Ibycter
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The red-throated caracara (''Ibycter americanus'') is a social species of
bird of prey Birds of prey or predatory birds, also known as (although not the same as) raptors, are hypercarnivorous bird species that actively predation, hunt and feed on other vertebrates (mainly mammals, reptiles and smaller birds). In addition to speed ...
in the family
Falconidae The falcons and caracaras are around 65 species of Diurnality, diurnal birds of prey that make up the family Falconidae (representing all extant species in the order (biology), order Falconiformes). The family likely originated in South America d ...
. It is placed in the
monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unisp ...
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
''Ibycter'', or sometimes united in ''
Daptrius The black caracara (''Daptrius ater'') is a species of bird of prey in the family Falconidae found in Amazonian and French Guianan lowlands, occurring commonly along rivers. They are locally referred to as ''Ger' futu busikaka'' in the Republ ...
'' with the
black caracara The black caracara (''Daptrius ater'') is a species of bird of prey in the family Falconidae found in Amazonian and French Guianan lowlands, occurring commonly along rivers. They are locally referred to as ''Ger' futu busikaka'' in the Republi ...
. Unique among caracaras, it mainly feeds on the
larva A larva (; : larvae ) is a distinct juvenile form many animals undergo before metamorphosis into their next life stage. Animals with indirect development such as insects, some arachnids, amphibians, or cnidarians typically have a larval phase ...
e of
bee Bees are winged insects closely related to wasps and ants, known for their roles in pollination and, in the case of the best-known bee species, the western honey bee, for producing honey. Bees are a monophyletic lineage within the superfamil ...
s and
wasp A wasp is any insect of the narrow-waisted suborder Apocrita of the order Hymenoptera which is neither a bee nor an ant; this excludes the broad-waisted sawflies (Symphyta), which look somewhat like wasps, but are in a separate suborder ...
s, but also takes the adult insects and
fruit In botany, a fruit is the seed-bearing structure in flowering plants (angiosperms) that is formed from the ovary after flowering. Fruits are the means by which angiosperms disseminate their seeds. Edible fruits in particular have long propaga ...
s and
berries A berry is a small, pulpy, and often edible fruit. Typically, berries are juicy, rounded, brightly colored, sweet, sour or tart, and do not have a stone fruit, stone or pit (fruit), pit although many wikt:pip#Etymology 2, pips or seeds may be p ...
. It is found from far southern Mexico through parts of Central and South America south to Peru, Bolivia, and Brazil. Its natural
habitat In ecology, habitat refers to the array of resources, biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species' habitat can be seen as the physical manifestation of its ...
s are
subtropical The subtropical zones or subtropics are geographical zone, geographical and Köppen climate classification, climate zones immediately to the Northern Hemisphere, north and Southern Hemisphere, south of the tropics. Geographically part of the Ge ...
or tropical moist lowland
forest A forest is an ecosystem characterized by a dense ecological community, community of trees. Hundreds of definitions of forest are used throughout the world, incorporating factors such as tree density, tree height, land use, legal standing, ...
s and subtropical or tropical moist
montane forest Montane ecosystems are found on the slopes of mountains. The alpine climate in these regions strongly affects the ecosystem because temperatures lapse rate, fall as elevation increases, causing the ecosystem to stratify. This stratification is ...
s.


