Intrafusal Muscle Fiber
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Intrafusal muscle fibers are skeletal muscle fibers that serve as specialized
sensory organs A sense is a biological system used by an organism for sensation, the process of gathering information about the surroundings through the detection of stimuli. Although, in some cultures, five human senses were traditionally identified as s ...
( proprioceptors). They detect the amount and rate of change in length of a
muscle Muscle is a soft tissue, one of the four basic types of animal tissue. There are three types of muscle tissue in vertebrates: skeletal muscle, cardiac muscle, and smooth muscle. Muscle tissue gives skeletal muscles the ability to muscle contra ...
.Casagrand, Janet (2008) ''Action and Movement: Spinal Control of Motor Units and Spinal Reflexes.'' University of Colorado, Boulder. They constitute the muscle spindle, and are innervated by both sensory (afferent) and motor (efferent) fibers. Intrafusal muscle fibers are not to be confused with extrafusal muscle fibers, which contract, generating skeletal movement and are innervated by alpha motor neurons.


Structure


Types

There are two types of intrafusal muscle fibers: nuclear bag fibers and nuclear chain fibers. They bear two types of sensory ending, known as annulospiral and flower-spray endings. Both ends of these fibers contract, but the central region only stretches and does not contract. Intrafusal muscle fibers are walled off from the rest of the muscle by an outer
connective tissue Connective tissue is one of the four primary types of animal tissue, a group of cells that are similar in structure, along with epithelial tissue, muscle tissue, and nervous tissue. It develops mostly from the mesenchyme, derived from the mesod ...
sheath consisting of flattened fibroblasts and
collagen Collagen () is the main structural protein in the extracellular matrix of the connective tissues of many animals. It is the most abundant protein in mammals, making up 25% to 35% of protein content. Amino acids are bound together to form a trip ...
. This sheath has a spindle or "fusiform" shape, hence the name "intrafusal".


Innervation

They are innervated by gamma motor neurons and beta motor neurons. Gamma efferents from small
multipolar neuron A multipolar neuron is a type of neuron that possesses a single axon and many dendrites (and dendritic branches), allowing for the integration of a great deal of information from other neurons. These processes are projections from the neuron cel ...
s from anterior gray column innervate it. These form a part of neuromuscular spindles.


Function

Intrafusal muscle fibers detect the amount and rate of change in muscle length. It is by the sensory information from gamma motor neurons and beta motor neurons that an individual is able to judge the position of their muscles.


See also

* Alpha motor neuron * Beta motor neuron * Extrafusal muscle fiber * Gamma motor neuron * Type Ia sensory fiber * Type II sensory fiber


References


External links


Histology at ucsd.edu


* {{Authority control Muscular system