ...
in 1970. In 1978, she accepted an invitation from Alec Panchen to obtain a Ph.D. from the same university, at the encouragement of her then new boyfriend Rob Clack. They married in 1980. Her doctorate was completed in 1984. She also held a Graduate Certificate in Museum Studies from the
University of Leicester The University of Leicester ( ) is a public university, public research university based in Leicester, England. The main campus is south of the city centre, adjacent to Victoria Park, Leicester, Victoria Park. The university's predecessor, Univ ...
and an MA from the
University of Cambridge The University of Cambridge is a Public university, public collegiate university, collegiate research university in Cambridge, England. Founded in 1209, the University of Cambridge is the List of oldest universities in continuous operation, wo ...
. On 9 December 2000, she was awarded a
Doctor of Science A Doctor of Science (; most commonly abbreviated DSc or ScD) is a science doctorate awarded in a number of countries throughout the world. Africa Algeria and Morocco In Algeria, Morocco, Libya and Tunisia, all universities accredited by the s ...
(ScD) degree by the
University of Cambridge The University of Cambridge is a Public university, public collegiate university, collegiate research university in Cambridge, England. Founded in 1209, the University of Cambridge is the List of oldest universities in continuous operation, wo ...
.


Academic career

In 1981, Clack joined the University Museum of Zoology,
University of Cambridge The University of Cambridge is a Public university, public collegiate university, collegiate research university in Cambridge, England. Founded in 1209, the University of Cambridge is the List of oldest universities in continuous operation, wo ...
, as an Assistant Curator. She was promoted to Senior Assistant Curator in 1995. Since 2005, she has been Curator in Vertebrate Palaeontology at the museum. In 2006, she was awarded a
personal chair Academic ranks in the United Kingdom are the titles, relative seniority and responsibility of employees in universities. In general the country has three academic career pathways: one focused on research, one on teaching, and one that combines ...
by the University of Cambridge, and took the title Professor of Vertebrate Palaeontology. She retired in 2015 and became Emeritus Professor of Vertebrate Palaeontology at the museum. In 1997, Clack was elected a
Fellow A fellow is a title and form of address for distinguished, learned, or skilled individuals in academia, medicine, research, and industry. The exact meaning of the term differs in each field. In learned society, learned or professional society, p ...
of
Darwin College, Cambridge Darwin College is a Colleges of the University of Cambridge, constituent college of the University of Cambridge. Founded on 28 July 1964, Darwin was Cambridge University's first graduate-only college, and also the first to admit both men and wo ...
; since 1 October 2015, she has been an
Emeritus Fellow ''Emeritus/Emerita'' () is an honorary title granted to someone who retires from a position of distinction, most commonly an academic faculty position, but is allowed to continue using the previous title, as in "professor emeritus". In some ca ...
. From 2000 to 2005, she was Reader in Vertebrate Palaeontology at the
University of Cambridge The University of Cambridge is a Public university, public collegiate university, collegiate research university in Cambridge, England. Founded in 1209, the University of Cambridge is the List of oldest universities in continuous operation, wo ...
. On 1 October 2006, Clack was awarded a
personal chair Academic ranks in the United Kingdom are the titles, relative seniority and responsibility of employees in universities. In general the country has three academic career pathways: one focused on research, one on teaching, and one that combines ...
by Cambridge, taking the title Professor of Vertebrate Palaeontology. Clack is best known for her extensive body of work on early tetrapods, much of which redefined how paleontologists conceived of the evolution of limbs and other features associated with tetrapods' transition from other lobe-finned fishes. She began her career working primarily on the ear of early tetrapods and later expanded to more broadly addressing the osteology and evolution of tetrapods. Together with Michael Coates (
