Iris Paradoxa
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''Iris paradoxa'' is a species of
flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (). The term angiosperm is derived from the Ancient Greek, Greek words (; 'container, vessel') and (; 'seed'), meaning that the seeds are enclosed with ...
native to western Asia. It has large upright petals and smaller lower petals, which is unique amongst most iris forms. They come in various shades from white, lavender, mauve, medium purple, violet, dark purple to black. It has a black or purplish black beard on the lower petals. It comes from the region of
Transcaucasia The South Caucasus, also known as Transcaucasia or the Transcaucasus, is a geographical region on the border of Eastern Europe and West Asia, straddling the southern Caucasus Mountains. The South Caucasus roughly corresponds to modern Armenia, ...
, and is found in the countries of
Iran Iran, officially the Islamic Republic of Iran (IRI) and also known as Persia, is a country in West Asia. It borders Iraq to the west, Turkey, Azerbaijan, and Armenia to the northwest, the Caspian Sea to the north, Turkmenistan to the nort ...
,
Turkey Turkey, officially the Republic of Türkiye, is a country mainly located in Anatolia in West Asia, with a relatively small part called East Thrace in Southeast Europe. It borders the Black Sea to the north; Georgia (country), Georgia, Armen ...
,
Armenia Armenia, officially the Republic of Armenia, is a landlocked country in the Armenian Highlands of West Asia. It is a part of the Caucasus region and is bordered by Turkey to the west, Georgia (country), Georgia to the north and Azerbaijan to ...
and in
Azerbaijan Azerbaijan, officially the Republic of Azerbaijan, is a Boundaries between the continents, transcontinental and landlocked country at the boundary of West Asia and Eastern Europe. It is a part of the South Caucasus region and is bounded by ...
.


Description

''I. paradoxa'' is a
rhizomatous In botany and dendrology, a rhizome ( ) is a modified subterranean plant stem that sends out roots and shoots from its nodes. Rhizomes are also called creeping rootstalks or just rootstalks. Rhizomes develop from axillary buds and grow hori ...
herbaceous
perennial In horticulture, the term perennial ('' per-'' + '' -ennial'', "through the year") is used to differentiate a plant from shorter-lived annuals and biennials. It has thus been defined as a plant that lives more than 2 years. The term is also ...
. The rhizomes are slender and usually less than 1 cm in diameter.British Iris Society (1997) It also has secondary roots underneath the rhizomes. It has greenish-gray, or blue-green leaves, that are recurved or falcate (sickle-like) shaped. They are narrow, between wide, and less than 4 cm long. The upright stem grows between long, and hold up to 2 blooms. It blooms in spring, or early summer, between April and June. The flowers are 10 cm in diameter, and come in various shades from white, lavender, mauve, medium purple, violet, dark purple to black. Like other irises, the flowers have 2 pairs of petals, 3 large
sepals A sepal () is a part of the flower of angiosperms (flowering plants). Usually green, sepals typically function as protection for the flower in bud, and often as support for the petals when in bloom., p. 106 Etymology The term ''sepalum'' ...
(outer petals), known as the 'falls' and 3 inner, smaller petals (or
tepals A tepal is one of the outer parts of a flower (collectively the perianth). The term is used when these parts cannot easily be classified as either sepals or petals. This may be because the parts of the perianth are undifferentiated (i.e. of ve ...
, known as the 'standards'. Compared to other irises, the paradoxa or strangeness of the iris, is that in most forms of irises, the standards are smaller than the falls, but on ''I. paradoxa'' the falls are much smaller than the standards. The erect standards are broadly obovate, or rounded, and long and wide. They are a pale shade with pale blue or deep blue veining. The smaller falls are a darker colour, blackish-violet, through to pale purple densely overlaid with black-purple veins. They are long and wide. The falls are also covered with blackish, blackish-purple or violet hairs or 'beard' with a pale v-shaped mark in centre. After flowering, it becomes dormant in late summer. The seed capsule produced after flower has not been described.


