Impala Limited
   HOME

TheInfoList



OR:

The impala or rooibok (''Aepyceros melampus'', lit. 'black-footed high-horn' in
Ancient Greek Ancient Greek (, ; ) includes the forms of the Greek language used in ancient Greece and the classical antiquity, ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Greek ...
) is a medium-sized
antelope The term antelope refers to numerous extant or recently extinct species of the ruminant artiodactyl family Bovidae that are indigenous to most of Africa, India, the Middle East, Central Asia, and a small area of Eastern Europe. Antelopes do ...
found in
eastern Eastern or Easterns may refer to: Transportation Airlines *China Eastern Airlines, a current Chinese airline based in Shanghai * Eastern Air, former name of Zambia Skyways *Eastern Air Lines, a defunct American airline that operated from 192 ...
and
southern Africa Southern Africa is the southernmost region of Africa. No definition is agreed upon, but some groupings include the United Nations geoscheme for Africa, United Nations geoscheme, the intergovernmental Southern African Development Community, and ...
. The only
extant Extant or Least-concern species, least concern is the opposite of the word extinct. It may refer to: * Extant hereditary titles * Extant literature, surviving literature, such as ''Beowulf'', the oldest extant manuscript written in English * Exta ...
member of the
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
'' Aepyceros'', and tribe Aepycerotini, it was first described to Europeans by German zoologist
Hinrich Lichtenstein Martin H nrich Carl Lichtenstein (10 January 1780 – 2 September 1857) was a German physician, List of explorers, explorer, botanist and zoologist. He explored parts of southern Africa and collected natural history specimens extensively and ...
in 1812. Two
subspecies In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...
are recognised—the grassland-dwelling common impala (sometimes referred to as the Kenyan impala), and the larger and darker black-faced impala, which lives in slightly more arid, scrubland environments. The impala reaches at the shoulder and weighs . It features a glossy, reddish brown
coat A coat is typically an outer garment for the upper body, worn by any gender for warmth or fashion. Coats typically have long sleeves and are open down the front, and closing by means of buttons, zippers, hook-and-loop fasteners (AKA velcro), ...
. The male's slender,
lyre The lyre () (from Greek λύρα and Latin ''lyra)'' is a string instrument, stringed musical instrument that is classified by Hornbostel–Sachs as a member of the History of lute-family instruments, lute family of instruments. In organology, a ...
-shaped
horns Horns or The Horns may refer to: * Plural of Horn (anatomy) * Plural of Horn (instrument), a group of musical instruments all with a horn-shaped bells * The Horns (Colorado), a summit on Cheyenne Mountain * Horns (novel), ''Horns'' (novel), a dar ...
are long. Active mainly during the day, the impala may be
gregarious Sociality is the degree to which individuals in an animal population tend to associate in social groups (gregariousness) and form cooperative societies. Sociality is a survival response to evolutionary pressures. For example, when a mother was ...
or
territorial A territory is an area of land, sea, or space, belonging or connected to a particular country, person, or animal. In international politics, a territory is usually a geographic area which has not been granted the powers of self-government, ...
depending upon the climate and geography. Three distinct social groups can be observed: the territorial males, bachelor herds and female herds. The impala is known for two characteristic leaps that constitute an anti-
predator Predation is a biological interaction in which one organism, the predator, kills and eats another organism, its prey. It is one of a family of common List of feeding behaviours, feeding behaviours that includes parasitism and micropredation ...
strategy.
Browsers Browse, browser, or browsing may refer to: Computing *Browser service, a feature of Microsoft Windows to browse shared network resources *Code browser, a program for navigating source code *File browser or file manager, a program used to manage f ...
as well as grazers, impala feed on
monocot Monocotyledons (), commonly referred to as monocots, (Lilianae ''sensu'' Chase & Reveal) are flowering plants whose seeds contain only one Embryo#Plant embryos, embryonic leaf, or cotyledon. A monocot taxon has been in use for several decades, but ...
s,
dicot The dicotyledons, also known as dicots (or, more rarely, dicotyls), are one of the two groups into which all the flowering plants (angiosperms) were formerly divided. The name refers to one of the typical characteristics of the group: namely, ...
s,
forb A forb or phorb is a herbaceous flowering plant that is not a graminoid (grass, sedge, or rush). The term is used in botany and in vegetation ecology especially in relation to grasslands and understory. Typically, these are eudicots without woo ...
s,
fruit In botany, a fruit is the seed-bearing structure in flowering plants (angiosperms) that is formed from the ovary after flowering. Fruits are the means by which angiosperms disseminate their seeds. Edible fruits in particular have long propaga ...
s and
acacia ''Acacia'', commonly known as wattles or acacias, is a genus of about of shrubs and trees in the subfamily Mimosoideae of the pea family Fabaceae. Initially, it comprised a group of plant species native to Africa, South America, and Austral ...
pods (whenever available). An annual, three-week-long rut takes place toward the end of the wet season, typically in May. Rutting males fight over dominance, and the victorious male courts females in
oestrus The estrous cycle (, originally ) is a set of recurring physiological changes induced by reproductive hormones in females of mammalian subclass Theria. Estrous cycles start after sexual maturity in females and are interrupted by anestrous phas ...
.
Gestation Gestation is the period of development during the carrying of an embryo, and later fetus, inside viviparous animals (the embryo develops within the parent). It is typical for mammals, but also occurs for some non-mammals. Mammals during pregn ...
lasts six to seven months, following which a single calf is born and immediately concealed in cover. Calves are suckled for four to six months; young males—forced out of the all-female groups—join bachelor herds, while females may stay back. The impala is found in woodlands and sometimes on the interface (
ecotone An ecotone is a transitional area between two plant communities, where these meet and integrate. Examples include areas between grassland and forest, estuaries and lagoon, freshwater and sea water etc. An ecotone may be narrow or wide, and it ma ...
) between woodlands and
savanna A savanna or savannah is a mixed woodland-grassland (i.e. grassy woodland) biome and ecosystem characterised by the trees being sufficiently widely spaced so that the canopy does not close. The open canopy allows sufficient light to reach th ...
hs; it inhabits places near water. While the black-faced impala is confined to southwestern
Angola Angola, officially the Republic of Angola, is a country on the west-Central Africa, central coast of Southern Africa. It is the second-largest Portuguese-speaking world, Portuguese-speaking (Lusophone) country in both total area and List of c ...
and
Kaokoland Kaokoland was an administrative unit and a Bantustan in northern South West Africa (now Namibia). Established in 1980 during the apartheid era, it was intended to be a self-governing homeland of the Ovahimba, but an actual government was nev ...
in northwestern
Namibia Namibia, officially the Republic of Namibia, is a country on the west coast of Southern Africa. Its borders include the Atlantic Ocean to the west, Angola and Zambia to the north, Botswana to the east and South Africa to the south; in the no ...
, the common impala is widespread across its range and has been reintroduced in
Gabon Gabon ( ; ), officially the Gabonese Republic (), is a country on the Atlantic coast of Central Africa, on the equator, bordered by Equatorial Guinea to the northwest, Cameroon to the north, the Republic of the Congo to the east and south, and ...
and southern Africa. The
International Union for Conservation of Nature The International Union for Conservation of Nature (IUCN) is an international organization working in the field of nature conservation and sustainable use of natural resources. Founded in 1948, IUCN has become the global authority on the stat ...
(IUCN) classifies the impala as a
species of least concern A least-concern species is a species that has been evaluated and categorized by the International Union for Conservation of Nature (IUCN) as not being a focus of wildlife conservation because the specific species is still plentiful in the wild ...
; the black-faced subspecies has been classified as a
vulnerable species A vulnerable species is a species which has been Conservation status, categorized by the International Union for Conservation of Nature as being threatened species, threatened with extinction unless the circumstances that are threatened species, ...
, with fewer than 1,000 individuals remaining in the wild as of 2008.


