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The jaguar (''Panthera onca'') is a large
cat The cat (''Felis catus''), also referred to as the domestic cat or house cat, is a small domesticated carnivorous mammal. It is the only domesticated species of the family Felidae. Advances in archaeology and genetics have shown that the ...
species and the only
living Living or The Living may refer to: Common meanings *Life, a condition that distinguishes organisms from inorganic objects and dead organisms ** Living species, one that is not extinct *Personal life, the course of an individual human's life * ...
member of the genus ''
Panthera ''Panthera'' is a genus within the family (biology), family Felidae, and one of two extant genera in the subfamily Pantherinae. It contains the largest living members of the cat family. There are five living species: the jaguar, leopard, lion, ...
'' that is native to the
Americas The Americas, sometimes collectively called America, are a landmass comprising the totality of North America and South America.''Webster's New World College Dictionary'', 2010 by Wiley Publishing, Inc., Cleveland, Ohio. When viewed as a sing ...
. With a body length of up to and a weight of up to , it is the biggest cat species in the Americas and the third largest in the world. Its distinctively marked
coat A coat is typically an outer garment for the upper body, worn by any gender for warmth or fashion. Coats typically have long sleeves and are open down the front, and closing by means of buttons, zippers, hook-and-loop fasteners (AKA velcro), ...
features pale yellow to tan colored fur covered by spots that transition to rosettes on the sides, although a
melanistic Melanism is the congenital excess of melanin in an organism resulting in dark pigment. Pseudomelanism, also called abundism, is another variant of pigmentation, identifiable by dark spots or enlarged stripes, which cover a large part of the bod ...
black coat appears in some individuals. The jaguar's powerful bite allows it to pierce the
carapace A carapace is a dorsal (upper) section of the exoskeleton or shell in a number of animal groups, including arthropods, such as crustaceans and arachnids, as well as vertebrates, such as turtles and tortoises. In turtles and tortoises, the unde ...
s of
turtle Turtles are reptiles of the order (biology), order Testudines, characterized by a special turtle shell, shell developed mainly from their ribs. Modern turtles are divided into two major groups, the Pleurodira (side necked turtles) and Crypt ...
s and
tortoise Tortoises ( ) are reptiles of the family Testudinidae of the order Testudines (Latin for "tortoise"). Like other turtles, tortoises have a shell to protect from predation and other threats. The shell in tortoises is generally hard, and like o ...
s, and to employ an unusual killing method: it bites directly through the skull of
mammal A mammal () is a vertebrate animal of the Class (biology), class Mammalia (). Mammals are characterised by the presence of milk-producing mammary glands for feeding their young, a broad neocortex region of the brain, fur or hair, and three ...
ian
prey Predation is a biological interaction in which one organism, the predator, kills and eats another organism, its prey. It is one of a family of common feeding behaviours that includes parasitism and micropredation (which usually do not ki ...
between the ears to deliver a fatal blow to the brain. The modern jaguar's ancestors probably entered the Americas from
Eurasia Eurasia ( , ) is a continental area on Earth, comprising all of Europe and Asia. According to some geographers, Physical geography, physiographically, Eurasia is a single supercontinent. The concept of Europe and Asia as distinct continents d ...
during the
Early Pleistocene The Early Pleistocene is an unofficial epoch (geology), sub-epoch in the international geologic timescale in chronostratigraphy, representing the earliest division of the Pleistocene Epoch within the ongoing Quaternary Period. It is currently esti ...
via the
land bridge In biogeography, a land bridge is an isthmus or wider land connection between otherwise separate areas, over which animals and plants are able to cross and colonize new lands. A land bridge can be created by marine regression, in which sea le ...
that once spanned the
Bering Strait The Bering Strait ( , ; ) is a strait between the Pacific and Arctic oceans, separating the Chukchi Peninsula of the Russian Far East from the Seward Peninsula of Alaska. The present Russia–United States maritime boundary is at 168° 58' ...
. Today, the jaguar's range extends from the
Southwestern United States The Southwestern United States, also known as the American Southwest or simply the Southwest, is a geographic and cultural list of regions of the United States, region of the United States that includes Arizona and New Mexico, along with adjacen ...
across
Mexico Mexico, officially the United Mexican States, is a country in North America. It is the northernmost country in Latin America, and borders the United States to the north, and Guatemala and Belize to the southeast; while having maritime boundar ...
and much of
Central America Central America is a subregion of North America. Its political boundaries are defined as bordering Mexico to the north, Colombia to the southeast, the Caribbean to the east, and the Pacific Ocean to the southwest. Central America is usually ...
, the
Amazon rainforest The Amazon rainforest, also called the Amazon jungle or Amazonia, is a Tropical and subtropical moist broadleaf forests, moist broadleaf tropical rainforest in the Amazon biome that covers most of the Amazon basin of South America. This basin ...
and south to
Paraguay Paraguay, officially the Republic of Paraguay, is a landlocked country in South America. It is bordered by Argentina to the Argentina–Paraguay border, south and southwest, Brazil to the Brazil–Paraguay border, east and northeast, and Boli ...
and northern
Argentina Argentina, officially the Argentine Republic, is a country in the southern half of South America. It covers an area of , making it the List of South American countries by area, second-largest country in South America after Brazil, the fourt ...
. It inhabits a variety of forested and open terrains, but its preferred
habitat In ecology, habitat refers to the array of resources, biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species' habitat can be seen as the physical manifestation of its ...
is
tropical and subtropical moist broadleaf forest Tropical and subtropical moist broadleaf forests (TSMF), also known as tropical moist forest, is a subtropical and tropical forest habitat type defined by the World Wide Fund for Nature (WWF). Description TSMF is generally found in large ...
,
wetland A wetland is a distinct semi-aquatic ecosystem whose groundcovers are flooded or saturated in water, either permanently, for years or decades, or only seasonally. Flooding results in oxygen-poor ( anoxic) processes taking place, especially ...
s and wooded regions. It is adept at swimming and is largely a solitary, opportunistic, stalk-and-ambush
apex predator An apex predator, also known as a top predator or superpredator, is a predator at the top of a food chain, without natural predators of its own. Apex predators are usually defined in terms of trophic dynamics, meaning that they occupy the hig ...
. As a
keystone species A keystone species is a species that has a disproportionately large effect on its natural environment relative to its abundance. The concept was introduced in 1969 by the zoologist Robert T. Paine. Keystone species play a critical role in main ...
, it plays an important role in stabilizing
ecosystem An ecosystem (or ecological system) is a system formed by Organism, organisms in interaction with their Biophysical environment, environment. The Biotic material, biotic and abiotic components are linked together through nutrient cycles and en ...
s and in regulating prey populations. The jaguar is threatened by
habitat loss Habitat destruction (also termed habitat loss or habitat reduction) occurs when a natural habitat is no longer able to support its native species. The organisms once living there have either moved elsewhere, or are dead, leading to a decrease ...
,
habitat fragmentation Habitat fragmentation describes the emergence of discontinuities (fragmentation) in an organism's preferred environment (habitat), causing population fragmentation and ecosystem decay. Causes of habitat fragmentation include geological proces ...
,
poaching Poaching is the illegal hunting or capturing of wild animals, usually associated with land use rights. Poaching was once performed by impoverished peasants for subsistence purposes and to supplement meager diets. It was set against the huntin ...
for trade with its body parts and killings in
human–wildlife conflict Human–wildlife conflict (HWC) refers to the negative interactions between humans and wild animals, with undesirable consequences both for people and their resources on the one hand, and wildlife and their habitats on the other. HWC, caused by ...
situations, particularly with
ranch A ranch (from /Mexican Spanish) is an area of landscape, land, including various structures, given primarily to ranching, the practice of raising grazing livestock such as cattle and sheep. It is a subtype of farm. These terms are most often ap ...
ers in Central and
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a considerably smaller portion in the Northern Hemisphere. It can also be described as the southern Subregion#Americas, subregion o ...
. It has been listed as
Near Threatened A near-threatened species is a species which has been Conservation status, categorized as "Near Threatened" (NT) by the International Union for Conservation of Nature (IUCN) as that may be vulnerable to Endangered species, endangerment in the ne ...
on the
IUCN Red List The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is an inventory of the global conservation status and extinction risk of biological ...
since 2002. The wild population is thought to have declined since the late 1990s. Priority areas for jaguar
conservation Conservation is the preservation or efficient use of resources, or the conservation of various quantities under physical laws. Conservation may also refer to: Environment and natural resources * Nature conservation, the protection and manage ...
comprise 51 Jaguar Conservation Units (JCUs), defined as large areas inhabited by at least 50 breeding jaguars. The JCUs are located in 36 geographic regions ranging from Mexico to Argentina. The jaguar has featured prominently in the mythology of
indigenous peoples of the Americas In the Americas, Indigenous peoples comprise the two continents' pre-Columbian inhabitants, as well as the ethnic groups that identify with them in the 15th century, as well as the ethnic groups that identify with the pre-Columbian population of ...
, including those of the
Aztec The Aztecs ( ) were a Mesoamerican civilization that flourished in central Mexico in the Post-Classic stage, post-classic period from 1300 to 1521. The Aztec people included different Indigenous peoples of Mexico, ethnic groups of central ...
and
Maya civilization The Maya civilization () was a Mesoamerican civilization that existed from antiquity to the early modern period. It is known by its ancient temples and glyphs (script). The Maya script is the most sophisticated and highly developed writin ...
s.


