Iho Eleru
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Iho Eleru, formerly known as Iwo Eleeru, is an
archaeological site An archaeological site is a place (or group of physical sites) in which evidence of past activity is preserved (either prehistoric or recorded history, historic or contemporary), and which has been, or may be, investigated using the discipline ...
and
rock shelter A rock shelter (also rockhouse, crepuscular cave, bluff shelter, or abri) is a shallow cave-like opening at the base of a bluff or cliff. In contrast to solutional caves (karst), which are often many miles long or wide, rock shelters are alm ...
that features
Later Stone Age The Later Stone Age (LSA) is a period in African prehistory that follows the Middle Stone Age. The Later Stone Age is associated with the advent of modern human behavior in Africa, although definitions of this concept and means of studyi ...
artifacts from during the Late
Pleistocene The Pleistocene ( ; referred to colloquially as the ''ice age, Ice Age'') is the geological epoch (geology), epoch that lasted from to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was fin ...
-
Holocene The Holocene () is the current geologic time scale, geological epoch, beginning approximately 11,700 years ago. It follows the Last Glacial Period, which concluded with the Holocene glacial retreat. The Holocene and the preceding Pleistocene to ...
transition, which is located in the forest–savanna village of Isarun in
Ondo State Ondo () is a States of Nigeria, state in Points of the compass, southwestern Nigeria. It was created on 3 February 1976 from the former Western State (Nigeria), Western State. Ondo borders Ekiti State to the north, Kogi State to the northeast fo ...
,
Nigeria Nigeria, officially the Federal Republic of Nigeria, is a country in West Africa. It is situated between the Sahel to the north and the Gulf of Guinea in the Atlantic Ocean to the south. It covers an area of . With Demographics of Nigeria, ...
. The site was initially discovered by Chief Officer J. Akeredolu, in 1961 during a large-scale survey of hilly landscapes around the town of Akure in Ondo State, Nigeria. The name was formerly and commonly known as Iwo Eleru, but the correct name is now regarded as Ihò Eléérú, or Iho Eleru, meaning "Cave of Ashes." The Iho Eleru skull is a notable archaeological discovery from the site which dates to approximately 13,000 years old. It may be evidence of modern humans possessing possible archaic human admixture or of a late-persisting
early modern human Early modern human (EMH), or anatomically modern human (AMH), are terms used to distinguish ''Homo sapiens'' (Homo sapiens sapiens, sometimes ''Homo sapiens sapiens'') that are Human anatomy, anatomically consistent with the Human variability, r ...
.


Name of archaeological site

The correct name for the archaeological site is Ihò Eléérú, or Iho Eleru, meaning "Cave of Ashes." This name originates from its common use for fire camps which resulted in a thick ash floor covering most of its superficial surface. The site was previously known as "Iwo Eleru", first reported by Chief Officer J. Akeredolu with the Department of Antiquities in Benin, Nigeria, and published by T. Shaw and S.G.H. Daniels, most probably as an incorrect anglicized translation from its original Yoruba name.


Archaeology


Archaeobotany

Remnants of endocarps have been directly dated; the results revealed that
Canarium schweinfurthii ''Canarium schweinfurthii'' (commonly known as the bush candle, African olive, African elemi, Empafu, or canarium), is a species of large tree native to tropical Africa.ICRAF Names in many African languages are variations of ''mupafu''. Descrip ...
was utilized in 11,300 cal BP as the earliest in the region of
West Africa West Africa, also known as Western Africa, is the westernmost region of Africa. The United Nations geoscheme for Africa#Western Africa, United Nations defines Western Africa as the 16 countries of Benin, Burkina Faso, Cape Verde, The Gambia, Gha ...
, and that the utilization of canarium, as well as likely
oil palm ''Elaeis'' () is a genus of palms, called oil palms, containing two species, native to Africa and the Americas. They are used in commercial agriculture in the production of palm oil. Description Mature palms are single-stemmed, and can gro ...
, occurred prior to 10,000 BP.


Ceramics

Following the emergence of pottery traditions in the Ounjougou region of
Mali Mali, officially the Republic of Mali, is a landlocked country in West Africa. It is the List of African countries by area, eighth-largest country in Africa, with an area of over . The country is bordered to the north by Algeria, to the east b ...
around 11,900 BP and in the Bosumpra region of
Ghana Ghana, officially the Republic of Ghana, is a country in West Africa. It is situated along the Gulf of Guinea and the Atlantic Ocean to the south, and shares borders with Côte d’Ivoire to the west, Burkina Faso to the north, and Togo to t ...
soon after, ceramics later arrived in the Iho Eleru region of
Nigeria Nigeria, officially the Federal Republic of Nigeria, is a country in West Africa. It is situated between the Sahel to the north and the Gulf of Guinea in the Atlantic Ocean to the south. It covers an area of . With Demographics of Nigeria, ...
.


