''Leptochilus pteropus'' (

synonym A synonym is a word, morpheme, or phrase that means precisely or nearly the same as another word, morpheme, or phrase in a given language. For example, in the English language, the words ''begin'', ''start'', ''commence'', and ''initiate'' are a ...

''Microsorum pteropus'') is a species of aquatic or semi-aquatic

fern The ferns (Polypodiopsida or Polypodiophyta) are a group of vascular plants (plants with xylem and phloem) that reproduce via spores and have neither seeds nor flowers. They differ from mosses by being vascular, i.e., having specialized tissue ...

, commonly known as Java fern (after the

Indonesia Indonesia, officially the Republic of Indonesia, is a country in Southeast Asia and Oceania, between the Indian Ocean, Indian and Pacific Ocean, Pacific oceans. Comprising over List of islands of Indonesia, 17,000 islands, including Sumatra, ...

n island of

Java Java is one of the Greater Sunda Islands in Indonesia. It is bordered by the Indian Ocean to the south and the Java Sea (a part of Pacific Ocean) to the north. With a population of 156.9 million people (including Madura) in mid 2024, proje ...

). It is predominantly known from

Malaysia Malaysia is a country in Southeast Asia. Featuring the Tanjung Piai, southernmost point of continental Eurasia, it is a federation, federal constitutional monarchy consisting of States and federal territories of Malaysia, 13 states and thre ...

Thailand Thailand, officially the Kingdom of Thailand and historically known as Siam (the official name until 1939), is a country in Southeast Asia on the Mainland Southeast Asia, Indochinese Peninsula. With a population of almost 66 million, it spa ...

Northeastern India Northeast India, officially the North Eastern Region (NER), is the easternmost region of India representing both a geographic and political administrative division of the country. It comprises eight states—Arunachal Pradesh, Assam, Manipur, M ...

and parts of

South China South China ( zh, s=, p=Huá'nán, j=jyut6 naam4) is a geographical and cultural region that covers the southernmost part of China. Its precise meaning varies with context. A notable feature of South China in comparison to the rest of China is ...

. It is a highly variable plant, with several different geographic varieties and physical forms that vary in leaf size, texture and shape (such as the crested-leaved 'Windelov' or 'Trident' varieties). Found in its natural habitat growing attached to

riparian A riparian zone or riparian area is the interface between land and a river or stream. In some regions, the terms riparian woodland, riparian forest, riparian buffer zone, riparian corridor, and riparian strip are used to characterize a ripar ...

roots and rocks, it can grow fully or partially submerged, provided its roots and rhizome remain wet. The plant will readily propagate asexually by producing small,

adventitious Important structures in plant development are buds, shoots, roots, leaves, and flowers; plants produce these tissues and structures throughout their life from meristems located at the tips of organs, or between mature tissues. Thus, a living plant ...

plantlets from the leaf tips and margins of established, mature foliage. The small sprouts quickly develop roots of their own, and, if water currents facilitate it, may attach themselves to surrounding objects before the "mother" leaf expires. In the event that the mother leaf wilts or detaches prior to the plantlet establishing itself, the small fern will float on its own to a new location to start anew.

Cultivation and uses

Java fern is one of the most popular plants in the aquarium hobby, due to its aesthetic appeal and ease of care. Several cultivars of Java fern exist, including the "narrow leaf", "needle leaf", "Windelov", "trident", and "lance leaf" variants. Cultivation in the aquarium is usually performed by tying or glueing the roots coming from the

rhizome In botany and dendrology, a rhizome ( ) is a modified subterranean plant stem that sends out roots and Shoot (botany), shoots from its Node (botany), nodes. Rhizomes are also called creeping rootstalks or just rootstalks. Rhizomes develop from ...

to rock or driftwood, instead of planting it directly into the substrate. It can be cultivated in tap water, dim or bright light, with or without gravel. It is more snail resistant and grows better with higher fish loads.

Biology

Java ferns are

epiphyte An epiphyte is a plant or plant-like organism that grows on the surface of another plant and derives its moisture and nutrients from the air, rain, water (in marine environments) or from debris accumulating around it. The plants on which epiphyt ...

s: they attach to surfaces rather than rooting in substrate.

References

Ferns of Asia Freshwater plants Plants described in 1933 Polypodiaceae Aquarium plants {{Polypodiaceae-stub