Etymology

The word "jaguar" is possibly derived from the Tupi-Guarani word meaning 'wild beast that overcomes its prey at a bound'. In North America, the word is pronounced disyllabic , while in British English, it is pronounced with three syllables . Because that word also applies to other animals,
indigenous peoples in Guyana Indigenous peoples in Guyana, Native Guyanese or Amerindian Guyanese are Guyanese people who are of Indigenous peoples in South America, Indigenous ancestry. They comprise approximately 9.16% of Guyana's population. Amerindians are credited with ...
call it , with the added sufix ''eté'', meaning "true beast". "Onca" is derived from the Portuguese name for a spotted cat that is larger than a
lynx A lynx ( ; : lynx or lynxes) is any of the four wikt:extant, extant species (the Canada lynx, Iberian lynx, Eurasian lynx and the bobcat) within the medium-sized wild Felidae, cat genus ''Lynx''. The name originated in Middle Engl ...
; cf. ''ounce''. The word "panther" is derived from
classical Latin Classical Latin is the form of Literary Latin recognized as a Literary language, literary standard language, standard by writers of the late Roman Republic and early Roman Empire. It formed parallel to Vulgar Latin around 75 BC out of Old Latin ...
, itself from the
ancient Greek Ancient Greek (, ; ) includes the forms of the Greek language used in ancient Greece and the classical antiquity, ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Greek ...
().


