Hugh Iltis
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Hugh Iltis (April 7, 1925 – December 19, 2016) was a professor of
botany Botany, also called plant science, is the branch of natural science and biology studying plants, especially Plant anatomy, their anatomy, Plant taxonomy, taxonomy, and Plant ecology, ecology. A botanist or plant scientist is a scientist who s ...
and director of the
herbarium A herbarium (plural: herbaria) is a collection of preserved plant biological specimen, specimens and associated data used for scientific study. The specimens may be whole plants or plant parts; these will usually be in dried form mounted on a sh ...
at the
University of Wisconsin–Madison The University of Wisconsin–Madison (University of Wisconsin, Wisconsin, UW, UW–Madison, or simply Madison) is a public land-grant research university in Madison, Wisconsin, United States. It was founded in 1848 when Wisconsin achieved st ...
. While he is most noted as a scientist for his role in the discovery of perennial
teosinte ''Zea'' is a genus of flowering plants in the Poaceae, grass family. The best-known species is ''Z. mays'' (variously called maize, corn, or Indian corn), one of the most important crops for human societies throughout much of the world. The four ...
(''
Zea diploperennis ''Zea diploperennis'', the diploperennial teosinte, is a species of grass (family: Poaceae) in the genus '' Zea'' and a teosinte (wild relative of maize or corn). It is perennial. Conservation Virtually all populations of this teosinte are eithe ...
''), a wild
diploid Ploidy () is the number of complete sets of chromosomes in a cell, and hence the number of possible alleles for autosomal and pseudoautosomal genes. Here ''sets of chromosomes'' refers to the number of maternal and paternal chromosome copies, ...
relative of modern maize (''
Zea mays Maize (; ''Zea mays''), also known as corn in North American English, is a tall stout Poaceae, grass that produces cereal grain. It was domesticated by indigenous peoples of Mexico, indigenous peoples in southern Mexico about 9,000 years ago ...
''), he is also remembered as an outspoken
environmental Environment most often refers to: __NOTOC__ * Natural environment, referring respectively to all living and non-living things occurring naturally and the physical and biological factors along with their chemical interactions that affect an organism ...
conservationist.


