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''Isoetes histrix'', the land quillwort, is an aquatic
pteridophyte A pteridophyte is a vascular plant (with xylem and phloem) that disperses spores. Because pteridophytes produce neither flowers nor seeds, they are sometimes referred to as " cryptogams", meaning that their means of reproduction is hidden. Ferns, ...
native to the
Mediterranean region In biogeography, the Mediterranean Basin (; also known as the Mediterranean Region or sometimes Mediterranea) is the region of lands around the Mediterranean Sea that have mostly a Mediterranean climate, with mild to cool, rainy winters and ...
, northwestern
Africa Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area ...
, and the coasts of western
Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a subcontinent of Eurasia and it is located enti ...
northwest to
Cornwall Cornwall (; kw, Kernow ) is a historic county and ceremonial county in South West England. It is recognised as one of the Celtic nations, and is the homeland of the Cornish people. Cornwall is bordered to the north and west by the Atlan ...
. It occurs mainly in temporarily wet habitats, otherwise called
vernal pool Vernal pools, also called vernal ponds or ephemeral pools, are seasonal pools of water that provide habitat for distinctive plants and animals. They are considered to be a distinctive type of wetland usually devoid of fish, and thus allow the safe ...
s. The leaves are 3–8 cm long, and summer-deciduous. It differs from '' Isoetes durieui'' by its large black scales between the leaves, though these are not always present, and a surface of megaspores with tubercles, whereas ''I. durieui'' has a network of ridges.


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Flora Europaea: ''Isoetes histrix''
histrix Flora of England Flora of Malta {{Lycophyte-stub