Taxonomy and evolution


Taxonomy

In 1758,
Carl Linnaeus Carl Linnaeus (23 May 1707 – 10 January 1778), also known after ennoblement in 1761 as Carl von Linné,#Blunt, Blunt (2004), p. 171. was a Swedish biologist and physician who formalised binomial nomenclature, the modern system of naming o ...
described the jaguar in his work ''
Systema Naturae ' (originally in Latin written ' with the Orthographic ligature, ligature æ) is one of the major works of the Sweden, Swedish botanist, zoologist and physician Carl Linnaeus (1707–1778) and introduced the Linnaean taxonomy. Although the syste ...
'' and gave it the
scientific name In Taxonomy (biology), taxonomy, binomial nomenclature ("two-term naming system"), also called binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, both of which use Latin gramm ...
''Felis onca''. In the 19th and 20th centuries, several jaguar
type specimen In biology, a type is a particular wikt:en:specimen, specimen (or in some cases a group of specimens) of an organism to which the scientific name of that organism is formally associated. In other words, a type is an example that serves to ancho ...
s formed the basis for descriptions of
subspecies In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...
. In 1939,
Reginald Innes Pocock Reginald Innes Pocock, (4 March 1863 – 9 August 1947) was a British zoologist. Pocock was born in Clifton, Bristol, the fourth son of Rev. Nicholas Pocock and Edith Prichard. He began showing interest in natural history at St. Edward's ...
recognized eight subspecies based on the geographic origins and skull
morphology Morphology, from the Greek and meaning "study of shape", may refer to: Disciplines *Morphology (archaeology), study of the shapes or forms of artifacts *Morphology (astronomy), study of the shape of astronomical objects such as nebulae, galaxies, ...
of these specimens. Pocock did not have access to sufficient
zoological specimen A zoological specimen is an animal or part of an animal preserved for scientific use. Various uses are: to verify the identity of a (species), to allow study, increase public knowledge of zoology. Zoological specimens are extremely diverse. Exampl ...
s to critically evaluate their subspecific status but expressed doubt about the status of several. Later consideration of his work suggested only three subspecies should be recognized. The description of ''P. o. palustris'' was based on a fossil skull. By 2005, nine subspecies were considered to be Valid name (zoology), valid taxa: *''P. o. onca'' was a jaguar from Brazil. *''P. o. peruviana'' was a jaguar skull from Peru. *''P. o. hernandesii'' was a jaguar from Mazatlán in Mexico. *''P. o. palustris'' was a fossil jaguar mandible excavated in the Sierras Pampeanas of Córdova District, Argentina. *''P. o. centralis'' was a skull of a male jaguar from Talamanca (canton), Talamanca, Costa Rica. *''P. o. goldmani'' was a jaguar skin from Yohatlan in Campeche, Mexico. *''P. o. paraguensis'' was a skull of a male jaguar from Paraguay. *''P. o. arizonensis'' was a skin and skull of a male jaguar from the vicinity of Cibecue, Arizona. *''P. o. veraecrucis'' was a skull of a male jaguar from San Andrés Tuxtla in Mexico. Reginald Innes Pocock placed the jaguar in the genus ''Panthera'' and observed that it shares several morphological features with the leopard (''P. pardus''). He, therefore, concluded that they are most closely related to each other. Results of Morphology (biology), morphological and genetics, genetic research indicate a cline (population genetics), clinal north–south variation between populations, but no evidence for subspecific differentiation. DNA analysis of 84 jaguar samples from South America revealed that the gene flow between jaguar populations in Colombia was high in the past. Since 2017, the jaguar is considered to be a monotypic taxon, though the modern ''Panthera onca onca'' is still distinguished from two fossil subspecies, ''Panthera onca augusta'' and ''Panthera onca mesembrina''. However, the 2024 study suggested that the validity of subspecific assignments on both ''P. o. augusta'' and ''P. o. mesembrina'' remains unresolved, since both fossil and living jaguars show a considerable variation in morphometry.


Evolution

The ''Panthera'' Lineage (evolution), lineage is estimated to have Genetic divergence, genetically diverged from the common ancestor of the Felidae around to . Some genetic analyzes place the jaguar as a sister species to the lion with which it diverged , but other studies place the lion closer to the leopard. The lineage of the jaguar appears to have originated in Africa and spread to Eurasia 1.95–1.77 mya. The living jaguar species is often suggested to have descended from the Eurasian ''Panthera gombaszoegensis.'' The ancestor of the jaguar entered the American continent via Beringia, the land bridge that once spanned the
Bering Strait The Bering Strait ( , ; ) is a strait between the Pacific and Arctic oceans, separating the Chukchi Peninsula of the Russian Far East from the Seward Peninsula of Alaska. The present Russia–United States maritime boundary is at 168° 58' ...
, Some authors have disputed the close relationship between ''P. gombaszoegensis'' (which is primarily known from Europe) and the modern jaguar. The oldest fossils of modern jaguars (''P. onca'') have been found in North America dating between 850,000-820,000 years ago. Results of mitochondrial DNA analysis of 37 jaguars indicate that current populations evolved between 510,000 and 280,000 years ago in northern South America and subsequently recolonized North and Central America after the extinction of jaguars there during the Late Pleistocene. Two extinct subspecies of jaguar are recognized in the fossil record: the North American ''Panthera onca augusta, P. o. augusta'' and South American ''Panthera onca mesembrina, P. o. mesembrina''.