Biochemistry

As most irises are
diploid Ploidy () is the number of complete sets of chromosomes in a cell, and hence the number of possible alleles for autosomal and pseudoautosomal genes. Here ''sets of chromosomes'' refers to the number of maternal and paternal chromosome copies, ...
, meaning having two sets of
chromosomes A chromosome is a package of DNA containing part or all of the genetic material of an organism. In most chromosomes, the very long thin DNA fibers are coated with nucleosome-forming packaging proteins; in eukaryotic cells, the most importa ...
. This can be used to identify hybrids and classification of groupings. ''Iris paradoxa'' has been counted as 2n=20 .


Taxonomy

''Iris paradoxa'' is classified in the
subgenus In biology, a subgenus ( subgenera) is a taxonomic rank directly below genus. In the International Code of Zoological Nomenclature, a subgeneric name can be used independently or included in a species name, in parentheses, placed between the ge ...
''Iris'' sub. ''Iris'', ''Iris'' sect. ''Oncocyclus''. There is a known
form Form is the shape, visual appearance, or configuration of an object. In a wider sense, the form is the way something happens. Form may also refer to: *Form (document), a document (printed or electronic) with spaces in which to write or enter dat ...
from Turkey called ''I. paradoxa'' f. ''choschab''. The Latin
specific epithet In Taxonomy (biology), taxonomy, binomial nomenclature ("two-term naming system"), also called binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, both of which use Latin gramm ...
''paradoxa'' refers to the
Greek Greek may refer to: Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group *Greek language, a branch of the Indo-European language family **Proto-Greek language, the assumed last common ancestor of all kno ...
word for 'unusual', This is due to being thought being entirely unlike any other species. It was discovered in the
Caucus A caucus is a group or meeting of supporters or members of a specific political party or movement. The exact definition varies between different countries and political cultures. The term originated in the United States, where it can refer to ...
region, and then was first published and described by
Steven Stephen or Steven is an English first name. It is particularly significant to Christians, as it belonged to Saint Stephen ( ), an early disciple and deacon who, according to the Book of Acts, was stoned to death; he is widely regarded as the firs ...
in 'Mémoires de la Société Impériale des Naturalistes de Moscou' (Mém. Soc. Imp. Naturalistes Moscou) Vol.5 on page 355 in 1817. It is occasionally called the 'Unusual iris', or 'Velvet Iris' in America.


Forms

There is a known
form Form is the shape, visual appearance, or configuration of an object. In a wider sense, the form is the way something happens. Form may also refer to: *Form (document), a document (printed or electronic) with spaces in which to write or enter dat ...
called ''I. paradoxa'' f. ''choschab'' . It was published in Fl. Iran. Vol.112 on page 31 in 1975. Originally ''Iris paradoxa'' f. ''choschab'' was thought by
Johannes Marius Cornelis Hoog Johannes is a Medieval Latin form of the personal name that usually appears as "John" in English language contexts. It is a variant of the Greek and Classical Latin variants (Ιωάννης, '' Ioannes''), itself derived from the Hebrew name '' Ye ...
to be a variety of ''I. paradoxa'', called ''I. paradoxa'' var. ''choschab'' and then published in Gard. Chron. III, 29: 104 in 1901, before being downgraded to a form in 1975. It can be found near
Lake Van Lake Van (; ; ) is the largest lake in Turkey. It lies in the Eastern Anatolia Region of Turkey in the provinces of Van Province, Van and Bitlis Province, Bitlis, in the Armenian highlands. It is a Salt lake, saline Soda lake, soda lake, receiv ...
, close to the town of
Başkale Başkale (, ) is a municipality and district of Van Province, Turkey. Its area is 2,727 km2, and its population is 44,168 (2022). The municipality Başkale was established in 1937. In the local elections of March 2019 Erkan Acar from the ...
, and it is common around the village of Hoşap, known for its
Hoşap Castle Hoşap Castle (, , ) is a large 17th-century castle located in the village of Hoşab, Gürpınar District, Van, Turkey. It is at a distance of approximately 50 km to the city center of Van. History The castle was built upon the founda ...
,
Gürpınar District Gürpınar may refer to: Places *Gürpınar, Van, a district and municipality in Van Province * Gürpınar, Alaplı, a village in Zonguldak Province * Gürpınar, Bartın, a village in Bartın Province * Gürpınar, Çivril, a neighbourhood in Deniz ...
,
Van Province Van Province (, , Armenian: Վանի մարզ) is a province and metropolitan municipality in the Eastern Anatolian region of Turkey, between Lake Van and the Iranian border. Its area is 20,921 km2, and its population is 1,128,749 (2022). ...
, Eastern Anatolia, Turkey. 'Choschab' was an old version of the
Aramaic Aramaic (; ) is a Northwest Semitic language that originated in the ancient region of Syria and quickly spread to Mesopotamia, the southern Levant, Sinai, southeastern Anatolia, and Eastern Arabia, where it has been continually written a ...
name of the castle, meaning 'good water' or 'beautiful water'. The village of Hoşap is now known by the Turkish name of 'Güzelsu'. The form is thought to be most used in cultivation. It has white standards, or pale lilac, lightly veined with blue. The beard is black and covers the upper half of the falls and the style arms are cream, speckled reddish brown. The internal tepals ("petals") are lined with purple on a white (choschab form) or purple to mauve (paradoxa form) background.


