The hylocitrea (''Hylocitrea bonensis''), also known as the yellow-flanked whistler or olive-flanked whistler, is a species of

bird Birds are a group of warm-blooded vertebrates constituting the class (biology), class Aves (), characterised by feathers, toothless beaked jaws, the Oviparity, laying of Eggshell, hard-shelled eggs, a high Metabolism, metabolic rate, a fou ...

that is

endemic Endemism is the state of a species being found only in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also foun ...

to montane forests on the

Indonesia Indonesia, officially the Republic of Indonesia, is a country in Southeast Asia and Oceania, between the Indian Ocean, Indian and Pacific Ocean, Pacific oceans. Comprising over List of islands of Indonesia, 17,000 islands, including Sumatra, ...

n island of

Sulawesi Sulawesi ( ), also known as Celebes ( ), is an island in Indonesia. One of the four Greater Sunda Islands, and the List of islands by area, world's 11th-largest island, it is situated east of Borneo, west of the Maluku Islands, and south of Min ...

.Boles, W. E. (2007). Yellow-flanked Whistler (Hylocitrea bonensis). pp. 411 in: Del Hoyo, J.; Elliot, A., & Christie D. eds. (2007). ''

Handbook of the Birds of the World The ''Handbook of the Birds of the World'' (HBW) is a multi-volume series produced by the Spanish publishing house Lynx Edicions in partnership with BirdLife International. It is the first handbook to cover every known living species of bird. ...

. Vol. 12. Picathartes to Tits and Chickadees.''

Lynx Edicions Lynx Nature Books, based in Barcelona, is a publishing company specializing in ornithology and natural history. The company was founded in 1989. It was formerly named Lynx Edicions. History Lynx Nature Books was founded as Lynx Edicions in Barce ...

, Barcelona. It is

monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unisp ...

within the genus ''Hylocitrea'', and has traditionally been considered a member of the family

Pachycephalidae The Pachycephalidae are a family of bird species that includes the whistlers, shrikethrushes, and three of the pitohuis, and is part of the ancient Australo-Papuan radiation of songbirds. The family includes 69 species that are separated into fi ...

, but recent genetic evidence suggests it should be placed in a monotypic subfamily of the family

Bombycillidae The waxwings are three species of passerine birds classified in the genus ''Bombycilla''. They are pinkish-brown and pale grey with distinctive smooth plumage in which many body feathers are not individually visible, a black and white eyestripe ...

, or even its own family, Hylocitreidae. A 2019 study found it to be a

sister group In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an evolutionary tree. Definition The expression is most easily illustrated by a cladogram: Taxon A and ...

to a clade containing the hypocolius (Hypocoliidae) and the extinct Hawaiian honeyeaters (Mohoidae), with the clade containing all three being a sister group to the silky-flycatchers (Ptiliogonatidae). The divergences forming these families occurred in the early

Miocene The Miocene ( ) is the first epoch (geology), geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and mea ...

, about 20-23 million years ago.

References

{{Taxonbar, from1=Q10770215, from2=Q1585397, from3=Q10770216 Bombycilloidea Birds described in 1894 Endemic birds of Sulawesi Extant Miocene first appearances Taxonomy articles created by Polbot