Etymology

The first attested English name, in 1802, was palla or pallah, from the
Tswana Tswana may refer to: * Tswana people, the Bantu languages, Bantu speaking people in Botswana, South Africa, Namibia, Zimbabwe, Zambia, and other Southern Africa regions * Tswana language, the language spoken by the (Ba)Tswana people * Tswanaland, ...
' 'red antelope'; the name ''impala'', also spelled ''impalla'' or ''mpala'', is first attested in 1875, and is directly from Zulu. Its
Afrikaans Afrikaans is a West Germanic languages, West Germanic language spoken in South Africa, Namibia and to a lesser extent Botswana, Zambia, Zimbabwe and also Argentina where there is a group in Sarmiento, Chubut, Sarmiento that speaks the Pat ...
name, ' 'red buck', is also sometimes used in English. The
scientific Science is a systematic discipline that builds and organises knowledge in the form of testable hypotheses and predictions about the universe. Modern science is typically divided into twoor threemajor branches: the natural sciences, which stu ...
generic name ''Aepyceros'' ( ‘high-horned’) comes from
Ancient Greek Ancient Greek (, ; ) includes the forms of the Greek language used in ancient Greece and the classical antiquity, ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Greek ...
(', 'high, steep') + (', 'horn'); the
specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...
''melampus'' ( ‘black-foot’) from (', 'black') + (', 'foot').


Taxonomy and evolution

The impala is the sole member of the genus ''Aepyceros'' and belongs to the
family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
Bovidae The Bovidae comprise the family (biology), biological family of cloven-hoofed, ruminant mammals that includes Bos, cattle, bison, Bubalina, buffalo, antelopes (including Caprinae, goat-antelopes), Ovis, sheep and Capra (genus), goats. A member o ...
. It was first described by German zoologist Martin Hinrich Carl Lichtenstein in 1812. In 1984, palaeontologist
Elisabeth Vrba Elisabeth S. Vrba (May 27, 1942 – February 5, 2025) was an American paleontologist at Yale University who developed the turnover-pulse hypothesis. Background Vrba was born in Hamburg, Germany. In 1944, following the death of her father, sh ...
opined that the impala is a
sister taxon In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an evolutionary tree. Definition The expression is most easily illustrated by a cladogram: Taxon A and ...
to the
alcelaphines The subfamily Alcelaphinae (or tribe Alcelaphini), of the family Bovidae, contains the wildebeest, tsessebe, topi, hartebeest, blesbok and bontebok, and several other related species. Depending on the classification, there are 6–10 species plac ...
, given its resemblance to the
hartebeest The hartebeest (; ''Alcelaphus buselaphus''), also known as kongoni or kaama, is an Fauna of Africa, African antelope. It is the Monotypic taxon, only member of the genus ''Alcelaphus''. Eight subspecies have been described, including two som ...
. A 1999
phylogenetic In biology, phylogenetics () is the study of the evolutionary history of life using observable characteristics of organisms (or genes), which is known as phylogenetic inference. It infers the relationship among organisms based on empirical dat ...
study by Alexandre Hassanin (of the National Centre for Scientific Research, Paris) and colleagues, based on
mitochondrial A mitochondrion () is an organelle found in the cells of most eukaryotes, such as animals, plants and fungi. Mitochondria have a double membrane structure and use aerobic respiration to generate adenosine triphosphate (ATP), which is used ...
and
nuclear Nuclear may refer to: Physics Relating to the nucleus of the atom: *Nuclear engineering *Nuclear physics *Nuclear power *Nuclear reactor *Nuclear weapon *Nuclear medicine *Radiation therapy *Nuclear warfare Mathematics * Nuclear space *Nuclear ...
analyses, showed that the impala forms a
clade In biology, a clade (), also known as a Monophyly, monophyletic group or natural group, is a group of organisms that is composed of a common ancestor and all of its descendants. Clades are the fundamental unit of cladistics, a modern approach t ...
with the
suni The suni (''Nesotragus moschatus'') is a small antelope of the family Bovidae, and one of the smallest ungulates on earth. It occurs in dense underbrush from central Kenya to KwaZulu-Natal in South Africa. It is also found on the island of Zanzi ...
(''Neotragus moschatus''). This clade is sister to another formed by the
bay duiker The bay duiker (''Cephalophus dorsalis''), also known as the black-striped duiker and the black-backed duiker, is a forest-dwelling duiker native to western and southern Africa. It was first described by British zoologist John Edward Gray in 18 ...
(''Cephalophus dorsalis'') and the
klipspringer The klipspringer (; ''Oreotragus oreotragus'') is a small antelope found in eastern and southern Africa. The sole member of its genus and subfamily/tribe, the klipspringer was first described by German zoologist Eberhard August Wilhelm von Zi ...
(''Oreotragus oreotragus''). An
rRNA Ribosomal ribonucleic acid (rRNA) is a type of non-coding RNA which is the primary component of ribosomes, essential to all cells. rRNA is a ribozyme which carries out protein synthesis in ribosomes. Ribosomal RNA is transcribed from ribosomal ...
and β-spectrin nuclear sequence analysis in 2003 also supported an association between ''Aepyceros'' and ''
Neotragus ''Neotragus'' is a genus of dwarf antelope, native to Africa. The genus includes only a single species without any dispute, namely '' Neotragus pygmaeus''. This species, the royal antelope, is the smallest antelope in the world, usually weighing ...
''. The following cladogram is based on the 1999 study: Up to six
subspecies In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...
have been described, although only two are generally recognised on the basis of mitochondrial data. Though morphologically similar, the subspecies show a significant
genetic distance Genetic distance is a measure of the genetics, genetic divergence between species or between population#Genetics, populations within a species, whether the distance measures time from common ancestor or degree of differentiation. Populations with ...
between them, and no hybrids between them have been reported. * ''A. m. melampus'' Lichtenstein, 1812: Known as the common impala, it occurs across eastern and southern Africa. The range extends from central
Kenya Kenya, officially the Republic of Kenya, is a country located in East Africa. With an estimated population of more than 52.4 million as of mid-2024, Kenya is the 27th-most-populous country in the world and the 7th most populous in Africa. ...
to
South Africa South Africa, officially the Republic of South Africa (RSA), is the Southern Africa, southernmost country in Africa. Its Provinces of South Africa, nine provinces are bounded to the south by of coastline that stretches along the Atlantic O ...
and westward into southeastern
Angola Angola, officially the Republic of Angola, is a country on the west-Central Africa, central coast of Southern Africa. It is the second-largest Portuguese-speaking world, Portuguese-speaking (Lusophone) country in both total area and List of c ...
. * ''A. m. petersi''
Bocage Bocage (, ) is a terrain of mixed woodland and pasture characteristic of parts of northern France, southern England, Ireland, the Netherlands, northern Spain and northern Germany, in regions where pastoral farming is the dominant land use. ' ...
, 1879
: Known as the black-faced impala, it is restricted to southwestern Africa, occurring in northwestern
Namibia Namibia, officially the Republic of Namibia, is a country on the west coast of Southern Africa. Its borders include the Atlantic Ocean to the west, Angola and Zambia to the north, Botswana to the east and South Africa to the south; in the no ...
and southwestern Angola. According to Vrba, the impala evolved from an alcelaphine ancestor. She noted that while this ancestor has diverged at least 18 times into various morphologically different forms, the impala has continued in its basic form for at least five million years. Several
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...
species have been discovered, including '' A. datoadeni'' from the
Pliocene The Pliocene ( ; also Pleiocene) is the epoch (geology), epoch in the geologic time scale that extends from 5.33 to 2.58Ethiopia Ethiopia, officially the Federal Democratic Republic of Ethiopia, is a landlocked country located in the Horn of Africa region of East Africa. It shares borders with Eritrea to the north, Djibouti to the northeast, Somalia to the east, Ken ...
. The oldest fossil discovered suggests its ancient ancestors were slightly smaller than the modern form, but otherwise very similar in all aspects to the latter. This implies that the impala has efficiently adapted to its environment since prehistoric times. Its gregarious nature, variety in diet, positive population trend, defence against
tick Ticks are parasitic arachnids of the order Ixodida. They are part of the mite superorder Parasitiformes. Adult ticks are approximately 3 to 5 mm in length depending on age, sex, and species, but can become larger when engorged. Ticks a ...
s and
symbiotic Symbiosis (Ancient Greek : living with, companionship < : together; and ''bíōsis'': living) is any type of a close and long-term biolo ...
relationship with the tick-feeding
oxpecker The oxpeckers are two species of bird which make up the genus ''Buphagus'', from Ancient Greek βοῦς (''boûs''), meaning "ox", and φάγος (''phágos''), meaning "eater", and family Buphagidae. The oxpeckers were formerly usually treat ...
s could have played a role in preventing major changes in morphology and behaviour.