Life and work

He was born Hugo Hellmut Iltis to Anni (née Liebscher) and Hugo Iltis, a botanist and geneticist who was a life sciences teacher at the German-language
secondary school A secondary school, high school, or senior school, is an institution that provides secondary education. Some secondary schools provide both ''lower secondary education'' (ages 11 to 14) and ''upper secondary education'' (ages 14 to 18), i.e., b ...
of Brünn (Brno). His father was also the first biographer of
Gregor Mendel Gregor Johann Mendel Order of Saint Augustine, OSA (; ; ; 20 July 1822 – 6 January 1884) was an Austrian Empire, Austrian biologist, meteorologist, mathematician, Augustinians, Augustinian friar and abbot of St Thomas's Abbey, Brno, St. Thom ...
and a vocal opponent of Nazi "racial science". In the fall of 1938, the Iltis family was granted visas to enter the United States thanks to the intercession of the
Emergency Committee in Aid of Displaced Foreign Scholars The Emergency Committee in Aid of Displaced Foreign Scholars (1933–1945) assisted scholars who were barred from teaching, persecuted and threatened with imprisonment by the Nazis. The program began in Germany soon after Hitler took power and expa ...
, along with affidavits of endorsement from
Albert Einstein Albert Einstein (14 March 187918 April 1955) was a German-born theoretical physicist who is best known for developing the theory of relativity. Einstein also made important contributions to quantum mechanics. His mass–energy equivalence f ...
and
Franz Boas Franz Uri Boas (July 9, 1858 – December 21, 1942) was a German-American anthropologist and ethnomusicologist. He was a pioneer of modern anthropology who has been called the "Father of American Anthropology". His work is associated with the mov ...
. In January 1939, when
Hitler Adolf Hitler (20 April 1889 – 30 April 1945) was an Austrian-born German politician who was the dictator of Nazi Germany from 1933 until Death of Adolf Hitler, his suicide in 1945. Adolf Hitler's rise to power, He rose to power as the lea ...
's military was preparing the
invasion of Czechoslovakia On 20–21 August 1968, the Czechoslovak Socialist Republic was jointly invaded by four fellow Warsaw Pact countries: the Soviet Union, the Polish People's Republic, the People's Republic of Bulgaria, and the Hungarian People's Republic. The in ...
, thirteen-year-old Hugo escaped with his mother and his older brother Wilfred on a harrowing train ride that traversed
Nazi Germany Nazi Germany, officially known as the German Reich and later the Greater German Reich, was the German Reich, German state between 1933 and 1945, when Adolf Hitler and the Nazi Party controlled the country, transforming it into a Totalit ...
to France. He recalled that during a midnight stop at the
Stuttgart Stuttgart (; ; Swabian German, Swabian: ; Alemannic German, Alemannic: ; Italian language, Italian: ; ) is the capital city, capital and List of cities in Baden-Württemberg by population, largest city of the States of Germany, German state of ...
station,
Gestapo The (, ), Syllabic abbreviation, abbreviated Gestapo (), was the official secret police of Nazi Germany and in German-occupied Europe. The force was created by Hermann Göring in 1933 by combining the various political police agencies of F ...
officers combed the train, removing ten passengers; the Iltises survived because the boys pretended to be asleep while their mother bluffed that she was the wife of a French diplomat. In
Cherbourg Cherbourg is a former Communes of France, commune and Subprefectures in France, subprefecture located at the northern end of the Cotentin peninsula in the northwestern French departments of France, department of Manche. It was merged into the com ...
, they were joined by Hugo Iltis and boarded the passenger ship
RMS Aquitania RMS ''Aquitania'' was an ocean liner of the Cunard Line in service from 1914 to 1950. She was designed by Leonard Peskett and built by John Brown & Company in Clydebank, Scotland. She was launched on 21 April 1913 and sailed on her maiden vo ...
for the Atlantic crossing. They settled in
Fredericksburg, Virginia Fredericksburg is an Independent city (United States), independent city in Virginia, United States. As of the 2020 United States census, 2020 census, the population was 27,982. It is south of Washington, D.C., and north of Richmond, Virginia, R ...
, where the senior Hugo Iltis was soon appointed to a professorship in biology at
Mary Washington College University of Mary Washington (UMW) is a public liberal arts university in Fredericksburg, Virginia. Established in 1908 as the State Normal and Industrial School for Women at Fredericksburg, the institution later became known as Fredericksbu ...
and the younger Hugo Americanized his name to Hugh Iltis. Iltis' undergraduate enrollment at the
University of Tennessee The University of Tennessee, Knoxville (or The University of Tennessee; UT; UT Knoxville; or colloquially UTK or Tennessee) is a Public university, public Land-grant university, land-grant research university in Knoxville, Tennessee, United St ...