Fauna

Some
fauna Fauna (: faunae or faunas) is all of the animal life present in a particular region or time. The corresponding terms for plants and fungi are ''flora'' and '' funga'', respectively. Flora, fauna, funga and other forms of life are collectively ...
found at Iho Eleru includes:
bushpig :''"Bush pig" may also refer to the red river hog.'' The bushpig (''Potamochoerus larvatus'') is a member of the pig family that inhabits forests, woodland, riverine vegetation and cultivated areas in East and Southern Africa. Probably introd ...
(
Potamochoerus porcus The red river hog (''Potamochoerus porcus'') or bushpig (a name also used for ''Potamochoerus larvatus'') is a wild member of the Suidae, pig family living in Africa, with most of its distribution in the Guinean Forests of West Africa, Guinean a ...
), dwarf antelope ( Neotragus batesi?), giant pouched rat (Cricetomys sp.),
rock hyrax The rock hyrax (; ''Procavia capensis''), also called dassie, Cape hyrax, rock rabbit, and (from some interpretations of a word used in the King James Bible) coney, is a medium-sized terrestrial mammal native to Africa and the Middle East. Common ...
(Procavia capensis), West African black turtle (Pelusios niger), and
yellow-backed duiker The yellow-backed duiker (''Cephalophus silvicultor'') is a shy, forest-dwelling antelope of the order Artiodactyla, from the family Bovidae. Yellow-backed duikers are the most widely-distributed of all duikers. They are found mainly in Central a ...
(Cephalophus silvicultor). Human activity, such as
foraging Foraging is searching for wild food resources. It affects an animal's fitness because it plays an important role in an animal's ability to survive and reproduce. Foraging theory is a branch of behavioral ecology that studies the foraging behavi ...
and
butchery A butcher is a person who may slaughter animals, dress their flesh, sell their meat, or participate within any combination of these three tasks. They may prepare standard cuts of meat and poultry for sale in retail or wholesale food establishme ...
, occurred at Iho Eleru. Animals that have been confirmed to have been consumed at the site include:
African buffalo The African buffalo (''Syncerus caffer)'' is a large sub-Saharan African bovine. The adult African buffalo's horns are its characteristic feature: they have fused bases, forming a continuous bone shield across the top of the head, referred to ...
(Syncerus caffer),
African savanna hare The African savanna hare (''Lepus victoriae'') is a mammal species in the family Leporidae, native to Africa. It is listed as "least concern" on the IUCN Red List. Distribution and habitat It is native to diverse regions and habitats of Africa, ...
(Lepus microtis),
crested porcupine The crested porcupine (''Hystrix cristata''), also known as the African crested porcupine, is a species of rodent in the family Hystricidae native to Italy, North Africa and sub-Saharan Africa. It is the largest porcupine species in the world. ...
(Hystrix cristata), Nile monitor (Varanus niloticus), and
Ostrich Ostriches are large flightless birds. Two living species are recognised, the common ostrich, native to large parts of sub-Saharan Africa, and the Somali ostrich, native to the Horn of Africa. They are the heaviest and largest living birds, w ...
(
Struthio camelus The common ostrich (''Struthio camelus''), or simply ostrich, is a species of flightless bird native to certain areas of Africa. It is one of two extant species of ostriches, the only living members of the genus '' Struthio'' in the ratite group ...
).


Iho Eleru skull


Discovery

The Iho Eleru site is a large rock shelter in southwestern Nigeria. The skull was found in 1965 by Thurstan Shaw and his team among over half a million
Later Stone Age The Later Stone Age (LSA) is a period in African prehistory that follows the Middle Stone Age. The Later Stone Age is associated with the advent of modern human behavior in Africa, although definitions of this concept and means of studyi ...
artifacts at the site. It was found as part of a skeleton that was buried with a thin covering of soil. The skeleton was excavated and encased in plaster, and the skull was separated from the rest of the body.


Dating

On the basis of charcoal remains that were found surrounding the skeleton, it was initially dated 9250 BC ± 200. However, in a 2011 study conducted by
Katerina Harvati Katerina Harvati (; born 1970 in Athens) is a Greek paleoanthropologist and expert in human evolution. She specializes in the broad application of 3-D geometric morphometric and virtual anthropology methods to paleoanthropology. Since 2009, she is ...
,
Chris Stringer Christopher Brian Stringer is a British physical anthropologist noted for his work on human evolution. Biography Growing up in a working-class family in the East End of London, Stringer first took an interest in anthropology during primary s ...
and others, the dating of the remains was revised: with the help of
uranium–thorium dating Uranium–thorium dating, also called thorium-230 dating, uranium-series disequilibrium dating or uranium-series dating, is a radiometric dating technique established in the 1960s which has been used since the 1970s to determine the age of calcium ...
, a time span of 11.7–16.3 ka was suggested.


