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''Icelinus''is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial n ...
of marine
ray-finned fish Actinopterygii (; ), members of which are known as ray-finned fishes, is a class of bony fish. They comprise over 50% of living vertebrate species. The ray-finned fishes are so called because their fins are webs of skin supported by bony or h ...
es belonging to the
family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
Cottidae The Cottidae are a family of fish in the superfamily Cottoidea, the sculpins. It is the largest sculpin family, with about 275 species in 70 genera.Kane, E. A. and T. E. Higham. (2012)Life in the flow lane: differences in pectoral fin morphology ...
, the typical sculpins. These fishes are found in the northern and eastern
Pacific Ocean The Pacific Ocean is the largest and deepest of Earth's five oceanic divisions. It extends from the Arctic Ocean in the north to the Southern Ocean (or, depending on definition, to Antarctica) in the south, and is bounded by the contine ...
.


Taxonomy

''Icelinus'' was first proposed as a monospecific genus in 1885 by the American
ichthyologist Ichthyology is the branch of zoology devoted to the study of fish, including bony fish (Osteichthyes), cartilaginous fish (Chondrichthyes), and jawless fish (Agnatha). According to FishBase, 33,400 species of fish had been described as of October ...
David Starr Jordan with its only and
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen( ...
being ''Artedius quadriseriatus''. This species had been described in 1880 by the English zoologist
William Neale Lockington William Neale Lockington (1840 in Rugby, Warwickshire, England – 1902 in Worthing, Sussex) was an English zoologist. California Lockington was the curator of the California Academy of Sciences museum in San Francisco, California ...
from
San Francisco San Francisco (; Spanish language, Spanish for "Francis of Assisi, Saint Francis"), officially the City and County of San Francisco, is the commercial, financial, and cultural center of Northern California. The city proper is the List of Ca ...
. The 5th edition of ''
Fishes of the World ''Fishes of the World'' by the American Ichthyology, ichthyologist Joseph S. Nelson (1937–2011) is a standard reference for fish systematics. Now in its fifth edition (2016), the work is a comprehensive overview of the diversity and classificat ...
'' classifies this genus within the subfamily
Cottinae Cottinae is a subfamily of ray-finned fishes belonging to the family Cottidae, the typical sculpin A sculpin is a type of fish that belongs to the superfamily Cottoidea in the order Scorpaeniformes.Kane, E. A. and T. E. Higham. (2012)Life in ...
of the family Cottidae, however, other authors classify the genus within the subfamily Oligocottinae of the family
Psychrolutidae The fish family Psychrolutidae (commonly known as blobfishes, toadfishes, flathead sculpins, tadpole sculpins,) contains over 35 recognized species in 8 genera. This family consists of bottom-dwelling marine sculpins shaped like tadpoles, with la ...
. The genus ''Icelinus'' may not be
monophyletic In cladistics for a group of organisms, monophyly is the condition of being a clade—that is, a group of taxa composed only of a common ancestor (or more precisely an ancestral population) and all of its lineal descendants. Monophyletic ...
as a study found that the 9 eastern Pacific species were in a clade which was a
sister taxon In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an evolutionary tree. Definition The expression is most easily illustrated by a cladogram: Taxon A and ...
to the genera ''
Furcina ''Furcina'' is a small genus of marine ray-finned fishes belonging to the family Cottidae, the typical sculpins. This species is found in the northwestern Pacific Ocean from around Japan and the Republic of Korea. Taxonomy ''Furcina'' was first ...
'' and ''
Antipodocottus ''Antipodocottus'' is a genus of marine ray-finned fishes belonging to the family Cottidae, the typical sculpins. These fishes are found in the western Pacific Ocean. This is the only genus of Cottids represented in the Southern Hemisphere, altho ...
'' while the 2 northwestern Pacific species (''I. japonicus'' and ''I. pietschi'') were found to be so closely related to the genus ''
Stlengis ''Stlengis'' is a genus of marine ray-finned fishes belonging to the family Cottidae, the typical sculpins. These fishes are found in the northwestern Pacific Ocean where they are only known from the waters around Japan. Taxonomy ''Stlengis' ...
'' that they were placed within that genus, leaving ''Icelinus'' confined to the eastern Pacific.


Etymology

''Icelinus'' is a diminutive of ''
Icelus Icelus may refer to: * Icelus or Icelos, another name for Phobetor, in Ovid's ''Metamorphoses'' * ''Icelus'' (fish), a genus of fishes * Icelus, a freedman who served the Roman Emperor Galba before his downfall * A character in ''Saint Seiya: Th ...
'', the genus ''I. quadriseriatus'' was thought to belong to.