Description

The impala is a medium-sized, slender-bodied
antelope The term antelope refers to numerous extant or recently extinct species of the ruminant artiodactyl family Bovidae that are indigenous to most of Africa, India, the Middle East, Central Asia, and a small area of Eastern Europe. Antelopes do ...
, comparable to the
kob The kob (''Kobus kob'') is an antelope found across Central Africa and parts of West Africa and East Africa. Together with the closely related reedbucks, waterbucks, lechwe, Nile lechwe, and puku, it forms the Reduncinae tribe. Found along ...
,
puku The puku (''Kobus vardonii'') is a medium-sized antelope found in wet grasslands in southern Democratic Republic of Congo, Namibia, Tanzania, Zambia and more concentrated in the Okavango Delta in Botswana. Nearly one-third of all puku are found ...
and
Grant's gazelle Grant's gazelle (''Nanger granti'') is a relatively large species of gazelle antelope, distributed from northern Tanzania to South Sudan and Ethiopia, and from the Kenyan coast to Lake Victoria. Its Swahili name is ''swala granti''. It was named ...
in size and build. The head-and-body length is around . Males reach approximately at the shoulder, while females are tall. Males typically weigh and females .
Sexually dimorphic Sexual dimorphism is the condition where sexes of the same species exhibit different Morphology (biology), morphological characteristics, including characteristics not directly involved in reproduction. The condition occurs in most dioecy, di ...
, females are hornless and smaller than males. Males grow slender, lyre-shaped horns long. The horns, strongly ridged and divergent, are circular in section and hollow at the base. Their arch-like structure allows interlocking of horns, which helps a male throw off his opponent during fights; horns also protect the skull from damage. The glossy coat of the impala shows two-tone colourationthe reddish brown back and the tan flanks; these are in sharp contrast to the white underbelly. Facial features include white rings around the eyes and a light chin and
snout A snout is the protruding portion of an animal's face, consisting of its nose, mouth, and jaw. In many animals, the structure is called a muzzle, Rostrum (anatomy), rostrum, beak or proboscis. The wet furless surface around the nostrils of the n ...
. The ears, long, are tipped with black. Black streaks run from the buttocks to the upper hindlegs. The bushy white tail, long, features a solid black stripe along the midline. The impala's colouration bears a strong resemblance to the
gerenuk The gerenuk (''Litocranius walleri''), also known as the giraffe gazelle, is a long-necked, medium-sized antelope found in parts of East Africa. The sole member of the genus ''Litocranius'', the gerenuk was first described by the naturalist Vi ...
, which has shorter horns and lacks the black thigh stripes of the impala. The impala has
scent gland Scent gland are exocrine glands found in most mammals. They produce semi-viscous secretions which contain pheromones and other semiochemical compounds. These odor-messengers indicate information such as status, territorial marking, mood, and se ...
s covered by a black tuft of hair on the hindlegs. 2-Methylbutanoic Acid and 2-Nonanone have been identified from this gland.
Sebaceous gland A sebaceous gland or oil gland is a microscopic exocrine gland in the skin that opens into a hair follicle to secrete an oily or waxy matter, called sebum, which lubricates the hair and skin of mammals. In humans, sebaceous glands occur in ...
s concentrated on the forehead and dispersed on the torso of dominant males are most active during the mating season, while those of females are only partially developed and do not undergo seasonal changes. There are four
nipples The nipple is a raised region of tissue on the surface of the breast from which, in lactating females, milk from the mammary gland leaves the body through the lactiferous ducts to nurse an infant. The milk can flow through the nipple passively, ...
. Of the subspecies, the black-faced impala is significantly larger and darker than the common impala;
melanism Melanism is the congenital excess of melanin in an organism resulting in dark pigment. Pseudomelanism, also called abundism, is another variant of pigmentation, identifiable by dark spots or enlarged stripes, which cover a large part of the bod ...
is responsible for the black colouration. Distinctive of the black-faced impala is a dark stripe, on either side of the nose, that runs upward to the eyes and thins as it reaches the forehead. Other differences include the larger black tip on the ear, and a bushier and nearly 30% longer tail in the black-faced impala. The impala has a special dental arrangement on the front lower jaw similar to the
toothcomb A toothcomb (also called a tooth comb or dental comb) is a Dentition, dental structure found in some mammals, comprising a group of front teeth arranged in a manner that facilitates grooming, similar to a hair comb. The toothcomb occurs in Lem ...
seen in strepsirrhine primates, which is used during
allogrooming Social grooming is a behavior in which social animals, including humans, clean or maintain one another's bodies or appearances. A related term, allogrooming, indicates social grooming between members of the same species. Grooming is a major s ...
to comb the fur on the head and the neck and remove
ectoparasite Parasitism is a Symbiosis, close relationship between species, where one organism, the parasite, lives (at least some of the time) on or inside another organism, the Host (biology), host, causing it some harm, and is Adaptation, adapted str ...
s.