Ecology

Males of ''
Xylocopa valga ''Xylocopa valga'' is a species of carpenter bee common to: western, central and southern Europe, except for far northern latitudes; the Caucasus; Middle East; Central Asia; and Mongolia. The species has become extinct in Latvia and Lithuania. D ...
'' (a type of carpenter bee) have been observed pseudo-copulating on the reduced, velvety petals of the plant in Leriksky area, near Gosmaljan, Azerbaijan.


Distribution and habitat

It is
native Native may refer to: People * '' Jus sanguinis'', nationality by blood * '' Jus soli'', nationality by location of birth * Indigenous peoples, peoples with a set of specific rights based on their historical ties to a particular territory ** Nat ...
to temperate
Western Asia West Asia (also called Western Asia or Southwest Asia) is the westernmost region of Asia. As defined by most academics, UN bodies and other institutions, the subregion consists of Anatolia, the Arabian Peninsula, Iran, Mesopotamia, the Armenian ...
. It is found in the region of
Transcaucasia The South Caucasus, also known as Transcaucasia or the Transcaucasus, is a geographical region on the border of Eastern Europe and West Asia, straddling the southern Caucasus Mountains. The South Caucasus roughly corresponds to modern Armenia, ...
, and in the countries of
Iran Iran, officially the Islamic Republic of Iran (IRI) and also known as Persia, is a country in West Asia. It borders Iraq to the west, Turkey, Azerbaijan, and Armenia to the northwest, the Caspian Sea to the north, Turkmenistan to the nort ...
, (formerly
Persia Iran, officially the Islamic Republic of Iran (IRI) and also known as Persia, is a country in West Asia. It borders Iraq to the west, Turkey, Azerbaijan, and Armenia to the northwest, the Caspian Sea to the north, Turkmenistan to the nort ...
),
Turkey Turkey, officially the Republic of Türkiye, is a country mainly located in Anatolia in West Asia, with a relatively small part called East Thrace in Southeast Europe. It borders the Black Sea to the north; Georgia (country), Georgia, Armen ...
,
Armenia Armenia, officially the Republic of Armenia, is a landlocked country in the Armenian Highlands of West Asia. It is a part of the Caucasus region and is bordered by Turkey to the west, Georgia (country), Georgia to the north and Azerbaijan to ...
and in
Azerbaijan Azerbaijan, officially the Republic of Azerbaijan, is a Boundaries between the continents, transcontinental and landlocked country at the boundary of West Asia and Eastern Europe. It is a part of the South Caucasus region and is bounded by ...
, (including in the Nakhchivan region). It grows on arid, stony hillsides and they can be found at altitude between above sea level.


Cultivation

''I. paradoxa'' can be easily grown in an
alpine house A greenhouse is a structure that is designed to regulate the temperature and humidity of the environment inside. There are different types of greenhouses, but they all have large areas covered with transparent materials that let sunlight pass an ...
, or they can be placed in
rock gardens A rock garden, also known as a rockery and formerly as a rockwork, is a garden, or more often a part of a garden, with a landscaping framework of rocks, stones, and gravel, with planting appropriate to this setting. Usually these are small ...
, which should have a summer drought. They grow well in full sun with mildly acidic to mildly alkaline soils (of ph levels between 6.1–7.8). The flower and stem (if removed from the plant) may have a shelf life of two to three days. Irises can generally be propagated by division, or by seed growing. ''I. paradoxa'' has been listed with '' Iris orientalis'' and ''Iris spuria'' subsp. ''musulmanica'' as
halophyte A halophyte is a salt-tolerant plant that grows in soil or waters of high salinity, coming into contact with saline water through its roots or by salt spray, such as in saline semi-deserts, mangrove swamps, marshes and sloughs, and seashores. ...
(salt-tolerant) plants.