Description

The
cranial vault The cranial vault is the space in the skull within the neurocranium, occupied by the brain. Development In humans, the cranial vault is imperfectly composed in newborns, to allow the large human head to pass through the birth canal. During bir ...
is relatively long and low, and the
frontal bone In the human skull, the frontal bone or sincipital bone is an unpaired bone which consists of two portions.'' Gray's Anatomy'' (1918) These are the vertically oriented squamous part, and the horizontally oriented orbital part, making up the bo ...
shows moderate recession. The
brow ridge The brow ridge, or supraorbital ridge known as superciliary arch in medicine, is a bony ridge located above the eye sockets of all primates and some other animals. In humans, the eyebrows are located on their lower margin. Structure The brow ri ...
s are moderately developed for a male and there is no pronounced nasal root. What remains of the nasal area suggest that the
nasal bridge The nasal bridge is the upper part of the nose, where the nasal bones and surrounding soft tissues provide structural support. While commonly discussed in human anatomy, nasal bridges exist in various forms across many vertebrates, particularl ...
was relatively flat, and the evidence from X-rays points to little
frontal sinus The frontal sinuses are one of the four pairs of paranasal sinuses that are situated behind the brow ridges. Sinuses are mucosa-lined airspaces within the bones of the face and skull. Each opens into the anterior part of the corresponding middle ...
development. The upper face is missing except for a small collection of fragments. Parts of the maxillary-molar region have been identified (including the
infraorbital foramen In human anatomy, the infraorbital foramen is one of two small holes in the skull's upper jawbone ( maxillary bone), located below the eye socket and to the left and right of the nose. Both holes are used for blood vessels and nerves. In anatomic ...
) and, based on what survives, it is unlikely that the upper face was large. The
mandible In jawed vertebrates, the mandible (from the Latin ''mandibula'', 'for chewing'), lower jaw, or jawbone is a bone that makes up the lowerand typically more mobilecomponent of the mouth (the upper jaw being known as the maxilla). The jawbone i ...
is well developed and has a masculine appearance, although there is no pronounced chin. Apart from two lower premolars, the teeth are not attached to the jaws and it is uncertain where the surviving teeth were originally placed. All the
anterior teeth In dentistry, the term anterior teeth usually refers as a group to the incisors and canine teeth as distinguished from the posterior teeth, which are the premolars and molars. The distinction is one of anterior (front of the body) versus posterio ...
show noticeable attrition and most of the
crown A crown is a traditional form of head adornment, or hat, worn by monarchs as a symbol of their power and dignity. A crown is often, by extension, a symbol of the monarch's government or items endorsed by it. The word itself is used, parti ...
has been eroded by wear. Based on the evidence of
tooth wear Tooth wear refers to loss of tooth substance by means other than dental caries. Tooth wear is a very common condition that occurs in approximately 97% of the population. This is a normal physiological process occurring throughout life; but with i ...
, the age of Iho Eleru fossil has been estimated as over 30 years. What remains of the rest of the skeleton are generally crushed fragments of large bones. The shafts of the
humeri The humerus (; : humeri) is a long bone in the arm that runs from the shoulder to the elbow. It connects the scapula and the two bones of the lower arm, the radius and ulna, and consists of three sections. The humeral upper extremity consists of ...
appear robust and the
cortical bone A bone is a rigid organ that constitutes part of the skeleton in most vertebrate animals. Bones protect the various other organs of the body, produce red and white blood cells, store minerals, provide structure and support for the body, an ...
is moderately thick. The shafts of the
radius In classical geometry, a radius (: radii or radiuses) of a circle or sphere is any of the line segments from its Centre (geometry), center to its perimeter, and in more modern usage, it is also their length. The radius of a regular polygon is th ...
and
femur The femur (; : femurs or femora ), or thigh bone is the only long bone, bone in the thigh — the region of the lower limb between the hip and the knee. In many quadrupeds, four-legged animals the femur is the upper bone of the hindleg. The Femo ...
are also robust. The existing remains suggest he was of medium height and build, and was no taller than approximately 165cm.