Ecology and behaviour

The impala is diurnal (active mainly during the day), though activity tends to cease during the hot midday hours; they feed and rest at night. Three distinct social groups can be observedthe
territorial A territory is an area of land, sea, or space, belonging or connected to a particular country, person, or animal. In international politics, a territory is usually a geographic area which has not been granted the powers of self-government, ...
males, bachelor herds and female herds. The territorial males hold territories where they may form
harems A harem is a domestic space that is reserved for the women of the house in a Muslim family. A harem may house a man's wife or wives, their pre-pubescent male children, unmarried daughters, female domestic Domestic worker, servants, and other un ...
of females; territories are demarcated with urine and faeces and defended against juvenile or male intruders. Bachelor herds tend to be small, with less than 30 members. Individuals maintain distances of from one another; while young and old males may interact, middle-aged males generally avoid one another except to spar. Female herds vary in size from 6 to 100; herds occupy
home range A home range is the area in which an animal lives and moves on a periodic basis. It is related to the concept of an animal's territory which is the area that is actively defended. The concept of a home range was introduced by W. H. Burt in 1943. ...
s of . The mother–calf bond is weak, and breaks soon after weaning; juveniles leave the herds of their mothers to join other herds. Female herds tend to be loose and have no obvious leadership. Allogrooming is an important means of social interaction in bachelor and female herds; in fact, the impala appears to be the only
ungulate Ungulates ( ) are members of the diverse clade Euungulata ("true ungulates"), which primarily consists of large mammals with Hoof, hooves. Once part of the clade "Ungulata" along with the clade Paenungulata, "Ungulata" has since been determined ...
to display self-grooming as well as allogrooming. In allogrooming, females typically groom related impalas, while males associate with unrelated ones. Each partner grooms the other six to twelve times. Social behaviour is influenced by the climate and geography; as such, the impala are territorial at certain times of the year and gregarious at other times, and the length of these periods can vary broadly among populations. For instance, populations in southern Africa display territorial behaviour only during the few months of the rut, whereas in eastern African populations, territoriality is relatively minimal despite a protracted mating season. Moreover, territorial males often tolerate bachelors, and may even alternate between bachelorhood and territoriality at different times of the year. A study of impala in the
Serengeti National Park The Serengeti National Park is a large national park in northern Tanzania that stretches over . It is located in eastern Mara Region and northeastern Simiyu Region and contains over of virgin savanna. The park was established in 1940. The Se ...
showed that in 94% of the males, territoriality was observed for less than four months. The impala is an important prey species for Africa's large
carnivore A carnivore , or meat-eater (Latin, ''caro'', genitive ''carnis'', meaning meat or "flesh" and ''vorare'' meaning "to devour"), is an animal or plant Plants are the eukaryotes that form the Kingdom (biology), kingdom Plantae; they ar ...
s, such as
cheetah The cheetah (''Acinonyx jubatus'') is a large Felidae, cat and the Fastest animals, fastest land animal. It has a tawny to creamy white or pale buff fur that is marked with evenly spaced, solid black spots. The head is small and rounded, wit ...
s,
leopard The leopard (''Panthera pardus'') is one of the five extant cat species in the genus ''Panthera''. It has a pale yellowish to dark golden fur with dark spots grouped in rosettes. Its body is slender and muscular reaching a length of with a ...
s,
wild dogs Wild dogs may refer to: *African wild dog also called painted dogs * Dingo–dog hybrids in Australia * Free-ranging dog#Free-ranging unowned dogs, including feral dogs and "wild" dogs * Wild Dogs, an American heavy metal band from Portland, Oregon ...
,
lion The lion (''Panthera leo'') is a large Felidae, cat of the genus ''Panthera'', native to Sub-Saharan Africa and India. It has a muscular, broad-chested body (biology), body; a short, rounded head; round ears; and a dark, hairy tuft at the ...
s,
hyenas Hyenas or hyaenas ( ; from Ancient Greek , ) are feliformia, feliform carnivoran mammals belonging to the Family (biology), family Hyaenidae (). With just four extant species (each in its own genus), it is the fifth-smallest family in the orde ...
,
crocodiles Crocodiles (family Crocodylidae) or true crocodiles are large, semiaquatic reptiles that live throughout the tropics in Africa, Asia, the Americas and Australia. The term "crocodile" is sometimes used more loosely to include all extant member ...
and
pythons The Pythonidae, commonly known as pythons, are a family of nonvenomous snakes found in Africa, Asia, and Australia. Among its members are some of the largest snakes in the world. Ten genera and 39 species are currently recognized. Being natura ...
. The antelope displays two characteristic leapsit can jump up to , over vegetation and even other impala, covering distances of up to ; the other type of leap involves a series of jumps in which the animal lands on its forelegs, moves its hindlegs mid-air in a kicking fashion, lands on all fours (
stotting Stotting (also called pronking or pronging) is a behavior of quadrupeds, particularly gazelles, in which they spring into the air, lifting all four feet off the ground simultaneously. Usually, the legs are held in a relatively stiff position. Ma ...
) and then rebounds. It leaps in either manner in different directions, probably to confuse
predator Predation is a biological interaction in which one organism, the predator, kills and eats another organism, its prey. It is one of a family of common List of feeding behaviours, feeding behaviours that includes parasitism and micropredation ...
s. At times, the impala may also conceal itself in vegetation to escape the eye of the predator. The most prominent vocalisation is the loud
roar A roar is a type of animal vocalization that is loud, deep and resonating. Many mammals have evolved to produce roars and other roar-like vocals for purposes such as long-distance communication and intimidation. These include various species of ...
, delivered through one to three loud snorts with the mouth closed, followed by two to ten deep grunts with the mouth open and the chin and tail raised; a typical roar can be heard up to away. Scent gland secretions identify a territorial male. Impalas are sedentary; adult and middle-aged males, in particular, can hold their territories for years.


