Scent gland are

exocrine gland Exocrine glands are glands that secrete substances onto an epithelial surface by way of a duct. Examples of exocrine glands include sweat, salivary, mammary, ceruminous, lacrimal, sebaceous, prostate and mucous. Exocrine glands are one of ...

s found in most

mammal A mammal () is a vertebrate animal of the Class (biology), class Mammalia (). Mammals are characterised by the presence of milk-producing mammary glands for feeding their young, a broad neocortex region of the brain, fur or hair, and three ...

s. They produce semi-viscous

secretion Secretion is the movement of material from one point to another, such as a secreted chemical substance from a cell or gland. In contrast, excretion is the removal of certain substances or waste products from a cell or organism. The classical mec ...

s which contain

pheromone A pheromone () is a secreted or excreted chemical factor that triggers a social response in members of the same species. Pheromones are chemicals capable of acting like hormones outside the body of the secreting individual, to affect the behavio ...

s and other

semiochemical A semiochemical, from the Greek wiktionary:σημεῖον, σημεῖον (''semeion''), meaning "signal", is a chemical substance or mixture released by an organism that affects the behaviors of other individuals. Semiochemical communication c ...

compounds. These odor-messengers indicate information such as status,

territorial marking In ethology, territory is the sociographical area that an animal consistently defends against conspecific competition (or, occasionally, against animals of other species) using agonistic behaviors or (less commonly) real physical aggression. ...

, mood, and sexual behaviour. The odor may be subliminal—not consciously detectable. Though it is not their primary function, the

salivary gland The salivary glands in many vertebrates including mammals are exocrine glands that produce saliva through a system of ducts. Humans have three paired major salivary glands ( parotid, submandibular, and sublingual), as well as hundreds of min ...

s may also function as scent glands in some animals.

In even-toed ungulates

The

even-toed ungulate Artiodactyls are placental mammals belonging to the order Artiodactyla ( , ). Typically, they are ungulates which bear weight equally on two (an even number) of their five toes (the third and fourth, often in the form of a hoof). The other t ...

s (Artiodactyla) have many specialized skin glands, the

s of which are involved in semiochemical communication. These glands include the sudoriferous glands (located on the forehead, between the

antler Antlers are extensions of an animal's skull found in members of the Cervidae (deer) Family (biology), family. Antlers are a single structure composed of bone, cartilage, fibrous tissue, skin, nerves, and blood vessels. They are generally fo ...

s and eyes), the

preorbital gland The preorbital gland is a paired exocrine gland found in many species of artiodactyls, which is homologous to the lacrimal gland found in humans. These glands are trenchlike slits of dark blue to black, nearly bare skin extending from the media ...

s (extending from the medial

canthus The canthus (: canthi, palpebral commissures) is either corner of the eye where the upper and lower eyelids meet. More specifically, the inner and outer canthi are, respectively, the medial and lateral ends/angles of the palpebral fissure. The ...

of each eye), the nasal glands (located inside the nostrils), the interdigital glands (located between the toes), the preputial gland (located inside the foreskin of the penis), the metatarsal glands (located outside of the hind legs), the tarsal glands (located inside of the hind legs), and the inguinal glands in the lower belly or groin area. Like many other species of Artiodactyla,

deer A deer (: deer) or true deer is a hoofed ruminant ungulate of the family Cervidae (informally the deer family). Cervidae is divided into subfamilies Cervinae (which includes, among others, muntjac, elk (wapiti), red deer, and fallow deer) ...

have seven major external scent glands distributed throughout their bodies. Deer rely heavily on these scent glands to communicate with other members of their species, and possibly even with members of other species. For example, male

white-tailed deer The white-tailed deer (''Odocoileus virginianus''), also known Common name, commonly as the whitetail and the Virginia deer, is a medium-sized species of deer native to North America, North, Central America, Central and South America. It is the ...

(''Odocoileus virginianus'') are often seen working over a scrape. First, the animal scrapes at the dirt with its hooves, depositing the scent from his interdigital gland on the ground. After that, he may bite the tip off an overhanging branch, depositing secretions from his salivary glands onto the branch. He may then rub his face on the overhanging branch, depositing secretions from the sudoriferous and preorbital glands on it. The interdigital glands of male and female black-tailed deer contain three volatile ketones, 2-tridecanone, (''E'')-3-tridecen-2-one and (''E'')-4-tridecen-2-one. (''E'')-3-tridecen-2-one was shown to have antibiotic activity against some skin pathogens. These compounds are absent from white-tailed deer interdigital glands, which contain a number of 2-methylcarboxylic acids. The tarsal gland appears to operate by a different mechanism than the other external scent glands. A behavior called '' rub-urination'' is central to this mechanism. During rub-urination, the animal squats while urinating so that urine will run down the insides of its legs and onto its tarsal glands. The tarsal glands have a tuft of hair which is specially adapted to extract certain chemical compounds from the animal's urine. For example, in the

black-tailed deer Black-tailed deer or blacktail deer occupy coastal regions of western North America. There are two subspecies, the Columbian black-tailed deer (''Odocoileus hemionus columbianus'') which ranges from the Pacific Northwest of the United States and ...

(''Odocoileus hemionus columbianus''), the major constituent of the tarsal gland secretion is a

lipid Lipids are a broad group of organic compounds which include fats, waxes, sterols, fat-soluble vitamins (such as vitamins A, D, E and K), monoglycerides, diglycerides, phospholipids, and others. The functions of lipids include storing ...