was interrupted by service in the U.S. Army from 1944 to 1946, initially as a medic. Because of his native proficiency in German, he was transferred to an intelligence unit. After
World War II World War II or the Second World War (1 September 1939 – 2 September 1945) was a World war, global conflict between two coalitions: the Allies of World War II, Allies and the Axis powers. World War II by country, Nearly all of the wo ...
, Iltis was stationed in Germany, where he participated in the interrogation of captive
Wehrmacht The ''Wehrmacht'' (, ) were the unified armed forces of Nazi Germany from 1935 to 1945. It consisted of the German Army (1935–1945), ''Heer'' (army), the ''Kriegsmarine'' (navy) and the ''Luftwaffe'' (air force). The designation "''Wehrmac ...
and SS officers, including
Heinrich Himmler Heinrich Luitpold Himmler (; 7 October 1900 – 23 May 1945) was a German Nazism, Nazi politician and military leader who was the 4th of the (Protection Squadron; SS), a leading member of the Nazi Party, and one of the most powerful p ...
, and processed documents to prosecute
Nazi war crimes The governments of the German Empire and Nazi Germany (under Adolf Hitler) ordered, organized, and condoned a substantial number of war crimes, first in the Herero and Nama genocide and then in the First and Second World Wars. The most notable of ...
at
Nuremberg Nuremberg (, ; ; in the local East Franconian dialect: ''Nämberch'' ) is the Franconia#Towns and cities, largest city in Franconia, the List of cities in Bavaria by population, second-largest city in the States of Germany, German state of Bav ...
. Iltis returned to the University of Tennessee, where he studied botany under Aaron J. Sharp. He carried out graduate studies at
Washington University in St. Louis Washington University in St. Louis (WashU) is a private research university in St. Louis, Missouri, United States. Founded in 1853 by a group of civic leaders and named for George Washington, the university spans 355 acres across its Danforth ...
, where he received his Ph.D. in 1952 under the direction of
Edgar Anderson Edgar Shannon Anderson (November 9, 1897 – June 18, 1969) was an American botanist. He introduced the term '' introgressive hybridization'' and his 1949 book of that title was an original and important contribution to botanical genetics. His w ...
. He was primarily trained in plant systematics and taxonomy, with a focus on the caper family (''
Capparaceae The Capparaceae (or Capparidaceae), commonly known as the caper family, are a family of plants in the order Brassicales. As currently circumscribed, the family contains 15 genera and about 430 species. The largest genera are ''Capparis'' (about 1 ...
'') and the spider-flower family (''
Cleomaceae The Cleomaceae are a small family of flowering plants in the order Brassicales, comprising about 220 species in two genera, '' Cleome'' and '' Cleomella''. These genera were previously included in the family Capparaceae, but were raised to a dis ...
''). His first academic appointment was at the
University of Arkansas The University of Arkansas (U of A, UArk, or UA) is a Public university, public Land-grant university, land-grant research university in Fayetteville, Arkansas, United States. It is the Flagship campus, flagship campus of the University of Arkan ...
from 1952 to 1955, and here he completed a study of the ''Capparaceae'' of
Nevada Nevada ( ; ) is a landlocked state in the Western United States. It borders Oregon to the northwest, Idaho to the northeast, California to the west, Arizona to the southeast, and Utah to the east. Nevada is the seventh-most extensive, th ...
. Later works formed a series, "Studies in the ''Capparaceae''", which includes 24 publications, including newly described species and genera. An associated series of papers describes his research in the ''Cleomaceae''. (At the time when he did his studies, the ''Cleomaceae'' was included in the ''Capparaceae''.) In 1955, Iltis relocated to the Botany Department of the University of Wisconsin–Madison, where in addition to directing the herbarium he regularly taught plant geography, taxonomy, and grass systematics. He arranged to purchase much of the
Catholic University of America The Catholic University of America (CUA) is a private Catholic research university in Washington, D.C., United States. It is one of two pontifical universities of the Catholic Church in the United States – the only one that is not primarily a ...
herbarium when it was deaccessioned. By the time of his retirement in 1993, he had directed 37 candidates pursuing graduate degrees, and he and his students had collected thousands of specimens throughout the Upper Midwest to document distributions of plant species, leading to the publication of the ''Atlas of the Wisconsin Prairie and Savanna Flora'' (2000) coauthored with Herbarium Curator Theodore Cochrane. Anecdotes abounded concerning his colorful, often imperious manner. One colleague poked fun at Iltis by taping on his office door a cartoon that showed a boss dictating to a secretary and concluding, "Type that up, make ten thousand copies, and send them to all the important people in the world