Description

The jaguar is a compact and muscular animal. It is the largest cat native to the Americas and the third largest in the world, exceeded in size only by the tiger and the lion. It stands tall at the shoulders. Its size and weight vary considerably depending on sex and region: weights in most regions are normally in the range of . Exceptionally big males have been recorded to weigh as much as . The smallest females from Middle America (Americas), Middle America weigh about . It is sexually dimorphic, with females typically being 10–20% smaller than males. The length from the nose to the base of the tail varies from . The tail is long and the shortest of any big cat. Its muscular legs are shorter than the legs of other ''Panthera'' species with similar body weight. Size tends to increase from north to south. Jaguars in the Chamela-Cuixmala Biosphere Reserve on the Pacific coast of central Mexico weighed around . Jaguars in Venezuela and Brazil are much larger, with average weights of about in males and of about in females. The jaguar's
coat A coat is typically an outer garment for the upper body, worn by any gender for warmth or fashion. Coats typically have long sleeves and are open down the front, and closing by means of buttons, zippers, hook-and-loop fasteners (AKA velcro), ...
ranges from pale yellow to tan or reddish-yellow, with a whitish underside and covered in black spots. The spots and their shapes vary: on the sides, they become rosette (zoology), rosettes which may include one or several dots. The spots on the head and neck are generally solid, as are those on the tail where they may merge to form bands near the end and create a black tip. They are elongated on the middle of the back, often connecting to create a median stripe, and blotchy on the belly. These patterns serve as camouflage in areas with dense vegetation and patchy shadows. Jaguars living in forests are often darker and considerably smaller than those living in open areas, possibly due to the smaller numbers of large, herbivorous prey in forest areas. The jaguar closely resembles the leopard but is generally more robust, with stockier limbs and a more square head. The rosettes on a jaguar's coat are larger, darker, fewer in number and have thicker lines, with a small spot in the middle. It has powerful jaws with the third-highest bite force of all felids, after the tiger and the lion. It has an average bite force at the Canine tooth, canine tip of 887.0 Newton (unit), Newton and a bite force quotient at the canine tip of 118.6. A jaguar can bite with a force of with the canine teeth and at the carnassial notch.


Color variation

Melanistic jaguars are also known as black panthers. The black Morph (biology), morph is less common than the spotted one. Black jaguars have been documented in Central and South America. Melanism in the jaguar is caused by deletions in the melanocortin 1 receptor gene and inherited through a dominant allele. Black jaguars occur at higher densities in tropical rainforest and are more active during the daytime. This suggests that melanism provides camouflage in dense vegetation with high illumination. In 2004, a camera trap in the Sierra Madre Occidental mountains photographed the first documented black jaguar in Northern Mexico. Black jaguars were also photographed in Costa Rica's Alberto Manuel Brenes Biological Reserve, in the mountains of the Cordillera de Talamanca, in Barbilla National Park and in eastern Panama.


Distribution and habitat

In 1999, the jaguar's historic range at the turn of the 20th century was estimated at , stretching from the southern United States through Central America to southern Argentina. By the turn of the 21st century, its global range had decreased to about , with most declines occurring in the southern United States, northern Mexico, northern Brazil, and southern Argentina. Its present range extends from Mexico through Central America to South America comprising Belize, Guatemala, Honduras, Nicaragua, Costa Rica, particularly on the Osa Peninsula, Panama, Colombia, Venezuela, Guyana, Suriname, French Guiana, Ecuador, Peru, Bolivia, Brazil,
Paraguay Paraguay, officially the Republic of Paraguay, is a landlocked country in South America. It is bordered by Argentina to the Argentina–Paraguay border, south and southwest, Brazil to the Brazil–Paraguay border, east and northeast, and Boli ...
and
Argentina Argentina, officially the Argentine Republic, is a country in the southern half of South America. It covers an area of , making it the List of South American countries by area, second-largest country in South America after Brazil, the fourt ...
. It is considered to be locally extinct in El Salvador and Uruguay. Jaguars have been occasionally sighted in Arizona, New Mexico and Texas, with 62 accounts reported in the 20th century. Between 2012 and 2015, a male Vagrancy (biology), vagrant jaguar was recorded in 23 locations in the Santa Rita Mountains. Eight jaguars were photographed in the southwestern US between 1996 and 2024. The jaguar prefers dense forest and typically inhabits dry deciduous forests, tropical and subtropical moist broadleaf forests, rainforests and cloud forests in Central and South America; open, seasonally flooded
wetland A wetland is a distinct semi-aquatic ecosystem whose groundcovers are flooded or saturated in water, either permanently, for years or decades, or only seasonally. Flooding results in oxygen-poor ( anoxic) processes taking place, especially ...
s, dry grassland and historically also oak forests in the United States. It has been recorded at elevations up to but avoids montane forests. It favors riverine habitat and swamps with dense vegetation cover. In the Mayan forests of Mexico and Guatemala, 11 GPS-collared jaguars preferred undisturbed dense habitat away from roads; females avoided even areas with low levels of human activity, whereas males appeared less disturbed by human population density. A young male jaguar was also recorded in the Semi-arid climate, semi-arid Sierra de San Carlos at a waterhole.


