''Maxillaria'', abbreviated as Max in the

horticultural Horticulture (from ) is the art and science of growing fruits, vegetables, flowers, trees, shrubs and ornamental plants. Horticulture is commonly associated with the more professional and technical aspects of plant cultivation on a smaller and mo ...

trade, is a large

genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...

orchid Orchids are plants that belong to the family Orchidaceae (), a diverse and widespread group of flowering plants with blooms that are often colourful and fragrant. Orchids are cosmopolitan plants that are found in almost every habitat on Eart ...

s (family Orchidaceae). This is a diverse genus, with very different morphological forms. Their characteristics can vary widely. They are commonly called spider orchids, flame orchids or tiger orchids. Their scientific name is derived from the

Latin Latin ( or ) is a classical language belonging to the Italic languages, Italic branch of the Indo-European languages. Latin was originally spoken by the Latins (Italic tribe), Latins in Latium (now known as Lazio), the lower Tiber area aroun ...

word ''maxilla'', meaning

jawbone In jawed vertebrates, the mandible (from the Latin ''mandibula'', 'for chewing'), lower jaw, or jawbone is a bone that makes up the lowerand typically more mobilecomponent of the mouth (the upper jaw being known as the maxilla). The jawbone ...

, reflecting on the

column A column or pillar in architecture and structural engineering is a structural element that transmits, through compression, the weight of the structure above to other structural elements below. In other words, a column is a compression member ...

and the base of the

lip The lips are a horizontal pair of soft appendages attached to the jaws and are the most visible part of the mouth of many animals, including humans. Mammal lips are soft, movable and serve to facilitate the ingestion of food (e.g. sucklin ...

of some species, that may evoke a protruding jaw.

Description

Their

pseudobulb In botany, a pseudobulb is a storage organ found in many epiphytic and terrestrial sympodial orchids. It is derived from a thickening of the part of a stem between leaf nodes and may be composed of just one internode or several, termed ''heter ...

s are round or oblong and each carry one or two

lanceolate The following terms are used to describe leaf plant morphology, morphology in the description and taxonomy (biology), taxonomy of plants. Leaves may be simple (that is, the leaf blade or 'lamina' is undivided) or compound (that is, the leaf blade ...

leaves A leaf (: leaves) is a principal appendage of the stem of a vascular plant, usually borne laterally above ground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, stem, ...

. Some grow close together in a clustered manner on a short

rhizome In botany and dendrology, a rhizome ( ) is a modified subterranean plant stem that sends out roots and Shoot (botany), shoots from its Node (botany), nodes. Rhizomes are also called creeping rootstalks or just rootstalks. Rhizomes develop from ...

, while in other species the pseudobulbs keep some distance on an elongate rhizome. This rhizome is clothed in a somewhat transparent, silvery-gray

velamen The velamen or velamen radicum ("covering of the roots" in Latin) is a spongy, multiple epidermis that covers the roots of some epiphytic or semi-epiphytic plants, such as orchid and ''Clivia'' species. The velamen of an orchid is the white or gr ...

. The

flower Flowers, also known as blooms and blossoms, are the reproductive structures of flowering plants ( angiosperms). Typically, they are structured in four circular levels, called whorls, around the end of a stalk. These whorls include: calyx, m ...

s grow solitary on short stalks, called scapes, from the base of the pseudobulb. Most are small to very small, but some

species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...

carry large, showy flowers. The flowers are never longer than the leaves. Their free

petal Petals are modified leaves that form an inner whorl surrounding the reproductive parts of flowers. They are often brightly coloured or unusually shaped to attract pollinators. All of the petals of a flower are collectively known as the ''corol ...

s and

sepal A sepal () is a part of the flower of angiosperms (flowering plants). Usually green, sepals typically function as protection for the flower in bud, and often as support for the petals when in bloom., p. 106 Etymology The term ''sepalum'' ...

s have a typically curved and adnate labellum with three inconspicuous lobes. Or the lip may have a distinct

callus A callus (: calluses) is an area of thickened and sometimes hardened skin that forms as a response to repeated friction, pressure, or other irritation. Since repeated contact is required, calluses are most often found on the feet and hands, b ...

on the disc ( = central part of the lip from which the lobes radiate). The papillae (= small warts like glands) and the

trichome Trichomes (; ) are fine outgrowths or appendages on plants, algae, lichens, and certain protists. They are of diverse structure and function. Examples are hairs, glandular hairs, scales, and papillae. A covering of any kind of hair on a plant ...

s of the lip show great diversity. The most common form for the papillae is the conical form with rounded or pointed tips.

Taxonomy

The genus was established by

Hipólito Ruiz López Hipólito Ruiz López (August 8, 1754 in Belorado, Burgos, Spain – 1816 in Madrid), or Hipólito Ruiz, was a Spanish botanist known for researching the floras of Peru and Chile during an expedition under Charles III of Spain, Carlos III from 17 ...

and

José Antonio Pavón Jiménez José Antonio Pavón Jiménez or José Antonio Pavón (April 22, 1754 in Casatejada, Cáceres, Spain – 1840 in Madrid) was a Spanish botanist known for researching the flora of Peru and Chile. Biography During the reign of Charles III of ...

in 1794. In 2007, a

molecular phylogenetic Molecular phylogenetics () is the branch of phylogeny that analyzes genetic, hereditary molecular differences, predominantly in DNA sequences, to gain information on an organism's evolutionary relationships. From these analyses, it is possible to ...

study found that many of the genera then accepted in the subtribe

Maxillariinae Maxillariinae is an orchid subtribe in the tribe Cymbidieae. It was formerly treated as the tribe Maxillarieae, and divided into a number of subtribes. Genera Generic boundaries in the tribe have changed substantially with new molecular evidenc ...