Species

There are currently 11 recognized species in this genus: * ''
Icelinus borealis ''Icelinus borealis'', or the northern sculpin or comb sculpin, is a species of fish in the family Cottidae. It can be found in the northeastern Pacific Ocean along the western coast of North America. Taxonomy ''Icelinus borealis'' was first form ...
'' C. H. Gilbert, 1896 (Northern sculpin) * ''
Icelinus burchami ''Icelinus''is a genus of marine ray-finned fishes belonging to the family Cottidae, the typical sculpins. These fishes are found in the northern and eastern Pacific Ocean. Taxonomy ''Icelinus'' was first proposed as a monospecific genus in 1885 ...
''
Evermann Barton Warren Evermann (October 24, 1853 – September 27, 1932) was an American ichthyologist. Early life and education Evermann was born in Monroe County, Iowa in 1853. His family moved to Indiana while he was still a child and it was ...
& Goldsborough, 1907
(Dusky sculpin) * ''
Icelinus cavifrons ''Icelinus''is a genus of marine ray-finned fishes belonging to the family Cottidae, the typical sculpins. These fishes are found in the northern and eastern Pacific Ocean. Taxonomy ''Icelinus'' was first proposed as a monospecific genus in 1885 ...
'' C. H. Gilbert, 1890 (Pit-head sculpin) * ''
Icelinus filamentosus The threadfin sculpin (''Icelinus filamentosus'') is a fish, mostly found in the Pacific waters of North America. A large, deep-water sculpin of the family Cottidae, it occurs in the North Pacific Ocean, from Chirikof Island in the western Gulf ...
'' C. H. Gilbert, 1890 (Threadfin sculpin) * ''
Icelinus fimbriatus ''Icelinus''is a genus of marine ray-finned fishes belonging to the family Cottidae, the typical sculpins. These fishes are found in the northern and eastern Pacific Ocean. Taxonomy ''Icelinus'' was first proposed as a monospecific genus in 1885 ...
'' C. H. Gilbert, 1890 (Fringed sculpin) * ''
Icelinus japonicus ''Icelinus''is a genus of marine ray-finned fishes belonging to the Family (biology), family Cottidae, the typical sculpins. These fishes are found in the northern and eastern Pacific Ocean. Taxonomy ''Icelinus'' was first proposed as a monospec ...
''
Yabe was a village located in Yame District, Fukuoka Prefecture, Japan. Yabe is located at 330m above sea level. The average temperature is 14 °C and the average annual precipitation is 2,706mm. As of 2006, the village had an estimated popul ...
, Tsumura &
Katayama is a Japanese surname. Notable people with the surname include: Entertainers *, Japanese stage actress *, Japanese film actress *, Japanese film actress and singer, former AKB48 member Politicians *, Japanese-born member of the American Communi ...
, 1980
* ''
Icelinus limbaughi ''Icelinus''is a genus of marine ray-finned fishes belonging to the family Cottidae, the typical sculpins. These fishes are found in the northern and eastern Pacific Ocean. Taxonomy ''Icelinus'' was first proposed as a monospecific genus in 1885 ...
''
Rosenblatt Rosenblatt is a surname of German and Jewish origin, meaning " rose leaf". People with this surname include: *Albert Rosenblatt (born 1936), New York Court of Appeals judge *Dana Rosenblatt, known as "Dangerous" (born 1972), American boxer *Elie R ...
& W. L. Smith, 2004
(Canyon sculpin) * ''
Icelinus oculatus ''Icelinus''is a genus of marine ray-finned fishes belonging to the family Cottidae, the typical sculpins. These fishes are found in the northern and eastern Pacific Ocean. Taxonomy ''Icelinus'' was first proposed as a monospecific genus in 1885 ...
'' C. H. Gilbert, 1890 (Frogmouth sculpin) * '' Icelinus pietschi'' Yabe,
Soma Soma may refer to: Businesses and brands * SOMA (architects), a New York–based firm of architects * Soma (company), a company that designs eco-friendly water filtration systems * SOMA Fabrications, a builder of bicycle frames and other bicycl ...
& Amaoka, 2001
* ''
Icelinus quadriseriatus ''Icelinus''is a genus of marine ray-finned fishes belonging to the family Cottidae, the typical sculpins. These fishes are found in the northern and eastern Pacific Ocean. Taxonomy ''Icelinus'' was first proposed as a monospecific genus in 1885 ...
'' ( Lockington, 1880) (Yellowchin sculpin) * ''
Icelinus tenuis ''Icelinus''is a genus of marine ray-finned fishes belonging to the family Cottidae, the typical sculpins. These fishes are found in the northern and eastern Pacific Ocean. Taxonomy ''Icelinus'' was first proposed as a monospecific genus in 1885 ...
'' C. H. Gilbert, 1890 (Spotfin sculpin)


Characteristics

''Icelinus'' sculpins are characterised by having the fourth uppermost spine on the
preoperculum This glossary of ichthyology is a list of definitions of terms and concepts used in ichthyology, the study of fishes. A B C ...
having a number of points like the
antler Antlers are extensions of an animal's skull found in members of the Cervidae (deer) family. Antlers are a single structure composed of bone, cartilage, fibrous tissue, skin, nerves, and blood vessels. They are generally found only on ...
of a deer. They have a single spine and 2 soft rays in the
pelvic fin Pelvic fins or ventral fins are paired fins located on the ventral surface of fish. The paired pelvic fins are homologous to the hindlimbs of tetrapods. Structure and function Structure In actinopterygians, the pelvic fin consists of two e ...
and there are two rows of
ctenoid scale A fish scale is a small rigid plate that grows out of the skin of a fish. The skin of most jawed fishes is covered with these protective scales, which can also provide effective camouflage through the use of reflection and colouration, as w ...
s along the
dorsal fin A dorsal fin is a fin located on the back of most marine and freshwater vertebrates within various taxa of the animal kingdom. Many species of animals possessing dorsal fins are not particularly closely related to each other, though through c ...
bases. These are small fishes, the largest species in the genus is ''I. filamentosus'', which has a maximum published
total length Fish measurement is the measuring of individual fish and various parts of their anatomies. These data are used in many areas of ichthyology, including taxonomy and fisheries biology. Overall length * Standard length (SL) is the length of a fish ...
of , while the smallest is ''I. piestchi'' which has a maximum published
standard length Fish measurement is the measuring of individual fish and various parts of their anatomies. These data are used in many areas of ichthyology, including taxonomy and fisheries biology. Overall length * Standard length (SL) is the length of a fish ...
of .


Distribution

''Icelinus'' sculpins are found in the northern and eastern Pacific Ocean. They are inshore fishes.


References

{{Taxonbar, from=Q142573 Cottinae Marine fish genera Taxa named by David Starr Jordan