Cultivars and crosses

''Iris paradoxa'' has been used in various breeding programmes and various
cultivars A cultivar is a kind of cultivated plant that people have selected for desired traits and which retains those traits when propagated. Methods used to propagate cultivars include division, root and stem cuttings, offsets, grafting, tissue cult ...
have been produced including: 'Atrata', 'Choschab', 'Jolfa Form', 'Medwedewi', 'Mirabilis', 'Paradoxa' and 'Paradoxa Albo-Lutescens'. Various ''Iris paradoxa'' crossed Hybrids have also been produced; * ''Iris paradoxa'' X ''
Iris pallida ''Iris pallida'', the Dalmatian iris or sweet iris, is a hardy flowering perennial plant of the genus '' Iris'', family Iridaceae. It is native to the Dalmatian coast (Croatia) but widely naturalised elsewhere. It is a member of the subgenus '' I ...
'' : 'Alkmene', 'Hamadan', 'Lady Lilford' and 'Parpall' * ''Iris paradoxa'' X ''
Iris variegata ''Iris variegata'', commonly known as the Hungarian iris, is a plant species in the genus ''Iris'', also in the subgenus ''Iris''. It is a rhizomatous perennial from eastern Europe. It has dark green, ribbed leaves. The branched flowering stems c ...
'' : 'Parvar' and 'Shadrach' ''Iris paradoxa'' crossed with Section Oncocyclus Irises: *''Iris paradoxa'' and ''
Iris acutiloba ''Iris acutiloba'' is a species in the genus '' Iris'', it is also in the subgenus of '' Iris'' and section ''Oncocyclus''. It is a rhizomatous perennial, from the mountains of the Caucasus and found in Turkey, Armenia Armenia, officially ...
'' : 'Zuvendicus' *''
Iris camillae ''Iris camillae '' is a species in the genus ''Iris'', it is also in the subgenus ''Iris'' and in the section ''Oncocyclus''. It is a rhizomatous perennial, from Azerbaijan. It has narrow, falcate (sickle-shaped) leaves, medium-sized stem and la ...
'' X ''Iris paradoxa'': 'First Sergeant' *''
Iris iberica ''Iris iberica'' is a plant species in the genus ''Iris'', it is also in the subgenus ''Iris'' and in the section ''Oncocyclus''. It is a rhizomatous perennial, from the Caucasus Mountains of Armenia, eastern Georgia, and western Azerbaijan. It ...
'' X ''Iris paradoxa'': 'Ib-Parad', 'Koenigii' and 'Paradib' Iris paradoxa crossed with Section Regelia Irises : * ''Iris paradoxa'' X '' Iris korolkowii'' : 'Parkor', 'Camilla', 'Clytemnestra', 'Sirona' and 'Minerva', * ¼ ''Iris paradoxa'' crosses 'Cactiforium', 'Charming Chick', 'Clotho', 'Dear Me', 'Don Ricardo', 'Emily Pyke' and 'Regal Plus'


Toxicity

Like many other irises, most parts of the plant are poisonous (including rhizome and leaves), if mistakenly ingested, it can cause stomach pains and vomiting. Also handling the plant may cause a skin irritation or an allergic reaction.David G Spoerke and Susan C. Smolinske


References

* Gartenflora 12: 16. Tab. 386. 1863, illustrated in color. * Krel. 1892; 1913; Van W. 1906; Hocker 1938; * F.C.C., R.H.S. 1900, shown by Van T.; Hort Dir. 42: 60. 1901;


External links

* http://balearicgarden.es/iris/oncocyclus/ Has images of various Oncocyclus Irises * https://photos.v-d-brink.eu/Flora-and-Fauna/Asia/Iran-Elburz-mountains-Western/i-3BNnLQm/ {{Taxonbar, from=Q12843821 paradoxa Flora of Iran Flora of Armenia Flora of Azerbaijan Flora of Turkey Plants described in 1980 Garden plants