Taxonomy

The red-throated caracara was described by the French polymath
Georges-Louis Leclerc, Comte de Buffon Georges-Louis Leclerc, Comte de Buffon (; 7 September 1707 – 16 April 1788) was a French Natural history, naturalist, mathematician, and cosmology, cosmologist. He held the position of ''intendant'' (director) at the ''Jardin du Roi'', now ca ...
in 1770 in his ''Histoire Naturelle des Oiseaux'' from a specimen collected in
Cayenne Cayenne (; ; ) is the Prefectures in France, prefecture and capital city of French Guiana, an overseas region and Overseas department, department of France located in South America. The city stands on a former island at the mouth of the Caye ...
,
French Guiana French Guiana, or Guyane in French, is an Overseas departments and regions of France, overseas department and region of France located on the northern coast of South America in the Guianas and the West Indies. Bordered by Suriname to the west ...
. The bird was also illustrated in a hand-coloured plate engraved by
François-Nicolas Martinet François-Nicolas Martinet (1731 - c. 1800) was a French engineer, engraver and naturalist. Although trained as an engineer and draftsman, he began to produce engravings for books and it later became his primary profession. Martinet's year of b ...
in the ''Planches Enluminées D'Histoire Naturelle'' which was produced under the supervision of
Edme-Louis Daubenton Edme-Louis Daubenton (12 August 1730 – 12 December 1785) was a French naturalist. Daubenton was the cousin of another French naturalist, Louis Jean-Marie Daubenton. Georges-Louis Leclerc, the Comte de Buffon engaged Edme-Louis Daubenton to su ...
to accompany Buffon's text. Neither the plate caption nor Buffon's description included a scientific name but in 1783 the Dutch naturalist
Pieter Boddaert Pieter Boddaert (1730 – 6 May 1795) was a Dutch physician and naturalist. Early life, family and education Boddaert was the son of a Middelburg jurist and poet by the same name (1694–1760). The younger Pieter obtained his M.D. at the Univers ...
coined the
binomial name In taxonomy, binomial nomenclature ("two-term naming system"), also called binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, both of which use Latin grammatical forms, altho ...
''Falco americanus'' in his catalogue of the ''Planches Enluminées''. The red-throated caracara was for many years placed with the
black caracara The black caracara (''Daptrius ater'') is a species of bird of prey in the family Falconidae found in Amazonian and French Guianan lowlands, occurring commonly along rivers. They are locally referred to as ''Ger' futu busikaka'' in the Republi ...
in the
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
''
Daptrius The black caracara (''Daptrius ater'') is a species of bird of prey in the family Falconidae found in Amazonian and French Guianan lowlands, occurring commonly along rivers. They are locally referred to as ''Ger' futu busikaka'' in the Republ ...
'' but based on a molecular genetic study published in 1999 it was moved to be the only species in the resurrected genus ''Ibycter'' that had been introduced by the French ornithologist
Louis Pierre Vieillot Louis Pierre Vieillot (10 May 1748, Yvetot – 24 August 1830, Sotteville-lès-Rouen) was a French ornithologist. Vieillot is the author of the first scientific descriptions and Linnaean names of a number of birds, including species he collected ...
in 1816. The species is
monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unisp ...
. The genus name ''Ibycter '' is from the
Ancient Greek Ancient Greek (, ; ) includes the forms of the Greek language used in ancient Greece and the classical antiquity, ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Greek ...
''ibuktēr'' meaning "singer of war-songs".


Description

Males average 20.1 in (51 cm) long, while females average 22.1 in (56 cm); they are distinguished from the
black caracara The black caracara (''Daptrius ater'') is a species of bird of prey in the family Falconidae found in Amazonian and French Guianan lowlands, occurring commonly along rivers. They are locally referred to as ''Ger' futu busikaka'' in the Republi ...
by larger size and
plumage Plumage () is a layer of feathers that covers a bird and the pattern, colour, and arrangement of those feathers. The pattern and colours of plumage differ between species and subspecies and may vary with age classes. Within species, there can b ...
that is mainly black, with the belly, tail feathers, and undertail feathers being white. Both their faces and throats are bare with a few black feathers scattered on the throat; the exposed skin is red. Both male and female red-throated caracaras are similar in appearance. Males have a wing length of 35.55 cm, a tail length of 24.96 cm, a bill length of 2.5 cm, and a tarsus length of 5.41 cm. Females have a wing length of 35.93 cm, a tail length of 25.31 cm, a bill length of 2.58 cm, and a tarsus length of 5.62 cm.