Former range

In the 19th century, the jaguar was still sighted at the North Platte River north of Longs Peak in Colorado, in coastal Louisiana, northern Arizona and New Mexico. Multiple verified zoological reports of the jaguar are known in California, two as far north as Monterey, California, Monterey in 1814 and 1826. The only record of an active jaguar den with breeding adults and kittens in the United States was in the Tehachapi Mountains of California prior to 1860. The jaguar persisted in California until about 1860. The last confirmed jaguar in Texas was shot in 1948, southeast of Kingsville, Texas. In Arizona, a female was shot in the White Mountains (Arizona), White Mountains in 1963. By the late 1960s, the jaguar was thought to have been extirpated in the United States. Arizona outlawed jaguar hunting in 1969, but by then no females remained, and over the next 25 years only two males were sighted and killed in the state. In 1996, a rancher and hunting guide from Douglas, Arizona came across a jaguar in the Peloncillo Mountains (Cochise County), Peloncillo Mountains and became a researcher on jaguars, placing trail cameras, which recorded four more jaguars.


Behavior and ecology

The jaguar is mostly active at night and during twilight. However, jaguars living in densely forested regions of the Amazon Rainforest and the Pantanal are largely active by day, whereas jaguars in the Atlantic Forest are primarily active by night. The activity pattern of the jaguar coincides with the activity of its main prey species. Jaguars are good swimmers and play and hunt in the water, possibly more than tigers. They have been recorded moving between islands and the shore. Jaguars are also good at climbing trees but do so less often than cougars.


Ecological role

The adult jaguar is an
apex predator An apex predator, also known as a top predator or superpredator, is a predator at the top of a food chain, without natural predators of its own. Apex predators are usually defined in terms of trophic dynamics, meaning that they occupy the hig ...
, meaning it is at the top of the food chain and is not preyed upon in the wild. The jaguar has also been termed a
keystone species A keystone species is a species that has a disproportionately large effect on its natural environment relative to its abundance. The concept was introduced in 1969 by the zoologist Robert T. Paine. Keystone species play a critical role in main ...
, as it is assumed that it controls the population levels of prey such as herbivorous and seed-eating mammals and thus maintains the structural integrity of forest systems. However, field work has shown this may be natural variability, and the population increases may not be sustained. Thus, the keystone predator hypothesis is not accepted by all scientists. The jaguar is sympatric with the cougar. In central Mexico, both prey on white-tailed deer, which makes up 54% and 66% of jaguar and cougar's prey, respectively. In northern Mexico, the jaguar and the cougar share the same habitat, and their diet overlaps dependent on prey availability. Jaguars seemed to prefer deer and calves. In Mexico and Central America, neither of the two cats are considered to be the dominant predator. In South America, the jaguar is larger than the cougar and tends to take larger prey, usually over . The cougar's prey usually weighs between , which is thought to be the reason for its smaller size. This situation may be advantageous to the cougar. Its broader prey niche, including its ability to take smaller prey, may give it an advantage over the jaguar in human-altered landscapes.


Hunting and diet

The jaguar is an obligate carnivore and depends solely on flesh for its nutrient requirements. An analysis of 53 studies documenting the diet of the jaguar revealed that its prey ranges in weight from ; it prefers prey weighing , with the capybara and the giant anteater being the most selected. When available, it also preys on marsh deer, southern tamandua, collared peccary and black agouti. In floodplains, jaguars opportunistically take reptiles such as green anacondas,
turtle Turtles are reptiles of the order (biology), order Testudines, characterized by a special turtle shell, shell developed mainly from their ribs. Modern turtles are divided into two major groups, the Pleurodira (side necked turtles) and Crypt ...
s and caimans. Consumption of reptiles appears to be more frequent in jaguars than in other big cats. One remote population in the Brazilian Pantanal is recorded to primarily feed on aquatic reptiles and fish. The jaguar also preys on livestock in cattle
ranch A ranch (from /Mexican Spanish) is an area of landscape, land, including various structures, given primarily to ranching, the practice of raising grazing livestock such as cattle and sheep. It is a subtype of farm. These terms are most often ap ...
ing areas where wild prey is scarce. The daily food requirement of a captive jaguar weighing was estimated at of meat. The jaguar's bite force allows it to pierce the
carapace A carapace is a dorsal (upper) section of the exoskeleton or shell in a number of animal groups, including arthropods, such as crustaceans and arachnids, as well as vertebrates, such as turtles and tortoises. In turtles and tortoises, the unde ...
s of the yellow-spotted Amazon river turtle and the yellow-footed tortoise. It employs an unusual killing method: it bites mammalian prey directly through the skull between the ears to deliver a fatal bite to the brain. It kills capybara by piercing its canine tooth, canine teeth through the temporal bones of its skull, breaking its zygomatic arch and mandible and penetrating its brain, often through the ears. It has been hypothesized to be an adaptation to cracking open turtle shells; armored reptiles may have formed an abundant prey base for the jaguar following the late Pleistocene extinctions. However, this is disputed, as even in areas where jaguars prey on reptiles, they are still taken relatively infrequently compared to mammals in spite of their greater abundance. Between October 2001 and April 2004, 10 jaguars were monitored in the southern Pantanal. In the dry season from April to September, they killed prey at intervals ranging from one to seven days; and ranging from one to 16 days in the wet season from October to March. The jaguar uses a stalk-and-ambush strategy when hunting rather than chasing prey. The cat will slowly walk down forest paths, listening for and stalking prey before rushing or ambushing. The jaguar attacks from cover and usually from a target's blind spot with a quick pounce; the species' ambushing abilities are considered nearly peerless in the animal kingdom by both indigenous people and field researchers and are probably a product of its role as an apex predator in several different environments. The ambush may include leaping into water after prey, as a jaguar is quite capable of carrying a large kill while swimming; its strength is such that carcasses as large as a Heifer (cow), heifer can be hauled up a tree to avoid flood levels. After killing prey, the jaguar will drag the carcass to a thicket or other secluded spot. It begins eating at the neck and chest. The heart and lungs are consumed, followed by the shoulders.