University of Chicago The University of Chicago (UChicago, Chicago, or UChi) is a Private university, private research university in Chicago, Illinois, United States. Its main campus is in the Hyde Park, Chicago, Hyde Park neighborhood on Chicago's South Side, Chic ...
), Clack defined what is known as "
Romer's Gap Romer's gap is an apparent gap in the Paleozoic tetrapod fossil record noted in the studies of paleontology and evolutionary biology, which represent periods in the Early Carboniferous from which excavators have not yet found relevant transition ...
", a major gap in the fossil record of early tetrapods and one that she subsequently began to fill in collaboration with other paleontologists. Clack also undertook extensive fieldwork expeditions in order to search for further remains of early tetrapods. In 1987, during an expedition to
East Greenland Tunu, in Danish Østgrønland ("East Greenland"), was one of the three counties (''amter'') of Greenland until 31 December 2008. The county seat was at the main settlement, Tasiilaq. The county's population in 2005 was around 3,800. The county ...
, Clack and her team discovered the remains of the Devonian tetrapods ''
Acanthostega ''Acanthostega'', from Ancient Greek ἄκανθα (''ákantha''), meaning "spine", and στέγη (''stégē''), meaning "roof", is an extinct genus of stem tetrapoda, stem-tetrapod, among the first vertebrates, vertebrate animals to have recogn ...
'' and ''
Ichthyostega ''Ichthyostega'', from Ancient Greek ἰχθύς (''ikthús''), meaning "fish", and στέγη (''stégē''), meaning "roof", is an Extinction, extinct genus of limbed tetrapodomorpha, tetrapodomorphs from the Devonian, Late Devonian of what is ...
,'' following up on field notes of researchers who had collected material of ''Acanthostega'' in 1970. Additional surveys in 1998 led to the collection of substantial new material, including what is now recognized as ''
Ymeria ''Ymeria'' is an extinct genus of early stem tetrapod from the Devonian of Greenland. Of the two other genera of stem tetrapods from Greenland, ''Acanthostega'' and ''Ichthyostega'', ''Ymeria'' is most closely related to ''Ichthyostega'', though ...
.'' Most recently, she led a major consortium project (TW:eed) investigating some exciting new fossils from Northumberland and the
Borders Region of Scotland The Scottish Borders is one of 32 council areas of Scotland. It is bordered by West Lothian, Edinburgh, Midlothian, and East Lothian to the north, the North Sea to the east, Dumfries and Galloway to the south-west, South Lanarkshire to the we ...
which date from the
Tournaisian The Tournaisian is in the ICS geologic timescale the lowest stage or oldest age of the Mississippian, the oldest subsystem of the Carboniferous. The Tournaisian age lasted from Ma to Ma. It is preceded by the Famennian (the uppermost st ...
stage of the earliest
Carboniferous The Carboniferous ( ) is a Geologic time scale, geologic period and System (stratigraphy), system of the Paleozoic era (geology), era that spans 60 million years, from the end of the Devonian Period Ma (million years ago) to the beginning of the ...
period; this project has produced numerous publications furthering the understanding of early tetrapod evolution. Over the course of her lengthy career, Clack published in some of the most notable scientific journals, including
Nature Nature is an inherent character or constitution, particularly of the Ecosphere (planetary), ecosphere or the universe as a whole. In this general sense nature refers to the Scientific law, laws, elements and phenomenon, phenomena of the physic ...
,
Science Science is a systematic discipline that builds and organises knowledge in the form of testable hypotheses and predictions about the universe. Modern science is typically divided into twoor threemajor branches: the natural sciences, which stu ...
, and