Parasites

Common
ixodid The Ixodidae are the family of hard ticks or scale ticks, one of the three families of ticks, consisting of 771 species, . They are known as 'hard ticks' because they have a scutum or hard shield, which the other major family of ticks, the 'sof ...
ticks collected from impala include '' Amblyomma hebraeum'', '' Boophilus decoloratus'', ''
Hyalomma marginatum ''Hyalomma marginatum'' is a hard-bodied tick found on birds including the pale crag martin. This tick has been implicated in the transmission of Tete virus, Bahig virus, a pathogenic arbovirus previously thought to be transmitted only by mosquit ...
'', ''
Ixodes cavipalpus ''Ixodes'' is a genus of hard-bodied ticks (family Ixodidae). It includes important disease vectors of animals and humans (tick-borne disease), and some species (notably ''Ixodes holocyclus'') inject toxins that can cause paralysis. Some ti ...
'', ''
Rhipicephalus appendiculatus ''Rhipicephalus appendiculatus'', the brown ear tick, is a hard tick found in Africa where it spreads the parasite '' Theileria parva'', the cause of East Coast fever in cattle. The tick has a three-host life-cycle, spending around 10% of its li ...
'' and '' R. evertsi''. In
Zimbabwe file:Zimbabwe, relief map.jpg, upright=1.22, Zimbabwe, relief map Zimbabwe, officially the Republic of Zimbabwe, is a landlocked country in Southeast Africa, between the Zambezi and Limpopo Rivers, bordered by South Africa to the south, Bots ...
, heavy infestation by ticks such as ''R. appendiculatus'' has proved to be a major cause behind the high mortality of ungulates, as they can lead to
tick paralysis Tick paralysis is a type of paralysis caused by specific types of ticks. Unlike tick-borne diseases caused by infectious organisms, the illness is caused by a neurotoxin produced in the tick's salivary gland. After prolonged attachment, the en ...
. Impala have special adaptations for grooming, such as their characteristic dental arrangement, to manage ticks before they engorge; however, the extensive grooming needed to keep the tick load under control involves the risk of dehydration during summer, lower vigilance against predators and gradual wearing out of the teeth. A study showed that impala adjust the time devoted to grooming and the number of grooming bouts according to the seasonal prevalence of ticks. Impala are symbiotically related to
oxpecker The oxpeckers are two species of bird which make up the genus ''Buphagus'', from Ancient Greek βοῦς (''boûs''), meaning "ox", and φάγος (''phágos''), meaning "eater", and family Buphagidae. The oxpeckers were formerly usually treat ...
s, which feed on ticks from those parts of the antelope's body which the animal cannot access by itself (such as the ears, neck, eyelids, forehead and underbelly). The impala is the smallest ungulate with which oxpeckers are associated. In a study it was observed that oxpeckers selectively attended to impala despite the presence of other animals such as
Coke's hartebeest Coke's hartebeest (''Alcelaphus buselaphus cokii'') or ''Kongoni'' is a large migratory antelope that is native to Kenya and Tanzania. It can breed with Lelwel hartebeest to produce a hybrid known as the Kenya Highland hartebeest (''Alcelaphus ...
, Grant's gazelle,
Thomson's gazelle Thomson's gazelle (''Eudorcas thomsonii'') is one of the best known species of gazelles. It is named after explorer Joseph Thomson (explorer), Joseph Thomson and is sometimes referred to as a "tommie". It is considered by some to be a subspecies o ...
and
topi ''Damaliscus lunatus jimela'' is a subspecies of Damaliscus lunatus, topi, and is usually just called a topi. It is a highly social and fast type of antelope found in the savannas, semi-deserts, and floodplains of sub-Saharan Africa. Names The ...
. A possible explanation for this could be that because the impala inhabits woodlands (which can have a high density of ticks), the impala could have greater mass of ticks per unit area of the body surface. Another study showed that the oxpeckers prefer the ears over other parts of the body, probably because these parts show maximum tick infestation. The bird has also been observed to perch on the
udders An udder is an organ formed of two or four mammary glands on the females of dairy animals and ruminants such as cattle, goats, and sheep. An udder is equivalent to the breast in primates, elephantine pachyderms and other mammals. The udder i ...
of a female and pilfer its
milk Milk is a white liquid food produced by the mammary glands of lactating mammals. It is the primary source of nutrition for young mammals (including breastfeeding, breastfed human infants) before they are able to digestion, digest solid food. ...
.
Lice Louse (: lice) is the common name for any member of the infraorder Phthiraptera, which contains nearly 5,000 species of wingless parasitic insects. Phthiraptera was previously recognized as an order, until a 2021 genetic study determined th ...
recorded from impala include '' Damalinia aepycerus'', '' D. elongata'', ''
Linognathus aepycerus ''Linognathus'' is a genus of lice belonging to the family Linognathidae Linognathidae is a family of lice in the order Psocodea. There are at least 3 genera and 70 described species in Linognathidae. Genera These three genera Genus (; : ...
'' and '' L. nevilli''; in a study,
ivermectin Ivermectin is an antiparasitic drug. After its discovery in 1975, its first uses were in veterinary medicine to prevent and treat heartworm and acariasis. Approved for human use in 1987, it is used to treat infestations including head lice ...
(a medication against parasites) was found to have an effect on ''Boophilus decoloratus'' and ''Linognathus'' species, though not on ''Damalinia'' species. In a study of impala in South Africa, the number of
worm Worms are many different distantly related bilateria, bilateral animals that typically have a long cylindrical tube-like body, no limb (anatomy), limbs, and usually no eyes. Worms vary in size from microscopic to over in length for marine ...
s in juveniles showed an increase with age, reaching a peak when impala turned a year old. This study recorded worms of genera such as '' Cooperia'', '' Cooperoides'', ''
Fasciola ''Fasciola'', commonly known as the liver fluke, is a genus of parasitic trematodes. There are three species within the genus ''Fasciola'': ''Fasciola nyanzae,'' ''Fasciola'' ''hepatica'' and ''Fasciola'' ''gigantica''. ''Fasciola hepatica'' an ...
'', ''
Gongylonema ''Gongylonema'' is a genus of thread-like nematode that was described by Molin in 1857. It is the only currently valid genus in the family Gongylonematidae, though the mysterious '' Spiruroides'' – usually placed in the Subuluridae, which are ...
''. ''
Haemonchus ''Haemonchus'' is a genus of nematodes belonging to the family Trichostrongylidae. They are abomasal parasites that cause anemia, followed by white eye disease. The genus has cosmopolitan distribution. Species: *''Haemonchus contortus '' ...
'', '' Impalaia'', ''
Longistrongylus ''Longistrongylus'' is a genus of nematodes belonging to the family Trichostrongylidae. Species: * ''Longistrongylus muraschkinzevi'' (Shulz & Kadenatsii, 1950) References