Social activity

The jaguar is generally Solitary animal, solitary except for females with cubs. In 1977, groups consisting of a male, female and cubs, and two females with two males were sighted several times in a study area in the Paraguay River valley; a radio-collared female moved in a home range of , which partly overlapped with another female. The home range of the male in this study area overlapped with several females. In the Venezuelan Llanos and Brazilian Pantanal, male coalitions were detected, which marked, defended and invaded territories together, hunted together and mated with several females. The jaguar uses scrape marks, urine, and feces to Territorial marking, mark its territory. The size of home ranges depends on the level of deforestation and human population density. The home ranges of females vary from in the Pantanal to in the Amazon to in the Atlantic Forest. Male jaguar home ranges vary from in the Pantanal to in the Amazon to in the Atlantic Forest and in the Cerrado. Studies employing Global Positioning System, GPS telemetry in 2003 and 2004 found densities of only six to seven jaguars per in the Pantanal region, compared with 10 to 11 using traditional methods; this suggests the widely used sampling methods may inflate the actual numbers of individuals in a sampling area. Fights between males occur but are rare, and avoidance behavior has been observed in the wild. In one wetland population with degraded territorial boundaries and more social proximity, adults of the same sex are more tolerant of each other and engage in more friendly and co-operative interactions. The jaguar roars/grunts for long-distance communication; intensive bouts of counter-calling between individuals have been observed in the wild. This vocalization is described as "hoarse" with five or six guttural notes. prusten, Chuffing is produced by individuals when greeting, during Courtship display, courting, or by a mother comforting her cubs. This sound is described as low intensity snorts, possibly intended to signal tranquility and passivity. Cubs have been recorded bleating, gurgling and mewing.


Reproduction and life cycle

In captivity, the female jaguar is recorded to reach sexual maturity at the age of about 2.5 years. Estrus lasts 7–15 days with an estrus cycle of 41.8 to 52.6 days. During estrus, she exhibits increased restlessness with rolling and prolonged vocalizations. She is an induced ovulator but can also ovulate spontaneously. Gestation lasts 91 to 111 days. The male is sexually mature at the age of three to four years. His mean ejaculate volume is 8.6±1.3 ml. Generation time, Generation length of the jaguar is 9.8 years. In the Pantanal, breeding pairs were observed to stay together for up to five days. Females had one to two cubs. The young are born with closed eyes but open them after two weeks. Cubs are weaned at the age of three months but remain in the birth den for six months before leaving to accompany their mother on hunts. Jaguars remain with their mothers for up to two years. They appear to rarely live beyond 11 years, but captive individuals may live 22 years. In 2001, a male jaguar killed and partially consumed two cubs in Emas National Park. DNA paternity testing of blood samples revealed that the male was the father of the cubs. Two more cases of Infanticide (zoology), infanticide were documented in the northern Pantanal in 2013. To defend against infanticide, the female may hide her cubs and distract the male with courtship behavior.


Attacks on humans

The Spanish conquistadors feared the jaguar. According to Charles Darwin, the indigenous peoples of South America stated that people did not need to fear the jaguar as long as capybaras were abundant. The first official record of a jaguar killing a human in Brazil dates to June 2008. Two children were attacked by jaguars in Guyana. The majority of known attacks on people happened when it had been cornered or wounded.