(including ''Anthosiphon'', ''Cryptocentrum'', ''Chrysocycnis'', ''Cyrtidiorchis'', ''Mormolyca'', ''Pityphyllum'', and ''Trigonidium'') were embedded within ''Maxillaria'' as then

circumscribed In geometry, a circumscribed circle for a set of points is a circle passing through each of them. Such a circle is said to ''circumscribe'' the points or a polygon formed from them; such a polygon is said to be ''inscribed'' in the circle. * Circum ...

, rendering that genus

polyphyletic A polyphyletic group is an assemblage that includes organisms with mixed evolutionary origin but does not include their most recent common ancestor. The term is often applied to groups that share similar features known as Homoplasy, homoplasies ...

. The authors suggested recognizing 17 separate genera, including in addition ''Brasilorchis'', ''Camaridium'', ''Christensonella'', ''Heterotaxis'', ''Inti'', ''Mapinguari'', ''Maxillariella'', ''Nitobulbon'', ''Ornithidium'', ''Rhetinantha'' and ''Sauvetrea''.Pridgeon, A.M., Cribb, P.J., Chase, M.C. & Rasmussen, F.N. (2009). Epidendroideae (Part two). Genera Orchidacearum 5: 1-585. Oxford University Press, New York, Oxford. Other sources have preferred to take a more conservative approach, treating all the genera within an enlarged ''Maxillaria''. For example, ,

Plants of the World Online Plants of the World Online (POWO) is an online taxonomic database published by the Royal Botanic Gardens, Kew. History Following the Convention on Biological Diversity, the Royal Botanic Gardens in Kew launched Plants of the World Online i ...

listed over 40 other genus names as

synonyms A synonym is a word, morpheme, or phrase that means precisely or nearly the same as another word, morpheme, or phrase in a given language. For example, in the English language, the words ''begin'', ''start'', ''commence'', and ''initiate'' are a ...

of ''Maxillaria''.

Distribution and ecology

''Maxillaria'' species are distributed in the

rainforest Rainforests are forests characterized by a closed and continuous tree Canopy (biology), canopy, moisture-dependent vegetation, the presence of epiphytes and lianas and the absence of wildfire. Rainforests can be generally classified as tropi ...

at sea level to elevations of 3,500 m, in

Latin America Latin America is the cultural region of the Americas where Romance languages are predominantly spoken, primarily Spanish language, Spanish and Portuguese language, Portuguese. Latin America is defined according to cultural identity, not geogr ...

from central

Mexico Mexico, officially the United Mexican States, is a country in North America. It is the northernmost country in Latin America, and borders the United States to the north, and Guatemala and Belize to the southeast; while having maritime boundar ...

Bolivia Bolivia, officially the Plurinational State of Bolivia, is a landlocked country located in central South America. The country features diverse geography, including vast Amazonian plains, tropical lowlands, mountains, the Gran Chaco Province, w ...

, as well as in the

West Indies The West Indies is an island subregion of the Americas, surrounded by the Atlantic Ocean, North Atlantic Ocean and the Caribbean Sea, which comprises 13 independent island country, island countries and 19 dependent territory, dependencies in thr ...

.Kew World Checklist of Selected Plant Families
/ref> This is an indication for the different temperature requirements, from warm growing to cold growing, within the genus. They are mostly

epiphyte An epiphyte is a plant or plant-like organism that grows on the surface of another plant and derives its moisture and nutrients from the air, rain, water (in marine environments) or from debris accumulating around it. The plants on which epiphyt ...

s, rather large in size, but some are

terrestrial Terrestrial refers to things related to land or the planet Earth, as opposed to extraterrestrial. Terrestrial may also refer to: * Terrestrial animal, an animal that lives on land opposed to living in water, or sometimes an animal that lives on o ...

s or even

lithophyte Lithophytes are plants that grow in or on rocks. They can be classified as either epilithic (or epipetric) or endolithic; epilithic lithophytes grow on the surfaces of rocks, while endolithic lithophytes grow in the crevices of rocks (and are als ...

s (such as ''M. rupestris'').

Cultivation

''Maxillaria'' is not one of the popular genera among growers. Only a few species grow big, showy flowers. But some species are nevertheless sought by collectors, mostly for the

fragrance An aroma compound, also known as an odorant, aroma, fragrance, flavoring or flavor, is a chemical compound that has a smell or odor. For an individual chemical or class of chemical compounds to impart a smell or fragrance, it must be sufficien ...

of their blossoms, such as ''

Maxillaria tenuifolia ''Maxillaria tenuifolia'', the delicate-leafed maxillaria or coconut pie orchid, is a species of orchid ranging from Mexico to Nicaragua and possibly Costa Rica. These plants are easy to grow if kept moist and given good air movement in a high-li ...

''.

References

*Dr. Karlheinz Senghas - ''Maxillaria, un genre chaotique'' - Richardiana *Eric A. Christenson - ''Vue d’ensemble du genre Maxillaria'' - Richardiana

External links

* * {{Taxonbar, from=Q2719986 Maxillariinae genera Epiphytic orchids Taxa named by José Antonio Pavón Jiménez Taxa named by Hipólito Ruiz López