Analysis

Don Brothwell and Thurstan Shaw said in 1971 that the sloping frontal vault was more pronounced in Iho Eleru fossil than in both later Neolithic and recent sub-Saharan skull samples. However, they also found that the
occipital The occipital bone () is a cranial dermal bone and the main bone of the occiput (back and lower part of the skull). It is trapezoidal in shape and curved on itself like a shallow dish. The occipital bone lies over the occipital lobes of the cere ...
structure, nasal root and the frontal bone of the skull "would qualify for identification as that of a proto-West African negro." In 1974 Chris Stringer said that there were surprising similarities between the crania of the much older
Solo Man Solo Man (''Homo erectus soloensis'') is a subspecies of '' H. erectus'' that lived along the Solo River in Java, Indonesia, about 117,000 to 108,000 years ago in the Late Pleistocene. This population is the last known record of the species. I ...
and Omo II with that of Iho Eleru. The 2011 study found that "Iwo Eleru possesses neurocranial morphology intermediate in shape between archaic hominins (
Neanderthal Neanderthals ( ; ''Homo neanderthalensis'' or sometimes ''H. sapiens neanderthalensis'') are an extinction, extinct group of archaic humans who inhabited Europe and Western and Central Asia during the Middle Pleistocene, Middle to Late Plei ...
s and ''
Homo erectus ''Homo erectus'' ( ) is an extinction, extinct species of Homo, archaic human from the Pleistocene, spanning nearly 2 million years. It is the first human species to evolve a humanlike body plan and human gait, gait, to early expansions of h ...
'') and modern humans."Harvati et al, 'The Later Stone Age Calvaria from Iwo Eleru, Nigeria: Morphology and Chronology', p. 6. The authors of the study asserted that the dating of Iho Eleru fossil to the late
Pleistocene The Pleistocene ( ; referred to colloquially as the ''ice age, Ice Age'') is the geological epoch (geology), epoch that lasted from to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was fin ...
"implies that the transition to anatomical modernity in Africa was more complicated than previously thought, with late survival of “archaic” features and possibly deep population substructure in Africa during this time." It has been argued that the Iho Eleru fossil was an archaic hybrid or part of a relict archaic ''Homo'' population. In 2014 Christopher Stojanowski of
Arizona State University Arizona State University (Arizona State or ASU) is a public university, public research university in Tempe, Arizona, United States. Founded in 1885 as Territorial Normal School by the 13th Arizona Territorial Legislature, the university is o ...
summarised the three dominant explanations for Iho Eleru fossil's atypical cranial shape: the first, that Iho Eleru was a hybrid with archaic African populations; the second, that Iho Eleru fossil was a member of a relict archaic population that was replaced by more modern humans upon the onset of the Holocene era; and the third, that Iho Eleru fossil was part of a population that diverged from the rest of North Africa's populations during a time of acute aridness in the
Sahara The Sahara (, ) is a desert spanning across North Africa. With an area of , it is the largest hot desert in the world and the list of deserts by area, third-largest desert overall, smaller only than the deserts of Antarctica and the northern Ar ...
desert that made it impassable until the arrival of the
African humid period The African humid period (AHP; also known by other names) was a climate period in Africa during the late Pleistocene and Holocene geologic epochs, when northern Africa was wetter than today. The covering of much of the Sahara desert by grass ...
. In 2014 Peter J. Waddell of
Massey University Massey University () is a Public university, public research university in New Zealand that provides internal and distance education. The university has campuses in Auckland, Palmerston North, and Wellington. Data from Universities New Zealand ...
argued that Iho Eleru man descended from a lineage 200–400 kya and whose extinction may have been caused by humans. Waddell also said: "Such a long apparently distinct lineage that terminated in West Africa perhaps 12kya, with no obvious sign of living descendants, suggests that the Iho Eleru lineage quite probably represents a distinct species of near modern human. As such, the species name ''Homo iwoelerueensis'' suggests itself." However, the
University of Washington The University of Washington (UW and informally U-Dub or U Dub) is a public research university in Seattle, Washington, United States. Founded in 1861, the University of Washington is one of the oldest universities on the West Coast of the Uni ...
's Fred L. Bookstein cautions against naming the fossil as a new species until more confirmatory evidence is discovered.Fred L. Bookstein, '“Like Fixing an Airpline in Flight”: On Paleoanthropology as an Evolutionary Discipline, or, Paleoanthropology for What?', in Schwartz (ed.), ''Rethinking Human Evolution'', p. 198.


Stone tools

Roughly half of a million stone tools have been discovered at Iho Eleru.


See also

*
Asselar man Asselar man is a Neolithic (Later Stone Age) skeleton found at Adrar des Ifoghas, Mali, which has been dated to between 9500 BP and 7000 BP, amid the early Holocene Wet Phase. The Asselar skeleton was likely intentionally buried. Geography A ...


References


Bibliography

* {{DEFAULTSORT:Iho Eleru skull Peopling of Africa Homo fossils 1965 archaeological discoveries Archaeology of Nigeria Archaeological sites of Western Africa