Threats

The jaguar is threatened by habitat loss, loss and habitat fragmentation, fragmentation of habitat, illegal killing in retaliation for livestock depredation and for illegal trade in jaguar body parts. It is listed as
Near Threatened A near-threatened species is a species which has been Conservation status, categorized as "Near Threatened" (NT) by the International Union for Conservation of Nature (IUCN) as that may be vulnerable to Endangered species, endangerment in the ne ...
on the
IUCN Red List The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is an inventory of the global conservation status and extinction risk of biological ...
since 2002, as the jaguar population has probably declined by 20–25% since the mid-1990s. Deforestation is a major threat to the jaguar across its range. Habitat loss was most rapid in drier regions such as the Argentine pampas, the arid grasslands of Mexico and the southwestern United States. In 2002, it was estimated that the range of the jaguar had declined to about 46% of its range in the early 20th century. In 2018, it was estimated that its range had declined by 55% in the last century. The only remaining stronghold is the Amazon rainforest, a region that is rapidly being fragmented by deforestation. Between 2000 and 2012, forest loss in the jaguar range amounted to , with fragmentation increasing in particular in corridors between Jaguar Conservation Units (JCUs). By 2014, direct linkages between two JCUs in Bolivia were lost, and two JCUs in northern Argentina became completely isolated due to deforestation. In Mexico, the jaguar is primarily threatened by
poaching Poaching is the illegal hunting or capturing of wild animals, usually associated with land use rights. Poaching was once performed by impoverished peasants for subsistence purposes and to supplement meager diets. It was set against the huntin ...
. Its habitat is fragmented in northern Mexico, in the Gulf of Mexico and the Yucatán Peninsula, caused by changes in land use, construction of roads and tourism infrastructure. In Panama, 220 of 230 jaguars were killed in retaliation for predation on livestock between 1998 and 2014. In Venezuela, the jaguar was extirpated in about 26% of its range in the country since 1940, mostly in dry savannas and unproductive scrubland in the northeastern region of Anzoátegui. In Ecuador, the jaguar is threatened by reduced prey availability in areas where the expansion of the road network facilitated access of human hunters to forests. In the Alto Paraná Atlantic forests, at least 117 jaguars were killed in Iguaçu National Park and the adjacent Misiones Province between 1995 and 2008. Some Afro-Colombians in the Colombian Chocó Department hunt jaguars for consumption and sale of meat. Between 2008 and 2012, at least 15 jaguars were killed by livestock farmers in central Belize. The international trade of jaguar skins boomed between the end of the Second World War and the early 1970s. Significant declines occurred in the 1960s, as more than 15,000 jaguars were yearly killed for their skins in the Brazilian Amazon alone; the trade in jaguar skins decreased since 1973 when the CITES, Convention on International Trade in Endangered Species was enacted. Interview surveys with 533 people in the northwestern Bolivian Amazon revealed that local people killed jaguars out of fear, in retaliation, and for trade. Between August 2016 and August 2019, jaguar skins and body parts were seen for sale in tourist markets in the Peruvian cities of Lima, Iquitos and Pucallpa. Human-wildlife conflict, opportunistic hunting and hunting for trade in domestic markets are key drivers for killing jaguars in Belize and Guatemala. Seizure reports indicate that at least 857 jaguars were involved in trade between 2012 and 2018, including 482 individuals in Bolivia alone; 31 jaguars were seized in China. Between 2014 and early 2019, 760 jaguar fangs were seized that originated in Bolivia and were destined for China. Undercover investigations revealed that the smuggling of jaguar body parts is run by Chinese residents in Bolivia.


Conservation

The jaguar is listed on CITES Appendix I, which means that all international commercial trade in jaguars or their body parts is prohibited. Hunting jaguars is prohibited in Argentina, Brazil, Colombia, French Guiana, Honduras, Nicaragua, Panama, Paraguay, Suriname, the United States, and Venezuela. Hunting jaguars is restricted in Guatemala and Peru. In Ecuador, hunting jaguars is prohibited, and it is classified as threatened with extinction. In Guyana, it is protected as an endangered species, and hunting it is illegal. In 1986, the Cockscomb Basin Wildlife Sanctuary was established in Belize as the world's first protected area for jaguar conservation.


Jaguar Conservation Units

In 1999, field scientists from 18 jaguar range countries determined the most important areas for long-term jaguar conservation based on the status of jaguar population units, stability of prey base and quality of habitat. These areas, called "Jaguar Conservation Units" (JCUs), are large enough for at least 50 breeding individuals and range in size from ; 51 JCUs were designated in 36 geographic regions including: * the Sierra Madre Occidental and Sierra de Tamaulipas in Mexico * the Selva Maya tropical forests extending over Mexico, Belize and Guatemala * the Chocó–Darién moist forests from Honduras and Panama to Colombia * Venezuelan Llanos * northern Cerrado and Amazon basin in Brazil * Tropical Andes in Bolivia and Peru * Misiones Province in Argentina Optimal routes of travel between core jaguar population units were identified across its range in 2010 to implement wildlife corridors that connect JCUs. These corridors represent areas with the shortest distance between jaguar breeding populations, require the least possible energy input of dispersal (ecology), dispersing individuals and pose a low mortality risk. They cover an area of and range in length from in Mexico and Central America and from in South America. Cooperation with local landowners and municipal, state, or federal agencies is essential to maintain connected populations and prevent fragmentation in both JCUs and corridors. Seven of 13 corridors in Mexico are functioning with a width of at least and a length of no more than . The other corridors may hamper passage, as they are narrower and longer. In August 2012, the United States Fish and Wildlife Service set aside in Arizona and New Mexico for the protection of the jaguar. The Jaguar Recovery Plan was published in April 2019, in which Interstate 10 is considered to form the northern boundary of the Jaguar Recovery Unit in Arizona and New Mexico. In Mexico, a national conservation strategy was developed from 2005 on and published in 2016. The Mexican jaguar population increased from an estimated 4,000 individuals in 2010 to about 4,800 individuals in 2018. This increase is seen as a positive effect of conservation measures that were implemented in cooperation with governmental and non-governmental institutions and landowners. An evaluation of JCUs from Mexico to Argentina revealed that they overlap with high-quality habitats of about 1,500 mammals to varying degrees. Since co-occurring mammals benefit from the JCU approach, the jaguar has been called an umbrella species. Central American JCUs overlap with the habitat of 187 of 304 regional endemic amphibian and reptile species, of which 19 amphibians occur only in the jaguar range.