, (Z)-6-dodecen-4-olide. This compound does not originate in the tarsal gland itself, but rather it is extracted from the animal's urine by the tarsal hair tuft during the rub-urination process. In white-tailed deer, the presence and concentration of certain chemical compounds in the urine depend on the season, gender, reproductive status and

social rank A social class or social stratum is a grouping of people into a set of Dominance hierarchy, hierarchical social categories, the most common being the working class and the Bourgeoisie, capitalist class. Membership of a social class can for exam ...

of the animals. This fact, along with the observation of rub-urination behavior in this animal (at least in the male) indicates that urine probably plays a role in

olfactory communication Olfactic communication is a channel of nonverbal communication referring to the various ways people and animals Communication, communicate and engage in social interaction through their sense of Sense of smell, smell. Our human Olfactory system ...

in deer.

In carnivorans

Canid Canidae (; from Latin, ''canis'', "dog") is a family (biology), biological family of caniform carnivorans, constituting a clade. A member of this family is also called a canid (). The family includes three subfamily, subfamilies: the Caninae, a ...

s have several scent glands that are used in

. The fossa has several scent glands. Like herpestids it has a perianal skin gland inside an anal sac which surrounds the anus like a pocket. The pocket opens to the exterior with a horizontal slit below the tail. Other glands are located near the penis or vagina, with the penile glands emitting a strong odor. Like the herpestids, it has no prescrotal glands.

Anal gland The anal glands or anal sacs are small glands near the anus in many mammals. They are situated in between the external anal sphincter muscle and internal anal sphincter muscle. In non-human mammals, the secretions of the anal glands contain most ...

s are found in all

carnivora Carnivora ( ) is an order of placental mammals specialized primarily in eating flesh, whose members are formally referred to as carnivorans. The order Carnivora is the sixth largest order of mammals, comprising at least 279 species. Carnivor ...

including

wolves The wolf (''Canis lupus''; : wolves), also known as the grey wolf or gray wolf, is a canine native to Eurasia and North America. More than thirty subspecies of ''Canis lupus'' have been recognized, including the dog and dingo, though gr ...

bear Bears are carnivoran mammals of the family (biology), family Ursidae (). They are classified as caniforms, or doglike carnivorans. Although only eight species of bears are extant, they are widespread, appearing in a wide variety of habitats ...

sea otter The sea otter (''Enhydra lutris'') is a marine mammal native to the coasts of the northern and eastern Pacific Ocean, North Pacific Ocean. Adult sea otters typically weigh between , making them the heaviest members of ...

s and kinkajous.

In other animals

Apocrine sweat gland An apocrine sweat gland (; from Greek ''apo'' 'away' and ''krinein'' 'to separate') is composed of a coiled secretory portion located at the junction of the dermis and subcutaneous fat, from which a straight portion inserts and secretes into th ...

s, such as in the armpits of humans *

Sebaceous gland A sebaceous gland or oil gland is a microscopic exocrine gland in the skin that opens into a hair follicle to secrete an oily or waxy matter, called sebum, which lubricates the hair and skin of mammals. In humans, sebaceous glands occur in ...

s, such as the

cranial Standard anatomical terms of location are used to describe unambiguously the anatomy of humans and other animals. The terms, typically derived from Latin or Greek language, Greek roots, describe something in its standard anatomical position. Thi ...

surface glands of the red-bellied lemur * Flank glands, such as in

vole Voles are small rodents that are relatives of lemmings and hamsters, but with a stouter body; a longer, hairy tail; a slightly rounder head; smaller eyes and ears; and differently formed molars (high-crowned with angular cusps instead of lo ...

s or

shrew Shrews ( family Soricidae) are small mole-like mammals classified in the order Eulipotyphla. True shrews are not to be confused with treeshrews, otter shrews, elephant shrews, West Indies shrews, or marsupial shrews, which belong to dif ...

s * Various glands producing pheromone signals found in non-avian reptiles, including

Amphisbaenia Amphisbaenia (called amphisbaenians or worm lizards) is a group of typically legless lizards, comprising over 200 extant species. Amphisbaenians are characterized by their long bodies, the reduction or loss of the limbs, and rudimentary eyes. A ...

ns,

lizard Lizard is the common name used for all Squamata, squamate reptiles other than snakes (and to a lesser extent amphisbaenians), encompassing over 7,000 species, ranging across all continents except Antarctica, as well as most Island#Oceanic isla ...

snake Snakes are elongated limbless reptiles of the suborder Serpentes (). Cladistically squamates, snakes are ectothermic, amniote vertebrates covered in overlapping scales much like other members of the group. Many species of snakes have s ...

s, and Crocodylians * Castor sacs, found in

beaver Beavers (genus ''Castor'') are large, semiaquatic rodents of the Northern Hemisphere. There are two existing species: the North American beaver (''Castor canadensis'') and the Eurasian beaver (''C. fiber''). Beavers are the second-large ...

s * Perineal glands, found in

viverrid Viverridae is a family (biology), family of small to medium-sized feliform mammals, comprising 14 genera with 33 species. This family was named and first described by John Edward Gray in 1821. Viverrids occur all over Africa, in southern Europe, ...

s, guinea pigs, and porcupines *

Preputial glands Preputial glands are exocrine glands in the prepuce in front of the penis. They occur in many mammals, including canids, mice, ferrets, rhinoceroses, and even-toed ungulates and produce pheromones. The preputial glands of female animals are so ...

, found in many species including mice, pigs, and wolves * Supracaudal glands, found in canidsFox, Michael W., and James A. Cohen.
Canid communication
" How animals communicate (1977): 728-748.

References

{{DEFAULTSORT:Scent Gland Glands Mammal anatomy

In even-toed ungulates

In carnivorans

In other animals

See also

References