The hymenium is the tissue layer on the

hymenophore A hymenophore refers to the hymenium-bearing structure of a fungal fruiting body. Hymenophores can be smooth surfaces, lamellae, folds, tubes, or teeth. The term was coined by Robert Hooke Robert Hooke (; 18 July 16353 March 1703) was an ...

of a

fungal A fungus (: fungi , , , or ; or funguses) is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and molds, as well as the more familiar mushrooms. These organisms are classified as one of the tradit ...

fruiting body The sporocarp (also known as fruiting body, fruit body or fruitbody) of fungi is a multicellular structure on which spore-producing structures, such as basidia or asci, are borne. The fruitbody is part of the sexual phase of a fungal life cyc ...

where the cells develop into

basidia A basidium (: basidia) is a microscopic spore-producing structure found on the hymenophore of reproductive bodies of basidiomycete fungi. The presence of basidia is one of the main characteristic features of the group. These bodies are also ...

or asci, which produce

spores In biology, a spore is a unit of sexual (in fungi) or asexual reproduction that may be adapted for dispersal and for survival, often for extended periods of time, in unfavourable conditions. Spores form part of the life cycles of many plant ...

. In some species all of the cells of the hymenium develop into basidia or asci, while in others some cells develop into sterile cells called

cystidia A cystidium (: cystidia) is a relatively large cell found on the sporocarp of a basidiomycete (for example, on the surface of a mushroom gill), often between clusters of basidia. Since cystidia have highly varied and distinct shapes that are o ...

(

basidiomycetes Basidiomycota () is one of two large division (mycology), divisions that, together with the Ascomycota, constitute the subkingdom Dikarya (often referred to as the "higher fungi") within the kingdom Fungi. Members are known as basidiomycetes. Mor ...

) or

paraphyses Paraphyses are erect sterile filament-like support structures occurring among the reproductive apparatuses of fungi, ferns, bryophytes and some thallophytes. The singular form of the word is paraphysis. In certain fungi, they are part of the f ...

(

ascomycetes Ascomycota is a phylum of the kingdom Fungi that, together with the Basidiomycota, forms the subkingdom Dikarya. Its members are commonly known as the sac fungi or ascomycetes. It is the largest phylum of Fungi, with over 64,000 species. The de ...

). Cystidia are often important for microscopic identification. The subhymenium consists of the supportive

hyphae A hypha (; ) is a long, branching, filamentous structure of a fungus, oomycete, or actinobacterium. In most fungi, hyphae are the main mode of vegetative growth, and are collectively called a mycelium. Structure A hypha consists of one o ...

from which the cells of the hymenium grow, beneath which is the hymenophoral trama, the hyphae that make up the mass of the hymenophore. The position of the hymenium is traditionally the first characteristic used in the classification and identification of mushrooms. Below are some examples of the diverse types which exist among the macroscopic

Basidiomycota Basidiomycota () is one of two large divisions that, together with the Ascomycota, constitute the subkingdom Dikarya (often referred to as the "higher fungi") within the kingdom Fungi. Members are known as basidiomycetes. More specifically, Basi ...

and

Ascomycota Ascomycota is a phylum of the kingdom Fungi that, together with the Basidiomycota, forms the subkingdom Dikarya. Its members are commonly known as the sac fungi or ascomycetes. It is the largest phylum of Fungi, with over 64,000 species. The def ...

. * In

agaric An agaric () is a type of fungal fruiting body characterized by the presence of a pileus (cap) that is clearly differentiated from the stipe (stalk), with lamellae (gills) on the underside of the pileus. It is a type of mushroom (or toadstool) ...

s, the hymenium is on the vertical faces of the

gill A gill () is a respiration organ, respiratory organ that many aquatic ecosystem, aquatic organisms use to extract dissolved oxygen from water and to excrete carbon dioxide. The gills of some species, such as hermit crabs, have adapted to allow r ...

s. * In

bolete A bolete is a type of mushroom, or fungal fruiting body. It can be identified thanks to a unique cap. On the underside of the cap there is usually a spongy surface with pores, instead of the gills typical of mushrooms. A similar pore surface i ...

s and

polypore Polypores, also called bracket or shelf fungi, are a morphological group of basidiomycete-like gilled mushrooms and hydnoid fungi that form large fruiting bodies called conks, which are typically woody, circular, shelf- or bracket-shaped, ...

s, it is in a spongy mass of downward-pointing tubes. * In

puffball Puffballs are a type of fungus featuring a ball-shaped fruit body that (when mature) bursts on contact or impact, releasing a cloud of dust-like spores into the surrounding area. Puffballs belong to the division Basidiomycota and encompass sever ...

s, it is internal. * In

stinkhorn Phallaceae is a family of fungi, commonly known as stinkhorns, within the order Phallales. Stinkhorns have a worldwide distribution, but are especially prevalent in tropical regions. They are known for their foul-smelling, sticky spore masses ...

s, it develops internally and then is exposed in the form of a foul-smelling gel. * In

cup fungi The Pezizaceae (commonly referred to as cup fungi) are a family of fungi in the Ascomycota which produce mushrooms that tend to grow in the shape of a "cup". Spores are formed on the inner surface of the fruit body ( ascoma). The cup shape typi ...

, it is on the concave surface of the cup. * In teeth fungi, it grows on the outside of tooth-like spines.

Gallery

Image:Lamellen.jpg, Gills of the fly agaric ''

Amanita muscaria ''Amanita muscaria'', commonly known as the fly agaric or fly amanita, is a basidiomycete fungus of the genus ''Amanita''. It is a large white-lamella (mycology), gilled, white-spotted mushroom typically featuring a bright red cap covered with ...

'' Image:Suillus luteus.JPG, The

slippery jack (''

Suillus luteus ''Suillus luteus'' is a bolete fungus, and the type species of the genus ''Suillus''. Commonly referred to as slippery jack or sticky bun in English-speaking countries, its names refer to the brown pileus (mycology), cap, which is characterist ...

'') with bright yellow pores Image:Riesenbovist.jpg, Giant

Calvatia gigantea ''Calvatia gigantea'', commonly known in English as the giant puffball, is a puffball mushroom commonly found in meadows, fields, and deciduous forests in late summer and autumn. It is found in temperate areas throughout the world. Taxonomy T ...

'' Image:Phallus impudicus7 Stinkhorn.jpg, A

, ''

Phallus impudicus ''Phallus impudicus'', known colloquially as the common stinkhorn, is a widespread fungus in the Phallaceae (stinkhorn) family. It is recognizable for its foul odor and its phallic shape when mature, the latter feature giving rise to several ...

'' Image:Scarlet elf cap cadnant dingle.jpg, The cup fungus ''

Sarcoscypha austriaca ''Sarcoscypha austriaca'' is a saprobic fungus in the family Sarcoscyphaceae of the order Pezizales of Ascomycota. It is commonly known as the scarlet elfcup, pézize écarlate and scharlachroter kelchbecherling.Polyporus alveolaris'', the hexagonal-pored polypore

References

*Régis Courtecuisse, Bernard Duhem : ''Guide des champignons de France et d'Europe'' (Delachaux & Niestlé, 1994–2000).

External links

IMA Mycological Glossary: Hymenium

Hymenium of an ascomycete, ''Monilinia fructicola''
Jack Murphy Mycological Images: Hymenium
Hymenium of a basidiomycete, ''Russula laurocerasi'' Fungal morphology and anatomy {{Fungi-stub