Proceedings of the National Academy of Sciences ''Proceedings of the National Academy of Sciences of the United States of America'' (often abbreviated ''PNAS'' or ''PNAS USA'') is a peer-reviewed multidisciplinary scientific journal. It is the official journal of the National Academy of Scie ...
and is one of the most published vertebrate paleontologists in Nature, arguably the leading scientific journal in the world, with over 15 papers in that journal alone. In addition to her ''Gaining Ground'' book, Clack also co-authored a volume of the Handbuch der Paläoherpetologie series on early tetrapods with Andrew Milner in 2015 and co-edited a volume on the evolution of hearing in 2016. Clack is probably best known for discovering that the earliest
tetrapod A tetrapod (; from Ancient Greek :wiktionary:τετρα-#Ancient Greek, τετρα- ''(tetra-)'' 'four' and :wiktionary:πούς#Ancient Greek, πούς ''(poús)'' 'foot') is any four-Limb (anatomy), limbed vertebrate animal of the clade Tetr ...
s had more than five toes per foot: the Upper Devonian tetrapods from East Greenland, ''
Ichthyostega ''Ichthyostega'', from Ancient Greek ἰχθύς (''ikthús''), meaning "fish", and στέγη (''stégē''), meaning "roof", is an Extinction, extinct genus of limbed tetrapodomorpha, tetrapodomorphs from the Devonian, Late Devonian of what is ...
'' had seven while ''
Acanthostega ''Acanthostega'', from Ancient Greek ἄκανθα (''ákantha''), meaning "spine", and στέγη (''stégē''), meaning "roof", is an extinct genus of stem tetrapoda, stem-tetrapod, among the first vertebrates, vertebrate animals to have recogn ...
'' had eight (compared to the six toes of the Russian Devonian tetrapod ''
Tulerpeton ''Tulerpeton'' is an extinct genus of Devonian four-limbed vertebrate, known from a fossil that was found in the Tula Region of Russia at a site named Andreyevka. This genus, ''Acanthostega'', and ''Ichthyostega'' represent the earliest tetrapods ...
''). This suggests that
pentadactyly In biology, dactyly is the arrangement of wikt:digit, digits (fingers and toes) on the hands, feet, or sometimes wings of a tetrapod animal. The term is derived from the Greek (language), Greek word () meaning "finger." Sometimes the suffix "- ...
was not an ancestral trait for tetrapods. Clack supervised many graduate students who went on to pursue successful careers in paleontology and evolutionary biology, including
Per Ahlberg Per Erik Ahlberg is a Sweden, Swedish palaeontologist working with the earliest tetrapods. He took his Ph.D. in zoology at the University of Cambridge in 1989 under English palaeontologist Jenny Clack. He is currently professor at the Department o ...
(
Uppsala University Uppsala University (UU) () is a public university, public research university in Uppsala, Sweden. Founded in 1477, it is the List of universities in Sweden, oldest university in Sweden and the Nordic countries still in operation. Initially fou ...
), Paul Upchurch (
University College London University College London (Trade name, branded as UCL) is a Public university, public research university in London, England. It is a Member institutions of the University of London, member institution of the Federal university, federal Uni ...
), Michael Lee (
Flinders University Flinders University, established as The Flinders University of South Australia is a public university, public research university based in Adelaide, South Australia, with a footprint extending across a number of locations in South Australia and ...
), and Matthew Friedman (
University of Michigan The University of Michigan (U-M, U of M, or Michigan) is a public university, public research university in Ann Arbor, Michigan, United States. Founded in 1817, it is the oldest institution of higher education in the state. The University of Mi ...
). In April 2012 she was featured in an episode of the BBC television series '' Beautiful Minds'', a set of documentaries about scientists who have made important discoveries. This may be viewed on YouTube. Clack was honored by her peers with a
festschrift In academia, a ''Festschrift'' (; plural, ''Festschriften'' ) is a book honoring a respected person, especially an academic, and presented during their lifetime. It generally takes the form of an edited volume, containing contributions from the h ...
published in 2019.


