''Isoetes macrospora'', the big-spore quillwort, is a species of quillwort in the

Isoetaceae Isoetaceae is a family including living quillworts ('' Isoetes'') and comparable extinct herbaceous lycopsids ('' Tomiostrobus''). References External links * * Lycophytes Plant families {{Lycophyte-stub ...

family. It can be found in the deep water of low nutrient lakes in the Precambrian Shield as well as in

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Nova Scotia Nova Scotia is a Provinces and territories of Canada, province of Canada, located on its east coast. It is one of the three Maritime Canada, Maritime provinces and Population of Canada by province and territory, most populous province in Atlan ...

Quebec Quebec is Canada's List of Canadian provinces and territories by area, largest province by area. Located in Central Canada, the province shares borders with the provinces of Ontario to the west, Newfoundland and Labrador to the northeast, ...

, and

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. In the United States, it has been found in

Minnesota Minnesota ( ) is a U.S. state, state in the Upper Midwestern region of the United States. It is bordered by the Canadian provinces of Manitoba and Ontario to the north and east and by the U.S. states of Wisconsin to the east, Iowa to the so ...

and south, through the

Appalachian Mountains The Appalachian Mountains, often called the Appalachians, are a mountain range in eastern to northeastern North America. The term "Appalachian" refers to several different regions associated with the mountain range, and its surrounding terrain ...

Virginia Virginia, officially the Commonwealth of Virginia, is a U.S. state, state in the Southeastern United States, Southeastern and Mid-Atlantic (United States), Mid-Atlantic regions of the United States between the East Coast of the United States ...

. It bears 3 to 17 long, stiff dark green leaves, sometimes with recurving tips. The

sporangium A sporangium (from Late Latin, ; : sporangia) is an enclosure in which spores are formed. It can be composed of a unicellular organism, single cell or can be multicellular organism, multicellular. Virtually all plants, fungus, fungi, and many ot ...

can be 5 millimeters long and 4 millimeters wide, covered from one sixth to one quarter by the velum. The triangular

ligule A ligule (from "strap", variant of ''lingula'', from ''lingua'' "tongue") is a thin outgrowth at the junction of leaf A leaf (: leaves) is a principal appendage of the plant stem, stem of a vascular plant, usually borne laterally above g ...

can grow up to 2 millimeters long. The spherical, white megaspores are 400 to 800 micrometers in diameter, and bear ridges that form honeycomb-like areas. The kidney-shaped microspores are 32 to 50 micrometers long, each with evenly spaced smooth papillae.

References

macrospora Flora of Northern America {{Lycophyte-stub