''Imagotaria'' is an

extinct Extinction is the termination of an organism by the death of its Endling, last member. A taxon may become Functional extinction, functionally extinct before the death of its last member if it loses the capacity to Reproduction, reproduce and ...

monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unisp ...

genus of

walrus The walrus (''Odobenus rosmarus'') is a large pinniped marine mammal with discontinuous distribution about the North Pole in the Arctic Ocean and subarctic seas of the Northern Hemisphere. It is the only extant species in the family Odobeni ...

with the sole species ''Imagotaria downsi''.

Fossils A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...

of ''Imagotaria'' are known from the early late

Miocene The Miocene ( ) is the first epoch (geology), geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and mea ...

California California () is a U.S. state, state in the Western United States that lies on the West Coast of the United States, Pacific Coast. It borders Oregon to the north, Nevada and Arizona to the east, and shares Mexico–United States border, an ...

(c. 10-12 million years ago).

Description

The long

pinniped Pinnipeds (pronounced ), commonly known as seals, are a widely range (biology), distributed and diverse clade of carnivorous, fin-footed, semiaquatic, mostly marine mammals. They comprise the extant taxon, extant families Odobenidae (whose onl ...

more closely resembled in its overall shape a

sea lion Sea lions are pinnipeds characterized by external ear flaps, long foreflippers, the ability to walk on all fours, short and thick hair, and a big chest and belly. Together with the fur seals, they make up the family Otariidae, eared seals. ...

rather than a walrus. Unlike the extant walrus (''Odobenus rosmarus''), ''Imagotaria'' did not possess elongate, ever-growing tusks, but instead bore enlarged canines (with respect to other pinnipeds). ''Imagotaria'' is an example of a primitive walrus that does not grossly appear similar to a modern walrus. However, the walrus family (the

Odobenidae Odobenidae is a family of pinnipeds, of which the only extant species is the walrus (''Odobenus rosmarus''). In the past, however, the group was much more diverse, and includes more than a dozen fossil genera. Taxonomy All genera, except '' Od ...

) is a more inclusive group, that includes walruses without tusks (e.g. ''Imagotaria''), walruses with upper and lower tusks (the subfamily Dusignathinae), and walruses with upper tusks like the extant walrus (subfamily Odobeninae, tribe Odobenini). It is possible to classify these pinnipeds as walruses because they share many other

skull The skull, or cranium, is typically a bony enclosure around the brain of a vertebrate. In some fish, and amphibians, the skull is of cartilage. The skull is at the head end of the vertebrate. In the human, the skull comprises two prominent ...

features (besides tusks) as well as many skeletal features, all of which indicate common ancestry.

Palaeobiology

The

teeth A tooth (: teeth) is a hard, calcified structure found in the jaws (or mouths) of many vertebrates and used to break down food. Some animals, particularly carnivores and omnivores, also use teeth to help with capturing or wounding prey, tear ...

of ''Imagotaria'' indicate that its feeding ecology was markedly different from that of modern walrus, and more similar to that of less specialized pinnipeds like seals, fur seals, and sea lions. Conical, unworn teeth and the lack of a vaulted palate indicate that ''Imagotaria'' did not feed on molluscs like modern walrus. Modern walruses do not use their teeth to chew

molluscs Mollusca is a phylum of protostome, protostomic invertebrate animals, whose members are known as molluscs or mollusks (). Around 76,000 extant taxon, extant species of molluscs are recognized, making it the second-largest animal phylum ...

sea otters The sea otter (''Enhydra lutris'') is a marine mammal native to the coasts of the northern and eastern North Pacific Ocean. Adult sea otters typically weigh between , making them the heaviest members of the weasel fa ...

do. Instead, they hold a clam in their lips, and the vaulted palate allows them to use their tongue as a powerful piston to suck the soft parts right out of the clam shell. The shell is then dropped to the seafloor, never entering the oral cavity. Additionally, fossils of ''Imagotaria'' (and the earlier '' Neotherium'', c. 15 million years ago) demonstrate that early walruses had, by the middle and late Miocene, already developed extreme

sexual dimorphism Sexual dimorphism is the condition where sexes of the same species exhibit different Morphology (biology), morphological characteristics, including characteristics not directly involved in reproduction. The condition occurs in most dioecy, di ...

(males and females having different body sizes). It is unclear whether extreme sexual dimorphism is ancestral to all pinnipeds, or if it has been independently acquired in multiple pinniped lineages.

References

{{Taxonbar, from=Q29381283 Miocene pinnipeds Prehistoric pinnipeds of North America Monotypic prehistoric carnivoran genera Odobenids Fossil taxa described in 1968