Trichostrongylidae Rhabditida genera {{rhabditida-stub ...
'' and ''
Trichostrongylus ''Trichostrongylus'' species are nematodes (round worms), which are ubiquitous among herbivores worldwide, including cattle, sheep, donkeys, goats, deer, and rabbits.Garcia LS, editor. Diagnostic Medical Parasitology. 5 ed. Washington, DC: ASM Pr ...
''; some of these showed seasonal variations in density. Impala show high frequency of
defensive behaviour Deimatic behaviour or startle display means any pattern of bluffing behaviour in an animal that lacks strong defences, such as suddenly displaying conspicuous eyespots, to scare off or momentarily distract a predator, thus giving the prey anima ...
s towards flying insects. This is probably the reason for Vale 1977 and Clausen et al 1998 only finding trace levels of feeding by ''Glossina'' (
tsetse fly Tsetse ( , or ) (sometimes spelled tzetze; also known as tik-tik flies) are large, biting flies that inhabit much of tropical Africa. Tsetse flies include all the species in the genus ''Glossina'', which are placed in their own family, Gloss ...
) upon impala. ''
Theileria ''Theileria'' is a genus of parasites that belongs to the phylum Apicomplexa, and is closely related to ''Plasmodium''. Two ''Theileria'' species, ''T. annulata'' and ''T. parva'', are important cattle parasites. ''T. annulata'' causes tropical ...
'' of impala in
Kenya Kenya, officially the Republic of Kenya, is a country located in East Africa. With an estimated population of more than 52.4 million as of mid-2024, Kenya is the 27th-most-populous country in the world and the 7th most populous in Africa. ...
are not cross infectious to
cattle Cattle (''Bos taurus'') are large, domesticated, bovid ungulates widely kept as livestock. They are prominent modern members of the subfamily Bovinae and the most widespread species of the genus '' Bos''. Mature female cattle are calle ...
: Grootenhuis et al 1975 were not able to induce cattle infection and Fawcett et al 1987 did not find it naturally occurring.


Diet

Impala
browse Browsing is a kind of orienting strategy. It is supposed to identify something of relevance for the browsing organism. In context of humans, it is a metaphor taken from the animal kingdom. It is used, for example, about people browsing open sh ...
as well as graze; either may predominate, depending upon the availability of resources. The diet comprises
monocot Monocotyledons (), commonly referred to as monocots, (Lilianae ''sensu'' Chase & Reveal) are flowering plants whose seeds contain only one Embryo#Plant embryos, embryonic leaf, or cotyledon. A monocot taxon has been in use for several decades, but ...
s,
dicot The dicotyledons, also known as dicots (or, more rarely, dicotyls), are one of the two groups into which all the flowering plants (angiosperms) were formerly divided. The name refers to one of the typical characteristics of the group: namely, ...
s,
forb A forb or phorb is a herbaceous flowering plant that is not a graminoid (grass, sedge, or rush). The term is used in botany and in vegetation ecology especially in relation to grasslands and understory. Typically, these are eudicots without woo ...
s, fruits and ''
acacia ''Acacia'', commonly known as wattles or acacias, is a genus of about of shrubs and trees in the subfamily Mimosoideae of the pea family Fabaceae. Initially, it comprised a group of plant species native to Africa, South America, and Austral ...
'' pods (whenever available). Impala prefer places close to water sources, and resort to succulent vegetation if water is scarce. An analysis showed that the diet of impala is composed of 45% monocots, 45% dicots and 10% fruits; the proportion of grasses in the diet increases significantly (to as high as 90%) after the first rains, but declines in the
dry season The dry season is a yearly period of low rainfall, especially in the tropics. The weather in the tropics is dominated by the tropical rain belt, which moves from the northern to the southern tropics and back over the course of the year. The t ...
. Browsing predominates in the late wet and dry season, and diets are nutritionally poor in the mid-dry season, when impala feed mostly on woody dicots. Another study showed that the dicot proportion in the diet is much higher in bachelors and females than in territorial males. Impala feed on soft and nutritious grasses such as ''
Digitaria macroblephara ''Digitaria'' is a genus of plants in the grass family native to tropical and warm temperate regions but can occur in tropical, subtropical, and cooler temperate regions as well. Common names include crabgrass, finger-grass, and fonio. They are ...
''; tough, tall grasses, such as ''
Heteropogon contortus ''Heteropogon contortus'' is a tropical, perennial tussock grass with a native distribution encompassing Southern Africa, southern Asia, Northern Australia, Oceania, and southwestern North America. The species has also become a naturalised weed ...
'' and ''
Themeda triandra } ''Themeda triandra'' is a species of perennial tussock-forming grass widespread in Africa, Australia, Asia and the Pacific. In Australia it is commonly known as kangaroo grass and in East Africa and South Africa it is known as red grass and ...
'', are typically avoided. Impala on the periphery of the herds are generally more vigilant against predators than those feeding in the centre; a foraging individual will try to defend the patch it is feeding on by lowering its head. A study revealed that time spent in foraging reaches a maximum of 75.5% of the day in the late dry season, decreases through the rainy season, and is minimal in the early dry season (57.8%).