Distribution and habitat

This species inhabits the humid lowland forests of
Brazil Brazil, officially the Federative Republic of Brazil, is the largest country in South America. It is the world's List of countries and dependencies by area, fifth-largest country by area and the List of countries and dependencies by population ...
,
Bolivia Bolivia, officially the Plurinational State of Bolivia, is a landlocked country located in central South America. The country features diverse geography, including vast Amazonian plains, tropical lowlands, mountains, the Gran Chaco Province, w ...
,
Colombia Colombia, officially the Republic of Colombia, is a country primarily located in South America with Insular region of Colombia, insular regions in North America. The Colombian mainland is bordered by the Caribbean Sea to the north, Venezuel ...
,
Costa Rica Costa Rica, officially the Republic of Costa Rica, is a country in Central America. It borders Nicaragua to the north, the Caribbean Sea to the northeast, Panama to the southeast, and the Pacific Ocean to the southwest, as well as Maritime bo ...
,
El Salvador El Salvador, officially the Republic of El Salvador, is a country in Central America. It is bordered on the northeast by Honduras, on the northwest by Guatemala, and on the south by the Pacific Ocean. El Salvador's capital and largest city is S ...
,
Ecuador Ecuador, officially the Republic of Ecuador, is a country in northwestern South America, bordered by Colombia on the north, Peru on the east and south, and the Pacific Ocean on the west. It also includes the Galápagos Province which contain ...
,
French Guiana French Guiana, or Guyane in French, is an Overseas departments and regions of France, overseas department and region of France located on the northern coast of South America in the Guianas and the West Indies. Bordered by Suriname to the west ...
,
Guatemala Guatemala, officially the Republic of Guatemala, is a country in Central America. It is bordered to the north and west by Mexico, to the northeast by Belize, to the east by Honduras, and to the southeast by El Salvador. It is hydrologically b ...
,
Guyana Guyana, officially the Co-operative Republic of Guyana, is a country on the northern coast of South America, part of the historic British West Indies. entry "Guyana" Georgetown, Guyana, Georgetown is the capital of Guyana and is also the co ...
,
Honduras Honduras, officially the Republic of Honduras, is a country in Central America. It is bordered to the west by Guatemala, to the southwest by El Salvador, to the southeast by Nicaragua, to the south by the Pacific Ocean at the Gulf of Fonseca, ...
,
Mexico Mexico, officially the United Mexican States, is a country in North America. It is the northernmost country in Latin America, and borders the United States to the north, and Guatemala and Belize to the southeast; while having maritime boundar ...
,
Nicaragua Nicaragua, officially the Republic of Nicaragua, is the geographically largest Sovereign state, country in Central America, comprising . With a population of 7,142,529 as of 2024, it is the third-most populous country in Central America aft ...
,
Panama Panama, officially the Republic of Panama, is a country in Latin America at the southern end of Central America, bordering South America. It is bordered by Costa Rica to the west, Colombia to the southeast, the Caribbean Sea to the north, and ...
,
Peru Peru, officially the Republic of Peru, is a country in western South America. It is bordered in the north by Ecuador and Colombia, in the east by Brazil, in the southeast by Bolivia, in the south by Chile, and in the south and west by the Pac ...
, and
Venezuela Venezuela, officially the Bolivarian Republic of Venezuela, is a country on the northern coast of South America, consisting of a continental landmass and many Federal Dependencies of Venezuela, islands and islets in the Caribbean Sea. It com ...
. The slow flight of the red-throated caracara makes it suited it to fly in the
understory In forestry and ecology, understory (American English), or understorey (English in the Commonwealth of Nations, Commonwealth English), also known as underbrush or undergrowth, includes plant life growing beneath the Canopy (biology), forest ca ...
of the forest where the vegetation is thin. The sparse vegetation gives the red-throated caracara greater visibility to spot food and predators. The
ornate hawk-eagle The ornate hawk-eagle (''Spizaetus ornatus'') is a fairly large bird of prey from the Tropics, tropical Americas. Formerly, some authorities referred to this species as the crested hawk-eagle, a name that may cause some confusion as it is more co ...
and the black-and-white hawk-eagle are predators of the red-throated caracara.