At the end of a public lecture, when an audience member asked flippantly, "What good is nature?" Iltis shot back, "What good are you?" Yet students flocked to his course on "Man's Need for Nature", and he was generous with his knowledge and his counsel. He cultivated strong ties with Latin American botanists, often hosting them for extended stays at his home located within the
University of Wisconsin–Madison Arboretum The University of Wisconsin–Madison Arboretum is a teaching and research facility of the University of Wisconsin–Madison and the site of historic research in ecological restoration. In addition to its in Madison, Wisconsin (located about fo ...
. An avid
naturalist Natural history is a domain of inquiry involving organisms, including animals, fungi, and plants, in their natural environment, leaning more towards observational than experimental methods of study. A person who studies natural history is cal ...
, Iltis conducted numerous expeditions to
Mexico Mexico, officially the United Mexican States, is a country in North America. It is the northernmost country in Latin America, and borders the United States to the north, and Guatemala and Belize to the southeast; while having maritime boundar ...
and Central and South America to search for new discoveries. High in the
Altiplano The Altiplano (Spanish language, Spanish for "high plain"), Collao (Quechuan languages, Quechua and Aymara language, Aymara: Qullaw, meaning "place of the Qulla people, Qulla") or Andean Plateau, in west-central South America, is the most extens ...
of southern
Peru Peru, officially the Republic of Peru, is a country in western South America. It is bordered in the north by Ecuador and Colombia, in the east by Brazil, in the southeast by Bolivia, in the south by Chile, and in the south and west by the Pac ...
in 1962, he noticed a tiny flower that had not yet been classified by taxonomists. A full fifty years later, he and Harvey Ballard finally named it '' Viola lilliputana'', and it was selected as one of the top ten new species of the year by the
International Institute for Species Exploration The International Institute for Species Exploration (IISE) was a research institute located in Syracuse, New York. Its mission was to improve Taxonomy (biology), taxonomical exploration and the cataloging of new species of flora and fauna. Betw ...
. Iltis' work was of economic importance, because he identified new sources of genetic variability that have been used by
horticultural Horticulture (from ) is the art and science of growing fruits, vegetables, flowers, trees, shrubs and ornamental plants. Horticulture is commonly associated with the more professional and technical aspects of plant cultivation on a smaller and mo ...
breeders. On the same 1962 expedition to Peru, he spotted a wild
tomato The tomato (, ), ''Solanum lycopersicum'', is a plant whose fruit is an edible Berry (botany), berry that is eaten as a vegetable. The tomato is a member of the nightshade family that includes tobacco, potato, and chili peppers. It originate ...
that he recorded as No. 832. He collected specimens for several herbaria, and sent samples and seeds to various specialists in the field. This plant turned out to be a new species of tomato with much higher sugar and solids content than domestically grown tomatoes. As a source for hybridization with domestic tomatoes, it has been used both to improve the flavor of tomatoes and to boost solids content. Iltis used taxonomic and morphological approaches to investigate the domestication of corn, tracing the changes that transformed an unpromising wild grass into one of the most important food crops. His work supported the view that domestic corn was derived from a species of
teosinte ''Zea'' is a genus of flowering plants in the Poaceae, grass family. The best-known species is ''Z. mays'' (variously called maize, corn, or Indian corn), one of the most important crops for human societies throughout much of the world. The four ...
