''Inguza predemersus'' is an

extinct Extinction is the termination of an organism by the death of its Endling, last member. A taxon may become Functional extinction, functionally extinct before the death of its last member if it loses the capacity to Reproduction, reproduce and ...

species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...

penguin Penguins are a group of aquatic flightless birds from the family Spheniscidae () of the order Sphenisciformes (). They live almost exclusively in the Southern Hemisphere. Only one species, the Galápagos penguin, is equatorial, with a sm ...

. It was formerly placed in the

genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...

'' Spheniscus'' and presumed to be a close relative of the African penguin, but after its well-distinct

tarsometatarsus The tarsometatarsus is a bone that is only found in the lower leg of birds and some non-avian dinosaurs. It is formed from the fusion of several bird bones found in other types of animals, and homologous to the mammalian tarsus (ankle bones) a ...

was found, it was moved into its present

monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unisp ...

genus. The known fossils specimens were found in Late

Pliocene The Pliocene ( ; also Pleiocene) is the epoch (geology), epoch in the geologic time scale that extends from 5.33 to 2.58quarry A quarry is a type of open-pit mining, open-pit mine in which dimension stone, rock (geology), rock, construction aggregate, riprap, sand, gravel, or slate is excavated from the ground. The operation of quarries is regulated in some juri ...

at Langebaanweg,

South Africa South Africa, officially the Republic of South Africa (RSA), is the Southern Africa, southernmost country in Africa. Its Provinces of South Africa, nine provinces are bounded to the south by of coastline that stretches along the Atlantic O ...

, from about 5 million years ago. What is known from molecular data is that the time at which the present species lived is not too distant from the arrival of the ancestors of the African penguin on the Atlantic coasts of southern Africa. On the other hand, it may be closer to ''

Pygoscelis The genus ''Pygoscelis'' ("rump-legged") contains three living species of penguins collectively known as "brush-tailed penguins". Taxonomy Mitochondrial and nuclear DNA evidence suggests the genus split from other penguins around 38 milli ...

''. This would mean that its ancestors diverged from those of the extant ''Pygoscelis'' most likely at an indeterminate point of time during the

Oligocene The Oligocene ( ) is a geologic epoch (geology), epoch of the Paleogene Geologic time scale, Period that extends from about 33.9 million to 23 million years before the present ( to ). As with other older geologic periods, the rock beds that defin ...

. Alternatively, it might not be close to extant penguins (the Spheniscinae), but a late survivor of an extinct lineage. This is not very likely given its age — it would be the last known survivor of the non-spheniscine penguins — but as some of these still lived a few million years ago, it cannot be ruled out.

References

{{Taxonbar, from=Q3798741 Extinct penguins Pliocene birds of Africa Monotypic prehistoric bird genera Fossil taxa described in 1975