''Icarops'' is an extinct, possibly

paraphyletic Paraphyly is a taxonomic term describing a grouping that consists of the grouping's last common ancestor and some but not all of its descendant lineages. The grouping is said to be paraphyletic ''with respect to'' the excluded subgroups. In co ...

genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...

mystacine __NOTOC__ Mystacinidae is a family of unusual bats, the New Zealand short-tailed bats. There is one living genus, ''Mystacina'', with two species, one of which could have possibly become extinct in the 1960s. They are medium-sized bats, about in ...

bat Bats are flying mammals of the order Chiroptera (). With their forelimbs adapted as wings, they are the only mammals capable of true and sustained flight. Bats are more agile in flight than most birds, flying with their very long spread-out ...

with three described species. The genus is known from fossils found at Riversleigh, north-western

Queensland Queensland ( , commonly abbreviated as Qld) is a States and territories of Australia, state in northeastern Australia, and is the second-largest and third-most populous state in Australia. It is bordered by the Northern Territory, South Austr ...

, Bullock Creek,

Northern Territory The Northern Territory (abbreviated as NT; known formally as the Northern Territory of Australia and informally as the Territory) is an states and territories of Australia, Australian internal territory in the central and central-northern regi ...

, and

Lake Ngapakaldi to Lake Palankarinna Fossil Area __NOTOC__ The Lake Ngapakaldi to Lake Palankarinna Fossil Area is a group of fossil sites located in the Australian state of South Australia within the Tirari Desert in the north-eastern part of the state's Far North region. The group has an ...

South Australia South Australia (commonly abbreviated as SA) is a States and territories of Australia, state in the southern central part of Australia. With a total land area of , it is the fourth-largest of Australia's states and territories by area, which in ...

Australia Australia, officially the Commonwealth of Australia, is a country comprising mainland Australia, the mainland of the Australia (continent), Australian continent, the island of Tasmania and list of islands of Australia, numerous smaller isl ...

. The

fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...

s date from the late

Oligocene The Oligocene ( ) is a geologic epoch (geology), epoch of the Paleogene Geologic time scale, Period that extends from about 33.9 million to 23 million years before the present ( to ). As with other older geologic periods, the rock beds that defin ...

to early

Miocene The Miocene ( ) is the first epoch (geology), geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and mea ...

. The name was derived from a figure of Greek mythology,

Icarus In Greek mythology, Icarus (; , ) was the son of the master craftsman Daedalus, the architect of the labyrinth of Crete. After Theseus, king of Athens and enemy of King Minos, escaped from the labyrinth, Minos suspected that Icarus and Daedalu ...

, who the authors noted, "flew towards the sun, in reference to the ancient mystacinid that flew eastwards from Australia to New Zealand". A study describing the genus ''

Vulcanops ''Vulcanops jennyworthyae'' is an extinct species of bat that lived during the Miocene in New Zealand, a large burrowing microchiropteran that probably ate arthropods and plant material around twenty million years before present. It is the ty ...

'' renders ''Icarops'' paraphyletic towards the rest of Mystacinidae, with ''I. paradox'' being closer to New Zealand mystacines than to other Australian mystacines, which form an independent clade. The described species are * ''Icarops'' :*''Icarops aenae'' :*''Icarops breviceps'' :*''Icarops paradox''

Terrestriality

Like its modern relatives, the ''

Mystacina ''Mystacina'' is the sole surviving genus of the Mystacinidae family of bats. It has three known species, of which only the New Zealand lesser short-tailed bat (''Mystacina tuberculata'') is confirmed to survive today. The closely related New Ze ...

''short-tailed bats, ''Icarops'' shows adaptations to foraging on the ground. This is in spite of occurring alongside various terrestrial tetrapods, including other mammals such as

marsupial Marsupials are a diverse group of mammals belonging to the infraclass Marsupialia. They are natively found in Australasia, Wallacea, and the Americas. One of marsupials' unique features is their reproductive strategy: the young are born in a r ...

s and

monotreme Monotremes () are mammals of the order Monotremata. They are the only group of living mammals that lay eggs, rather than bearing live young. The extant monotreme species are the platypus and the four species of echidnas. Monotremes are typified ...

s. This shows that the terrestrial habits of

s did not evolve due to lack of competition with other mammals in

New Zealand New Zealand () is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and List of islands of New Zealand, over 600 smaller islands. It is the List of isla ...

, predating the island's colonisation and having evolved on mainland Australia.

References

{{Taxonbar, from=Q5985076 Miocene mammals of Australia Miocene bats Oligocene bats Prehistoric bat genera Mystacinidae Riversleigh fauna Bats of New Zealand