Reproduction

Males are
sexually mature Sexual maturity is the capability of an organism to reproduce. In humans, it is related to both puberty and adulthood. ''Puberty'' is the biological process of sexual maturation, while ''adulthood'', the condition of being socially recognized as ...
by the time they are a year old, though successful mating generally occurs only after four years. Mature males start establishing territories and try to gain access to females. Females can conceive after they are a year and a half old;
oestrus The estrous cycle (, originally ) is a set of recurring physiological changes induced by reproductive hormones in females of mammalian subclass Theria. Estrous cycles start after sexual maturity in females and are interrupted by anestrous phas ...
lasts for 24 to 48 hours, and occurs every 12–29 days in non-pregnant females. The annual three-week-long rut (breeding season) begins toward the end of the
wet season The wet season (sometimes called the rainy season or monsoon season) is the time of year when most of a region's average annual rainfall occurs. Generally, the season lasts at least one month. The term ''green season'' is also sometimes used a ...
, typically in May.
Gonad A gonad, sex gland, or reproductive gland is a Heterocrine gland, mixed gland and sex organ that produces the gametes and sex hormones of an organism. Female reproductive cells are egg cells, and male reproductive cells are sperm. The male gon ...
al growth and
hormone A hormone (from the Ancient Greek, Greek participle , "setting in motion") is a class of cell signaling, signaling molecules in multicellular organisms that are sent to distant organs or tissues by complex biological processes to regulate physio ...
production in males begin a few months before the breeding season, resulting in greater aggressiveness and territoriality. The
bulbourethral gland The bulbourethral glands or Cowper's glands (named for English anatomist William Cowper) are two small exocrine and accessory glands in the reproductive system of many male mammals. They are homologous to Bartholin's glands in females. The bul ...
s are heavier,
testosterone Testosterone is the primary male sex hormone and androgen in Male, males. In humans, testosterone plays a key role in the development of Male reproductive system, male reproductive tissues such as testicles and prostate, as well as promoting se ...
levels are nearly twice as high in territorial males as in bachelors, and the neck of a territorial male tends to be thicker than that of a bachelor during the rut. Mating tends to take place between full moons. Rutting males fight over dominance, often giving out noisy roars and chasing one another; they walk stiffly and
display Display may refer to: Technology * Display device, output device for presenting information, including: ** Electronic visual display, output device to present information for visual or tactile reception *** Cathode-ray tube (CRT), that uses an el ...
their neck and horns. Males desist from feeding and allogrooming during the rut, probably to devote more time to garnering females in oestrus; the male checks the female's urine to ensure that she is in oestrus. On coming across such a female, the excited male begins the
courtship Courtship is the period wherein some couples get to know each other prior to a possible marriage or committed romantic, ''de facto'' relationship. Courtship traditionally may begin after a betrothal and may conclude with the celebration of marri ...
by pursuing her, keeping a distance of from her. The male flicks his tongue and may nod vigorously; the female allows him to lick her vulva, and holds her tail to one side. The male tries mounting the female, holding his head high and clasping her sides with his forelegs. Mounting attempts may be repeated every few seconds to every minute or two. The male loses interest in the female after the first copulation, though she is still active and can mate with other males. Gestation lasts six to seven months. Births generally occur in the midday; the female will isolate herself from the herd when labour pain begins. The perception that females can delay giving birth for an additional month if conditions are harsh may however not be realistic. A single calf is born, and is immediately concealed in cover for the first few weeks of its birth. The fawn then joins a nursery group within its mother's herd. Calves are suckled for four to six months; young males, forced out of the group, join bachelor herds, while females may stay back.


Distribution and habitat

The impala inhabits woodlands due to its preference for shade; it can also be found on the interface (ecotone) between woodlands and savannahs. Places near water sources are preferred. In southern Africa, populations tend to be associated with ''
Colophospermum mopane ''Colophospermum mopane'', commonly called mopane, mopani, butterfly tree, turpentine tree, or balsam tree, is a tree in the legume family (Fabaceae), that grows in hot, dry, low-lying areas, in elevation, in parts of Southern Africa. The tree ...
'' and ''Acacia'' woodlands. Habitat choices differ seasonally''
Acacia senegal ''Senegalia senegal'' (also known as ''Acacia senegal'') is a small thorny deciduous tree from the genus ''Senegalia'', which is known by several common names, including gum acacia, gum arabic tree, Sudan gum and Sudan gum arabic. In parts of In ...
'' woodlands are preferred in the wet season, and '' A. drepanolobium'' savannahs in the dry season. Another factor that could influence habitat choice is vulnerability to predators; impala tend to keep away from areas with tall grasses as predators could be concealed there. A study found that the reduction of woodland cover and creation of shrublands by the
African bush elephant The African bush elephant (''Loxodonta africana''), also known as the African savanna elephant, is a species of elephant native to sub-Saharan Africa. It is one of three extant elephant species and, along with the African forest elephant, one ...
s has favoured impala population by increasing the availability of more dry season browse. Earlier, the ''
Baikiaea ''Baikiaea'' is a genus of flowering plants in the legume family, Fabaceae. It includes six species of trees native to sub-Saharan Africa. Species range from Nigeria eastwards to Uganda and Tanzania and south to Zimbabwe, Botswana, and Namibia. H ...
'' woodland, which has now declined due to elephants, provided minimum browsing for impala. The newly formed ''
Capparis ''Capparis'' is a genus of flowering plants in the family Capparaceae. It includes 142 species of shrubs or lianas which are collectively known as caper shrubs or caperbushes. ''Capparis'' species occur over a wide range of habitat in the subtr ...
'' shrubland, on the other hand, could be a key browsing habitat. Impala are generally not associated with
montane Montane ecosystems are found on the slopes of mountains. The alpine climate in these regions strongly affects the ecosystem because temperatures lapse rate, fall as elevation increases, causing the ecosystem to stratify. This stratification is ...
habitats; however, in
KwaZulu-Natal KwaZulu-Natal (, also referred to as KZN) is a Provinces of South Africa, province of South Africa that was created in 1994 when the government merged the Zulu people, Zulu bantustan of KwaZulu ("Place of the Zulu" in Zulu language, Zulu) and ...
, impala have been recorded at altitudes of up to
above sea level Height above mean sea level is a measure of a location's vertical distance (height, elevation or altitude) in reference to a vertical datum based on a historic mean sea level. In geodesy, it is formalized as orthometric height. The zero level ...
. The historical range of the impala – spanning across southern and eastern Africa – has remained intact to a great extent, although it has disappeared from a few places, such as
Burundi Burundi, officially the Republic of Burundi, is a landlocked country in East Africa. It is located in the Great Rift Valley at the junction between the African Great Lakes region and Southeast Africa, with a population of over 14 million peop ...
. The range extends from central and southern Kenya and northeastern
Uganda Uganda, officially the Republic of Uganda, is a landlocked country in East Africa. It is bordered to the east by Kenya, to the north by South Sudan, to the west by the Democratic Republic of the Congo, to the south-west by Rwanda, and to the ...
in the east to northern KwaZulu-Natal in the south, and westward up to Namibia and southern Angola. The black-faced impala is confined to southwestern Angola and
Kaokoland Kaokoland was an administrative unit and a Bantustan in northern South West Africa (now Namibia). Established in 1980 during the apartheid era, it was intended to be a self-governing homeland of the Ovahimba, but an actual government was nev ...
in northwestern Namibia; the status of this subspecies has not been monitored since the 2000s. The common impala has a wider distribution, and has been introduced in protected areas in
Gabon Gabon ( ; ), officially the Gabonese Republic (), is a country on the Atlantic coast of Central Africa, on the equator, bordered by Equatorial Guinea to the northwest, Cameroon to the north, the Republic of the Congo to the east and south, and ...
and across southern Africa.