, a group of grasses that grows wild in many areas of Mexico. It was generally believed that the original wild corn was extinct in the wild. Iltis used a hypothetical illustration of this plant for a New Year's greeting card that he sent to family and friends in 1976. This drawing prompted a Mexican colleague, Luz María Villarreal de Puga (1913–2013), to launch an intensive search for just such a plant, and one of her students, Rafael Guzmán, found it (or so he thought) growing in the wild. In 1978, Iltis led a team of botanists to the site and determined that it was in fact a heretofore unknown species of teosinte, ''Zea diploperennis'', which is valued for its resistance to certain viruses. Iltis warned that the practice of collecting plants in tropical countries without involving local botanists and without depositing duplicate specimens in local herbaria would eventually cause trouble. And indeed, in recent years Brazil and some Andean countries have enacted laws that severely restrict field studies. Iltis was an outspoken
environmentalist Environmentalism is a broad Philosophy of life, philosophy, ideology, and social movement about supporting life, habitats, and surroundings. While environmentalism focuses more on the environmental and nature-related aspects of Green politics, g ...
and conservationist, championing the preservation of threatened habitats to protect
biodiversity Biodiversity is the variability of life, life on Earth. It can be measured on various levels. There is for example genetic variability, species diversity, ecosystem diversity and Phylogenetics, phylogenetic diversity. Diversity is not distribut ...
. Some species of teosinte are critically endangered, and all face a growing threat as agricultural land use expands in Mexico. He campaigned with colleagues at the
University of Guadalajara The University of Guadalajara () is a public university, public research university located in Guadalajara, Mexico. It was originally established in 1586 and officially founded on 12 February 1791 as the Royal and Pontifical University of Gu ...
to protect the natural environment of ''Zea diploperennis'' by creating the 345,000-acre
Sierra de Manantlán Biosphere Reserve The Sierra de Manantlán Biosphere Reserve () (established 1988) is a UNESCO Biosphere Reserve in the states of Colima and Jalisco, Mexico. The reserve is located in the transition of the Nearctic and Neotropical realms and encompasses parts of ...
. He cofounded the Wisconsin chapter of the
Nature Conservancy The Nature Conservancy (TNC) is a global environmental organization headquartered in Arlington, Virginia, United States. it works via affiliates or branches in 79 countries and territories, as well as across every state in the US. Founded in ...
in 1960 and helped establish Hawaii's Natural Areas Law of 1970. He was a leader in the campaign to ban
DDT Dichlorodiphenyltrichloroethane, commonly known as DDT, is a colorless, tasteless, and almost odorless crystalline chemical compound, an organochloride. Originally developed as an insecticide, it became infamous for its environmental impacts. ...
in Wisconsin, which in 1968 was the first U.S. state to do so. He also called for a moratorium on cutting virgin timber in the state. In a 1970 article, "Man First? Man Last? The Paradox of Human Ecology", he wrote: "If we are to usher in an Age of Ecologic Reason, we must accept the certainty of a radical economic and political restructuring as well as ethical and cultural restructuring of society. No more expanding populations.... We must stop and limit ourselves now." Iltis fathered four sons, Frank and Michael by his first wife, Grace Schaffel, and David and John by his second wife,
Carolyn Merchant Carolyn Merchant (born July 12, 1936 in Rochester, New York) is an American ecofeminist philosopher and historian of science most famous for her theory (and book of the same title) on '' The Death of Nature'', whereby she identifies the Scien ...
. He and his third wife, Sharyn Wisniewski (1950–2013), endowed a fund at the University of Wisconsin–Madison Department of Botany to support graduate student fieldwork in plant systematics. He remained active up to his death in Madison at age 91 from complications of
vascular disease Vascular disease is a class of diseases of the vessels of the circulatory system in the human body, body, including blood vessels – the arteries and veins, and the lymphatic vessels. Vascular disease is a subgroup of cardiovascular disease. Diso ...
. His papers are preserved in the archives of the University of Wisconsin–Madison.