Approaches

In setting up protected reserves, efforts generally also have to be focused on the surrounding areas, as jaguars are unlikely to confine themselves to the bounds of a reservation, especially if the population is increasing in size. Human attitudes in the areas surrounding reserves and laws and regulations to prevent poaching are essential to make conservation areas effective. To estimate population sizes within specific areas and to keep track of individual jaguars, camera trapping and Telemetry#Fishery and wildlife research and management, wildlife tracking telemetry are widely used, and feces are sought out with the help of detection dogs to study jaguar health and diet. Current conservation efforts often focus on educating ranch owners and promoting ecotourism. Ecotourism setups are being used to generate public interest in charismatic animals such as the jaguar while at the same time generating revenue that can be used in conservation efforts. A key concern in jaguar ecotourism is the considerable habitat space the species requires. If ecotourism is used to aid in jaguar conservation, some considerations need to be made as to how existing ecosystems will be kept intact, or how new ecosystems will be put into place that are large enough to support a growing jaguar population. Conservationists and professionals in Mexico and the United States have established the Northern Jaguar Reserve in northern Mexico. Advocacy for reintroduction of the jaguar to its former range in Arizona and New Mexico have been supported by documentation of natural migrations by individual jaguars into the southern reaches of both states, the recency of extirpation from those regions by human action, and supportive arguments pertaining to biodiversity, ecological, human, and practical considerations.


In culture and mythology

In the pre-Columbian era, pre-Columbian Americas, the jaguar was a symbol of power and strength. In the Andes, a jaguar cult disseminated by the early Chavín culture became accepted over most of today's Peru by 900 Before Christ, BC. The later Moche (culture), Moche culture in northern Peru used the jaguar as a symbol of power in many of their ceramics. In the Muisca religion in Altiplano Cundiboyacense, the jaguar was considered a sacred animal, and people dressed in jaguar skins during religious rituals. The skins were traded with peoples in the nearby Orinoquía Region. The name of the Muisca rulers, Muisca ruler Nemequene was derived from the Chibcha language, Chibcha words ''nymy'' and ''quyne'', meaning "force of the jaguar". Sculptures with "Olmec were-jaguar" motifs were found on the Yucatán Peninsula in Veracruz and Tabasco; they show stylized jaguars with half-human faces. In the later
Maya civilization The Maya civilization () was a Mesoamerican civilization that existed from antiquity to the early modern period. It is known by its ancient temples and glyphs (script). The Maya script is the most sophisticated and highly developed writin ...
, the jaguar was known as ''balam'' or ''bolom in many of the Mayan languages, and was used to symbolize warriors and the elite class for being brave, fierce and strong. The cat was associated with the Xibalba, underworld and its image was used to decorate tombs and grave-good vessels. The
Aztec The Aztecs ( ) were a Mesoamerican civilization that flourished in central Mexico in the Post-Classic stage, post-classic period from 1300 to 1521. The Aztec people included different Indigenous peoples of Mexico, ethnic groups of central ...
civilization called the jaguar ''ocelotl'' and considered it to be the king of the animals. It was believed to be fierce and courageous, but also wise, dignified and careful. The military had two classes of warriors, the ''ocelotl'' or jaguar warriors and the ''cuauhtli'' or eagle warriors and each dressed like their representative animal. In addition, members of the royal class would decorate in jaguar skins. The jaguar was considered to be the totem animal of the powerful deities Tezcatlipoca and Tepeyollotl. A conch shell gorget depicting a jaguar was found in a burial mound in Benton County, Missouri. The gorget shows evenly-engraved lines and measures . Rock drawings made by the Hopi, Anasazi and Pueblo all over the desert and chaparral regions of the American Southwest show an explicitly spotted cat, presumably a jaguar, as it is drawn much larger than an ocelot.


See also

* List of largest cats


References


External links

* *
People and Jaguars a Guide for Coexistence

Felidae Conservation Fund
* {{Authority control Jaguars, Apex predators Big cats Carnivorans of Brazil ESA endangered species Extant Middle Pleistocene first appearances Fauna of the Amazon Fauna of the Cerrado Fauna of the Pantanal Fauna of the Southwestern United States Felids of Central America Felids of North America Felids of South America Mammals described in 1758 Near threatened animals Near threatened biota of North America Near threatened biota of South America Panthera, onca Pleistocene mammals of North America Pleistocene mammals of South America Taxa named by Carl Linnaeus