''Isthmura naucampatepetl'', commonly known as the Cofre de Perote salamander, is a species of

salamander Salamanders are a group of amphibians typically characterized by their lizard-like appearance, with slender bodies, blunt snouts, short limbs projecting at right angles to the body, and the presence of a tail in both larvae and adults. All t ...

s in the family

Plethodontidae Plethodontidae, or lungless salamanders, are a family of salamanders. With over 500 species, lungless salamanders are by far the largest family of salamanders in terms of their diversity. Most species are native to the Western Hemisphere, from B ...

. It is

endemic Endemism is the state of a species being found only in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also foun ...

to the

Sierra Madre Oriental The Sierra Madre Oriental () is a mountain range in northeastern Mexico. The Sierra Madre Oriental is part of the American Cordillera, a chain of mountain ranges (cordillera) that consists of an almost continuous sequence of mountain ranges that ...

in central

Veracruz Veracruz, formally Veracruz de Ignacio de la Llave, officially the Free and Sovereign State of Veracruz de Ignacio de la Llave, is one of the 31 states which, along with Mexico City, comprise the 32 Political divisions of Mexico, Federal Entit ...

, Mexico, where it is known from between

Cofre de Perote Cofre de Perote, also known by its Nahuatl names Naupa-Tecutépetl (from ''Nāuhpa-Tēuctēpetl'') and Nauhcampatépetl, both meaning something like "Place of Four Mountains" or "Mountain of the Lord of Four Places", is an inactive volcano locate ...

and Cerro Volcancillo, a satellite peak of Cofre de Perote.

Etymology

The

specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...

''naucampatepetl'' is

Nahuatl Nahuatl ( ; ), Aztec, or Mexicano is a language or, by some definitions, a group of languages of the Uto-Aztecan language family. Varieties of Nahuatl are spoken by about Nahuas, most of whom live mainly in Central Mexico and have smaller popul ...

name for Cofre de Perote.

Description

Adult males measure and females up to in snout–vent length (SVL). The tail is slender and shorter than SVL; it tapers gradually but has a blunt tip. The body is moderately robust. The head is prominent and the eyes are large and relatively protuberant. The snout is large and broadly rounded. The limbs are long and robust. The digits are well developed, and there is no appreciable webbing. The coloration is striking, with solid black background color and with bright pink to pinkish-cream dorsal spots: there is a pair of rounded spots on the back of the head, about the size of the eyeball in diameter, a small mid-dorsal spot in the neck, and a pair of large spots at the level of the forelimbs. These larger spots are followed by 11 pairs of small spots. Finally, there is a conspicuous U-shaped mark behind the hips, pointing backward. The venter is pale gray to dark gray.

Habitat and conservation

Its natural

habitat In ecology, habitat refers to the array of resources, biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species' habitat can be seen as the physical manifestation of its ...

is pine-oak forest at elevations of

above sea level Height above mean sea level is a measure of a location's vertical distance (height, elevation or altitude) in reference to a vertical datum based on a historic mean sea level. In geodesy, it is formalized as orthometric height. The zero level ...

, with plenty of

bunch grass Tussock grasses or bunch grasses are a group of grass species in the family Poaceae. They usually grow as singular plants in clumps, tufts, hummocks, or bunches, rather than forming a sod or lawn, in meadows, grasslands, and prairies. As perennia ...

. All specimens in the

type series In biology, a type is a particular specimen (or in some cases a group of specimens) of an organism to which the scientific name of that organism is formally associated. In other words, a type is an example that serves to anchor or centralizes t ...

were found on roadside banks, under a surface layer of moist soil with a somewhat dry outer crust. This species is known from only six specimens. An individual was photographed in 2015, but surveys have failed to locate the species. Extensive logging, farming, and expanding human settlements have led to loss of much of the original habitat, and what remains is very degraded. The

IUCN SSC The International Union for Conservation of Nature (IUCN) is an international organization working in the field of nature conservation and sustainable use of natural resources. Founded in 1948, IUCN has become the global authority on the status ...

Amphibian Specialist Group considers ''Isthmura naucampatepetl'' critically endangered.

References

{{Taxonbar, from=Q28007374 naucampatepetl EDGE species Endemic amphibians of Mexico Fauna of the Trans-Mexican Volcanic Belt Amphibians described in 2001 Taxa named by David B. Wake Taxonomy articles created by Polbot