Threats and conservation

Impala tracks The
International Union for Conservation of Nature and Natural Resources The International Union for Conservation of Nature (IUCN) is an international organization working in the field of nature conservation and sustainable use of natural resources. Founded in 1948, IUCN has become the global authority on the status ...
(IUCN) classifies the impala as a
species of least concern A least-concern species is a species that has been evaluated and categorized by the International Union for Conservation of Nature (IUCN) as not being a focus of wildlife conservation because the specific species is still plentiful in the wild ...
overall. The black-faced impala, however, is classified as a
vulnerable species A vulnerable species is a species which has been Conservation status, categorized by the International Union for Conservation of Nature as being threatened species, threatened with extinction unless the circumstances that are threatened species, ...
; as of 2008, fewer than 1,000 were estimated in the wild. Though there are no major threats to the survival of the common impala, poaching and natural calamities have significantly contributed to the decline of the black-faced impala. As of 2008, the population of the common impala has been estimated at around two million. According to some studies, translocation of the black-faced impala can be highly beneficial in its conservation. Around a quarter of the common impala populations occur in protected areas, such as the
Okavango Delta The Okavango Delta or Okavango Grassland is a vast inland delta in Botswana formed where the Okavango River reaches a tectonic trough at an elevation of in the central part of the endorheic basin of the Kalahari Desert. It is a UNESCO Wor ...
(
Botswana Botswana, officially the Republic of Botswana, is a landlocked country in Southern Africa. Botswana is topographically flat, with approximately 70 percent of its territory part of the Kalahari Desert. It is bordered by South Africa to the sou ...
);
Masai Mara Maasai Mara, sometimes also spelt Masai Mara and locally known simply as The Mara, is a large national game reserve in Narok County, Kenya, contiguous with the Serengeti National Park in Tanzania. It is named in honour of the Maasai people, th ...
and
Kajiado Kajiado is a town in Kajiado County, Kenya. The town is located south of Nairobi, along the Nairobi – Arusha highway ( A104 road). Kajiado has an urban population of 24,678 (2019 census). Local people are predominantly of the Maasai tribe. ...
(Kenya);
Kruger National Park Kruger National Park () is a national park in South Africa covering an area of in the provinces of Limpopo and Mpumalanga in the country's northeast. It extends from north to south and from east to west. The administrative headquarters are i ...
(South Africa); the Ruaha and
Serengeti National Park The Serengeti National Park is a large national park in northern Tanzania that stretches over . It is located in eastern Mara Region and northeastern Simiyu Region and contains over of virgin savanna. The park was established in 1940. The Se ...
s and
Selous Game Reserve The Selous Game Reserve, now renamed as Nyerere National Park (in-part), is a protected nature reserve and wilderness area in southern Tanzania, East Africa. It covers a total area of , with additional buffer zones, as well. It was designated a ...
(
Tanzania Tanzania, officially the United Republic of Tanzania, is a country in East Africa within the African Great Lakes region. It is bordered by Uganda to the northwest; Kenya to the northeast; the Indian Ocean to the east; Mozambique and Malawi to t ...
);
Luangwa Valley The Luangwa River is one of the major tributaries of the Zambezi River, and one of the four biggest rivers of Zambia. The river generally floods in the rainy season (December to March) and then falls considerably in the dry season. It is one of ...
(
Zambia Zambia, officially the Republic of Zambia, is a landlocked country at the crossroads of Central Africa, Central, Southern Africa, Southern and East Africa. It is typically referred to being in South-Central Africa or Southern Africa. It is bor ...
);
Hwange Hwange, originally known as Wankie, is a town in Zimbabwe, located in Hwange District, in Matabeleland North Province, in northwestern Zimbabwe, close to the international borders with Botswana and Zambia. It lies approximately , by road, sou ...
, Sebungwe and
Zambezi Valley The Zambezi (also spelled Zambeze and Zambesi) is the fourth-longest river in Africa, the longest east-flowing river in Africa and the largest flowing into the Indian Ocean from Africa. Its drainage basin covers , slightly less than half of t ...
(
Zimbabwe file:Zimbabwe, relief map.jpg, upright=1.22, Zimbabwe, relief map Zimbabwe, officially the Republic of Zimbabwe, is a landlocked country in Southeast Africa, between the Zambezi and Limpopo Rivers, bordered by South Africa to the south, Bots ...
). The rare black-faced impala has been introduced into private farms in Namibia and the
Etosha National Park Etosha National Park is a national park in northwestern Namibia and one of the largest national parks in Africa. It was proclaimed a game reserve in March 1907 in Ordinance 88 by the Governor of German South West Africa, Friedrich von Lindequist. ...
. Population densities vary largely from place to place; from less than one impala per square kilometre in
Mkomazi National Park Mkomazi National Park is a national park in Tanzania, located in Same District of Kilimanjaro Region and Lushoto District, with a slither of the park in Mkinga District both of Tanga Region. It was established as a game reserve in 1951 and upg ...
(Tanzania) to as high as 135 per square kilometre near
Lake Kariba Lake Kariba is the world's largest artificial lake and reservoir by volume. It lies upstream from the mouth of the Zambezi river on the Indian Ocean, along the border between Zambia and Zimbabwe. Lake Kariba was filled between 1958 and 1963 fo ...
(Zimbabwe).


References


External links


Impala: Summary from the African Wildlife Foundation

ITIS Standard Report Page
{{Authority control Antelopes of Sub-Saharan Africa Fauna of East Africa Mammals of Southern Africa Mammals described in 1812