Iberomesornis
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''Iberomesornis'' ("Iberian intermediate bird") is a monotypic
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of enantiornithine
bird Birds are a group of warm-blooded vertebrates constituting the class (biology), class Aves (), characterised by feathers, toothless beaked jaws, the Oviparity, laying of Eggshell, hard-shelled eggs, a high Metabolism, metabolic rate, a fou ...
of the
Cretaceous The Cretaceous ( ) is a geological period that lasted from about 143.1 to 66 mya (unit), million years ago (Mya). It is the third and final period of the Mesozoic Era (geology), Era, as well as the longest. At around 77.1 million years, it is the ...
of
Spain Spain, or the Kingdom of Spain, is a country in Southern Europe, Southern and Western Europe with territories in North Africa. Featuring the Punta de Tarifa, southernmost point of continental Europe, it is the largest country in Southern Eur ...
.


Discovery

In 1985 the
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...
of ''Iberomesornis'' was discovered by Armando Díaz Romeral in the
Early Cretaceous The Early Cretaceous (geochronology, geochronological name) or the Lower Cretaceous (chronostratigraphy, chronostratigraphic name) is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 143.1 ...
Calizas de La Huérguina Formation at Las Hoyas, Cuenca Province, east central Spain, which dates to the late Barremian, roughly 125 million years ago. The find was first reported in 1988. In 1992 the
type species In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...
''Iberomesornis romerali'' was named and described by José Luis Sanz and José Fernando Bonaparte. The generic name is derived from Iberia and
Ancient Greek Ancient Greek (, ; ) includes the forms of the Greek language used in ancient Greece and the classical antiquity, ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Greek ...
, , "middle", and , , "bird", in reference to the intermediate status between the most basal and the modern birds. The specific name honours Romeral. The holotype specimen, LH-22, part of the Las Hoyas Collection, consists of a compressed articulated partial skeleton of an adult individual lacking the skull, the anterior neck and most of the hands. A second specimen, LH-8200, was referred to a ''Iberomesornis'' sp. in 1994, consisting of the left foot of an individual similar in size to the holotype. After further preparation of the fossil, the species was redescribed by Paul Sereno in 2000.


Biology and ecology

''Iberomesornis'' was quite small, no bigger than a large modern sparrow. On the assumption it had relatively short wings, the wingspan was about twenty centimetres; its weight has been estimated at fifteen to twenty grammes. It bore a single claw on each wing. The preserved axial column length is eighty-seven millimetres. Its ribcage was not strengthened by ossified uncinate processes but cartilaginous processes were likely present. The Las Hoyas Unit 3 site was once a forest surrounding a lake; the climate was warm with a distinct dry season. Since the skull is not known, the diet of ''Iberomesornis'' remains a subject of guesswork. It perhaps hunted insects and other small animals, plucking them out of the air or from the ground. It may have preferred to live near lakes, catching insects from the water's surface. When resting, it used its strong, clawed feet to perch on branches; the toes are long with curved claws and a low and long hallux improved the grasping ability.


Phylogeny

''Iberomesornis romerali'' was more derived physically than '' Archaeopteryx''. The basal characteristic of a long tail had been lost and the coracoid was longer. The vertebrate column of ''Iberomesornis'' ended in a pygostyle: a splint of bone made of fused tail vertebrae that supports the tail feathers. The shorter tail and the higher position of the shoulder indicated by the strut-like coracoid — allowing for a greater wing amplitude — improved manoeuvrability, turning and swooping at speed. Flying speed may have been impaired due to the lack of a bony keel on the sternum as attachment for the flight muscles; it is uncertain whether a true '' foramen triosseum'' was present, through which a tendon could elevate and supinate the wing. In 1992 ''Iberomesornis'' was assigned to the Iberomesornithidae. At that time it was considered to possibly have been very basal, outside of the
clade In biology, a clade (), also known as a Monophyly, monophyletic group or natural group, is a group of organisms that is composed of a common ancestor and all of its descendants. Clades are the fundamental unit of cladistics, a modern approach t ...
Ornithothoraces, as is reflected by its generic name. However, since the discovery of ''Iberomesornis'', many more bird fossils have come to light in the quarries of Liaoning province, China. Well preserved specimens of Chinese species such as '' Sinornis'' and '' Confuciusornis'' have permitted scientists to better understand the birds' early fossil history. Today, ''Iberomesornis'' is seen as positioned within Ornithothoraces, as a basal member of the Enantiornithes.


References


Sources

* {{Taxonbar, from=Q134376 Enantiornithes Monotypic prehistoric bird genera Cretaceous birds of Europe Cretaceous Spain Fossils of Spain La Huérguina Formation Fossil taxa described in 1992 Taxa named by José Bonaparte