Iteaceae is a flowering plant family of trees and shrubs native to the eastern

USA The United States of America (USA), also known as the United States (U.S.) or America, is a country primarily located in North America. It is a federal republic of 50 states and a federal capital district, Washington, D.C. The 48 contiguous ...

, southeastern

Africa Africa is the world's second-largest and second-most populous continent after Asia. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 20% of Earth's land area and 6% of its total surfac ...

, and

south South is one of the cardinal directions or compass points. The direction is the opposite of north and is perpendicular to both west and east. Etymology The word ''south'' comes from Old English ''sūþ'', from earlier Proto-Germanic ''*sunþa ...

and

Southeastern Asia Southeast Asia is the geographical southeastern region of Asia, consisting of the regions that are situated south of China, east of the Indian subcontinent, and northwest of the Australian mainland, which is part of Oceania. Southeast Asia is ...

. Some older taxonomic systems place the genus ''Itea'' in the family

Grossulariaceae ''Ribes'' () is a genus of about 200 known species of flowering plants, most of them native to the temperate regions of the Northern Hemisphere. The species may be known as various kinds of currants, such as redcurrants, blackcurrants, and whi ...

. The

APG III system The APG III system of flowering plant classification is the third version of a modern, mostly molecular-based, system of plant taxonomy being developed by the Angiosperm Phylogeny Group (APG). Published in 2009, it was superseded in 2016 by a f ...

of 2009 includes the former Pterostemonaceae in Iteaceae. Consequently, it now has two genera with a total of 18 known species.

Fossil record

The family is known from fossil flowers dating to the

Turonian The Turonian is, in the International Commission on Stratigraphy, ICS' geologic timescale, the second age (geology), age in the Late Cretaceous epoch (geology), Epoch, or a stage (stratigraphy), stage in the Upper Cretaceous series (stratigraphy), ...

age of the

Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''cre ...

that have been found in the

Raritan Formation The Raritan Formation is a Cretaceous (Turonian) sedimentary geologic formation of the Atlantic Coastal Plain. Overview The formation was first described in 1888 by G. H. Cook, who measured sections in the vicinity of Raritan Bay in New Jerse ...

New Jersey New Jersey is a U.S. state, state located in both the Mid-Atlantic States, Mid-Atlantic and Northeastern United States, Northeastern regions of the United States. Located at the geographic hub of the urban area, heavily urbanized Northeas ...

and from leaves dating to the

Eocene The Eocene ( ) is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (Ma). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene'' comes ...

found in the

Klondike Mountain Formation The Klondike Mountain Formation is an Early Eocene (Ypresian) geological formation located in the northeast central area of Washington (state), Washington state. The formation is composed of volcanic rocks in the upper unit and volcanic plus lacu ...

Washington Washington most commonly refers to: * George Washington (1732–1799), the first president of the United States * Washington (state), a state in the Pacific Northwest of the United States * Washington, D.C., the capital of the United States ** A ...

. Some seeds and one fruit of †''Itea europea'' were found in two samples of muddy sediment from the Fossil Forest of Dunarobba. In northwestern

Italy Italy, officially the Italian Republic, is a country in Southern Europe, Southern and Western Europe, Western Europe. It consists of Italian Peninsula, a peninsula that extends into the Mediterranean Sea, with the Alps on its northern land b ...

, it occurs with a some seeds and fruits in two sites of Early or

Middle Pliocene Middle or The Middle may refer to: * Centre (geometry), the point equally distant from the outer limits. Places * Middle (sheading), a subdivision of the Isle of Man * Middle Bay (disambiguation) * Middle Brook (disambiguation) * Middle Creek ...

age. These records suggest it was an accessory element of the mid-Pliocene

swamp forest Freshwater swamp forests, or flooded forests, are forests which are inundated with freshwater, either permanently or seasonally. They normally occur along the lower reaches of rivers and around freshwater lakes. Freshwater swamp forests are foun ...

Fossil pollen Paleobotany or palaeobotany, also known as paleophytology, is the branch of botany dealing with the recovery and identification of plant fossils from geological contexts, and their use for the biological reconstruction of past environments (pale ...

of ''Itea'' has been detected in a short section of the

Sarzana Sarzana (, ; ) is a town, ''comune'' (municipality) and former short-lived Catholic bishopric in the Province of La Spezia, Liguria, Italy. It is east of La Spezia, on the railway to Pisa, at the point where the railway to Parma diverges to the ...

Basin in north-western part of

central Italy Central Italy ( or ) is one of the five official statistical regions of Italy used by the National Institute of Statistics (ISTAT), a first-level NUTS region with code ITI, and a European Parliament constituency. It has 11,704,312 inhabita ...

tentatively assigned to the

Miocene The Miocene ( ) is the first epoch (geology), geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and mea ...

-Pliocene transition. In this site a high pollen percentages of 12% suggest that ''Itea'' was an important element in the local vegetation. The nearest living relative of †''Itea europaea'' is the American species ''

Itea virginica ''Itea virginica'', commonly known as Virginia willow or Virginia sweetspire, is a small North American flowering shrub that grows in low-lying woods and wetland margins. Virginia willow is a member of the Iteaceae family, and native to the south ...

''.The role of central Italy as a centre of refuge for

thermophilous A thermophile is a type of extremophile that thrives at relatively high temperatures, between . Many thermophiles are archaea, though some of them are bacteria and fungi. Thermophilic eubacteria are suggested to have been among the earliest bacte ...

plants in the late Cenozoic, Edoardo Martinetto, Acta Palaeobotan. 41(2): 299-319, 2001

References

External links

Iteaceae
in Stevens, P. F. (2001 onwards). Saxifragales Saxifragales families Turonian first appearances Extant Turonian first appearances {{Saxifragales-stub