Jeholosauridae
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Jeholosauridae is an extinct group of
herbivorous A herbivore is an animal anatomically and physiologically evolved to feed on plants, especially upon vascular tissues such as foliage, fruits or seeds, as the main component of its diet. These more broadly also encompass animals that eat n ...
neornithischia Neornithischia ("new ornithischians") is a clade of the dinosaur order Ornithischia. It is the sister group of the Thyreophora within the clade Genasauria. Neornithischians are united by having a thicker layer of asymmetrical enamel on the insi ...
n
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic Geological period, period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the #Evolutio ...
s from the
Cretaceous The Cretaceous ( ) is a geological period that lasted from about 143.1 to 66 mya (unit), million years ago (Mya). It is the third and final period of the Mesozoic Era (geology), Era, as well as the longest. At around 77.1 million years, it is the ...
Period (
Aptian The Aptian is an age (geology), age in the geologic timescale or a stage (stratigraphy), stage in the stratigraphic column. It is a subdivision of the Early Cretaceous, Early or Lower Cretaceous epoch (geology), Epoch or series (stratigraphy), S ...
-
Cenomanian The Cenomanian is, in the International Commission on Stratigraphy's (ICS) geological timescale, the oldest or earliest age (geology), age of the Late Cretaceous epoch (geology), Epoch or the lowest stage (stratigraphy), stage of the Upper Cretace ...
). The
family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
was first proposed by Han ''et al.'' in 2012. The jeholosaurids were defined as all neornithischians more closely related to ''Jeholosaurus shangyuanensis'' than to ''
Hypsilophodon foxii ''Hypsilophodon'' (; meaning "high-crested tooth") is a neornithischian dinosaur genus from the Early Cretaceous Period (geology), period of England. It has traditionally been considered an early member of the group Ornithopoda, but recent resear ...
'', ''
Iguanodon bernissartensis ''Iguanodon'' ( ; meaning 'iguana-tooth'), named in 1825, is a genus of iguanodontian dinosaur. While many species found worldwide have been classified in the genus ''Iguanodon'', dating from the Late Jurassic to Early Cretaceous, Taxonomy (bi ...
'', ''
Protoceratops andrewsi ''Protoceratops'' (; ) is a genus of small protoceratopsid dinosaurs that lived in Asia during the Late Cretaceous, around 75 to 71 million years ago. The genus ''Protoceratops'' includes two species: ''P. andrewsi'' and the larger ''P. helleni ...
'', ''
Pachycephalosaurus wyomingensis ''Pachycephalosaurus'' (; meaning "thick-headed lizard", from Ancient Greek, Greek ''pachys-/'' "thickness", ''kephalon/'' "head" and ''sauros/'' "lizard") is a genus of pachycephalosaurid Ornithischia, ornithischian dinosaur. The type species, ...
'', or ''
Thescelosaurus neglectus ''Thescelosaurus'' ( ) is a genus of ornithischian dinosaur that lived during the Late Cretaceous period in western North America. It was named and described in 1913 by the paleontologist Charles W. Gilmore; the type species is ''T. neglectus''. ...
''. The Jeholosauridae includes the
type genus In biological taxonomy, the type genus (''genus typica'') is the genus which defines a biological family and the root of the family name. Zoological nomenclature According to the International Code of Zoological Nomenclature, "The name-bearin ...
''
Jeholosaurus ''Jeholosaurus'' is a genus of neornithischian dinosaur from the Early Cretaceous Period. It is thought to have been a herbivorous small ornithopod. History of discovery The first two ''Jeholosaurus'' specimens were found in 2000 at Lujiatu ...
'', and also possibly ''
Yueosaurus ''Yueosaurus'' is an extinct genus of basal neornithischian dinosaur known from Zhejiang Province, China. Description ''Yueosaurus'' is known only from the holotype ZMNH M8620, an articulated, partial but well preserved postcranial skel ...
'', ''
Changchunsaurus ''Changchunsaurus'' (meaning "Changchun lizard") is an extinct genus of small herbivorous dinosaur from Early Cretaceous deposits of Gongzhuling, Jilin, China. It is the first named dinosaur genus from Jilin. Description ''Changchunsaurus'' was ...
'', and '' Haya''.Boyd CA. (2015) The systematic relationships and biogeographic history of ornithischian dinosaurs. PeerJ 3:e1523 https://doi.org/10.7717/peerj.1523 The
cladogram A cladogram (from Greek language, Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an Phylogenetic tree, evolutionary tree because it does not s ...
below results from analysis by Herne ''et al.'', 2019, which added ''
Changchunsaurus ''Changchunsaurus'' (meaning "Changchun lizard") is an extinct genus of small herbivorous dinosaur from Early Cretaceous deposits of Gongzhuling, Jilin, China. It is the first named dinosaur genus from Jilin. Description ''Changchunsaurus'' was ...
'' and '' Haya'' to Jeholosauridae, but moved ''
Yueosaurus ''Yueosaurus'' is an extinct genus of basal neornithischian dinosaur known from Zhejiang Province, China. Description ''Yueosaurus'' is known only from the holotype ZMNH M8620, an articulated, partial but well preserved postcranial skel ...
'' to the sister family
Thescelosauridae Thescelosauridae is a clade of neornithischians from the Cretaceous of East Asia and North America. The group was originally used as a name by Charles M. Sternberg in 1937, but was not formally defined until 2013, where it was used by Brown and ...
.


References

Neornithischia Cretaceous dinosaurs Dinosaur families {{ornithischian-stub