Behaviour and ecology


Food and feeding

The red-throated caracara hunts in the
canopy Canopy may refer to: Plants * Canopy (biology), aboveground portion of plant community or crop (including forests) * Canopy (grape), aboveground portion of grapes Religion and ceremonies * Baldachin or canopy of state, typically placed over an a ...
and the understory of the lowland jungle, foraging mainly for insect nests. Most red-throated caracaras hunt silently, but occasionally make soft caws and sometimes hunt in groups. When hunting in groups, one or two individuals scout for predators in the canopy, while the remaining flock hunts in the understory. The red-throated caracara is highly territorial, with four to eight individuals in a group.Life History - Red-throated Caracara (Ibycter americanus) - Neotropical Birds
/ref> The diet consists mainly of wasp and bee larvae, though it will eat mature insects and also forage on fruits and berries found in the humid subtropical and tropical lowlands, and mountainous regions of its Central and South American habitat. Biodiversity of the forest
ecosystem An ecosystem (or ecological system) is a system formed by Organism, organisms in interaction with their Biophysical environment, environment. The Biotic material, biotic and abiotic components are linked together through nutrient cycles and en ...
is paramount for the birds' special diet, since wasps and bees often make their nests in hollows or amongst branches of mature trees found in old-growth forests. Deforestation and intensive agriculture practices severely hamper the red-throated caracara's population, likely accounting for its rare sightings today. After the 1950s, both its population and range rapidly declined in Costa Rica, Honduras, Panama, Ecuador, and French Guiana, causing the species to be placed on the World Wildlife endangered list. Until 2013, very little was known of the red-throated caracara's feeding behavior until a team of Canadian biologists from the University of Simon Fraser spent months researching the birds using camera surveillance at the Nouragues Field Station in French Guiana. The scientific footage shows the birds using a rapid-fire "fly-by" aerial-diving attack strategy to knock nests down onto the forest floor, while skillfully evading most wasp stings. The birds use air squadron precision, repeatedly diving then scooping upward, to drive off or confuse angry defender swarms around the hive.Birds outsmart wasps to feed young - SFU News - Simon Fraser University
/ref> Researchers also found that neotropical defender wasps eventually abandon their damaged hives and retreat, alongside smaller worker wasps, to rebuild a new nest site. All predators evolve ways of hunting or trapping prey. Biologist Sean McCann observed that these intelligent birds have a highly specialized predation trait in response the wasps’ behavior to cut losses and rebuild elsewhere. The predation impact on the numbers of prey populations is undetermined. Furthermore, it is not clear how much the red-throated caracara's primary food source, wasp larvae, places constraints on the birds' ability to survive since their complex predation is interlinked with neotropical wasp behavior. Knowledge of the birds' chemical resistance to stings is also unknown. Chemical traces found on the birds’ feet are similar to those secreted from Azteca ants, likely contacted along tree branches and nest sites which both species inhabit.


References


Further reading

*


External links


Red-throated caracara videos
on the Internet Bird Collection

VIREO * ttp://www.pinkmoose.ic24.net/images/red-faced-caracara.jpg Photo-Medium Res ttp://www.pinkmoose.ic24.net/peru/amazonia.htm Articlepinkmoose–"Amazonia Lodge"
Photo-Medium Res
https://www.nhlstenden.com/;"Suriname Birds" {{Taxonbar, from1=Q517757, from2=Q10772622 Polyborinae Birds of Central America Birds of Brazil Birds of Colombia Birds of Venezuela Birds of Ecuador Birds of the Guiana Shield Birds of the Amazon rainforest Birds described in 1783 Taxonomy articles created by Polbot