Death

Clack died on 26 March 2020 at the age of 72, after a five-year battle with
endometrial cancer Endometrial cancer is a cancer that arises from the endometrium (the epithelium, lining of the uterus or womb). It is the result of the abnormal growth of cells (biology), cells that can invade or spread to other parts of the body. The first s ...
.


Honours

In 2008, Clack was awarded the
Daniel Giraud Elliot Medal The Daniel Giraud Elliot Medal is awarded by the U.S. National Academy of Sciences "for meritorious work in zoology or paleontology study published in a three to five year period." Named after Daniel Giraud Elliot, it was first awarded in 1917. L ...
from the U.S.
National Academy of Sciences The National Academy of Sciences (NAS) is a United States nonprofit, NGO, non-governmental organization. NAS is part of the National Academies of Sciences, Engineering, and Medicine, along with the National Academy of Engineering (NAE) and the ...
, the first woman to achieve the honor. In 2009, Clack was elected a
Fellow of the Royal Society Fellowship of the Royal Society (FRS, ForMemRS and HonFRS) is an award granted by the Fellows of the Royal Society of London to individuals who have made a "substantial contribution to the improvement of natural science, natural knowledge, incl ...
, the first female vertebrate paleontologist to achieve the honor. She has also been elected a Foreign Honorary Member of the
American Academy of Arts and Sciences The American Academy of Arts and Sciences (The Academy) is one of the oldest learned societies in the United States. It was founded in 1780 during the American Revolution by John Adams, John Hancock, James Bowdoin, Andrew Oliver, and other ...
. On 15 June 2013, Clack was awarded an
honorary An honorary position is one given as an honor, with no duties attached, and without payment. Other uses include: * Honorary Academy Award, by the Academy of Motion Picture Arts and Sciences, United States * Honorary Aryan, a status in Nazi Germany ...
Doctor of Science A Doctor of Science (; most commonly abbreviated DSc or ScD) is a science doctorate awarded in a number of countries throughout the world. Africa Algeria and Morocco In Algeria, Morocco, Libya and Tunisia, all universities accredited by the s ...
(DSc) degree by the
University of Chicago The University of Chicago (UChicago, Chicago, or UChi) is a Private university, private research university in Chicago, Illinois, United States. Its main campus is in the Hyde Park, Chicago, Hyde Park neighborhood on Chicago's South Side, Chic ...
. The university described her as "an internationally preeminent palaeontologist whose research has profoundly changed the understanding of the origin of terrestrial vertebrate life." Also in 2013, she was awarded the T Neville George Medal by the
Geological Society of Glasgow The Geological Society of Glasgow is a scientific society devoted to the study of geology in Scotland. The society contributed to the understanding of Scotland's glacial history, and the relationship between the Earth's rotation and climate ch ...
. On 17 July 2014, she was awarded an honorary Doctor of Science degree by the
University of Leicester The University of Leicester ( ) is a public university, public research university based in Leicester, England. The main campus is south of the city centre, adjacent to Victoria Park, Leicester, Victoria Park. The university's predecessor, Univ ...
. Also in 2014, she was made an Honorary Foreign Member of the
Royal Swedish Academy of Sciences The Royal Swedish Academy of Sciences () is one of the Swedish Royal Academies, royal academies of Sweden. Founded on 2 June 1739, it is an independent, non-governmental scientific organization that takes special responsibility for promoting nat ...
. In 2018, she won the
Palaeontological Association The Palaeontological Association (PalAss for short) is a charitable organisation based in the UK founded in 1957 for the promotion of the study of palaeontology and its allied sciences. Publications The Association publishes two main journals ...
's most prestigious award, the
Lapworth Medal The Lapworth Medal is the highest award of the Palaeontological Association, given to those who have made a significant contribution to the science by means of a substantial body of research. Recipients Source Palaeontological Association*2024 - P ...
.


References


External links


Jenny Clack on the Cambridge University websiteHome page - Jennifer "Jenny" ClackInterview on PBS websiteJenny Clack, Paleontologist: The First Vertebrate Walks on Land
on Vimeo, 2012 {{DEFAULTSORT:Clack, Jenny 1947 births 2020 deaths Alumni of Newcastle University Alumni of the University of Leicester English palaeontologists British women paleontologists Fellows of Darwin College, Cambridge Female fellows of the Royal Society Scientists from Manchester People educated at Bolton School 21st-century English women scientists Members of the Royal Swedish Academy of Sciences Fellows of the American Academy of Arts and Sciences Fellows of the Royal Society