Honors

Fellow botanists have honored Iltis by naming two genera and 19 species of plants after him. He received the
Asa Gray Award Asa Gray (November 18, 1810 – January 30, 1888) is considered the most important American botanist of the 19th century. His '' Darwiniana'' (1876) was considered an important explanation of how religion and science were not necessarily mutua ...
of the
American Society of Plant Taxonomists The American Society of Plant Taxonomists (ASPT) is a botany, botanical organization formed in 1935 to "foster, encourage, and promote education and research in the field of plant taxonomy, to include those areas and fields of study that contribut ...
(1994) and a Merit Award of the
Botanical Society of America The Botanical Society of America (BSA) represents professional and amateur botanists, researchers, educators and students in over 80 countries of the world. It functions as a United States nonprofit 501(c)(3) membership society. History The soc ...
(1996). Internationally, he received a Contribución Distinguida award from the president of Mexico for his role in establishing the Sierra de Manantlán Biosphere Reserve (1987), the Luz María Villarreal de Puga Medal from the University of Guadalajara (1994), and an honorary doctorate from the same university (2007). He received the Sol Feinstone Environmental Award conferred by the
State University of New York College of Environmental Science and Forestry State most commonly refers to: * State (polity), a centralized political organization that regulates law and society within a territory **Sovereign state, a sovereign polity in international law, commonly referred to as a country **Nation state, a ...
(1990), the
National Wildlife Federation The National Wildlife Federation (NWF) is the largest private, nonprofit conservation education and advocacy organization in the United States, with over six million members and supporters, and 51 state and territorial affiliated organizations (i ...
's Merit Award (1992), and the
Society for Conservation Biology The Society for Conservation Biology (SCB) is an 501(c)(3) non-profit international professional organization that is dedicated to conserving biodiversity. There are over 4,000 members worldwide, including students and those in related non-academ ...
Service Award (1994). On
Earth Day Earth Day is an annual event on April 22 to demonstrate support for environmental protection. First held on April 22, 1970, it now includes a wide range of events coordinated globally through earthday.org (formerly Earth Day Network) includin ...
2017, he was posthumously inducted into the Wisconsin Conservation Hall of Fame.


Selected publications

* Iltis, Hugh H. (1947). "A Visit to Gregor Mendel's Home". ''Journal of Heredity'', vol. 38. no. 6, pp. 163–166. https://doi.org/10.1093/oxfordjournals.jhered.a105719 * Iltis, Hugh H. (1950). "Studies in Virginia Plants. I. List of Bryophytes from the Vicinity of Fredericksburg, Virginia." ''Castanea. The Journal of the Southern Appalachian Botanical Club'', vol. 15, pp. 38–50. * Iltis, Hugh H. (1955). Capparidaceae' of Nevada''. Beltsville: National Arboretum and Section of Plant Introduction, Horticultural Corp Research Branch, Plant Industry Station. 24 pp. * Iltis, Hugh H. (1973)
"Can One Love a Plastic Tree?"
''Bulletin of the Ecological Society of America'', 54(4), 5–7,19. * Iltis, Hugh H., Doebley, John F., Guzmán, Rafael, & Pazy, Batia (1979). "''Zea diploperennis'' (Gramineae): A New Teosinte from Mexico". ''Science'', 203(4376), 186–188. * Iltis, Hugh H., & Doebley, John F. (1980). "Taxonomy of ''Zea'' (Gramineae). II. Subspecific Categories in the ''Zea mays'' Complex and a Generic Synopsis". ''American Journal of Botany'', 994–1004. * https://repository.arizona.edu/bitstream/handle/10150/550937/dp_03_04-175-192.pdf;jsessionid=15E022575DB1DF6E55009B91F11F29B3?sequence=1 * * Iltis, Hugh H. (2000). "Homeotic Sexual Translocations and the Origin of Maize (''Zea mays'', Poaceae): A New Look at an Old Problem". ''Economic Botany'', 54(1), 7–42. * Iltis, Hugh H., & Benz, Bruce F. (2000). "''Zea nicaraguensis'' (Poaceae), a New Teosinte from Pacific Coastal Nicaragua". ''Novon'', 382–390. * Cochrane, Theodore S., & Iltis, Hugh H. (2000). ''Atlas of the Wisconsin Prairie and Savanna Flora.'' Madison: Wisconsin Department of Natural Resources. 226 pp
Online
* Ballard, Harvey E., and Iltis, Hugh H. (2012). "''Viola lilliputana sp. Nov.'' (Viola sect. Andinium, Violaceae), One of the World’s Smallest Violets, from the Andes of Peru". ''Brittonia'', 64(4), 353-358.


References


External links


Hugh H. Iltis papers
UW Archives
Curriculum vitae 2017


of maize and teosinte
Photos
of Hugh Iltis {{DEFAULTSORT:Iltis, Hugh 1925 births 2016 deaths 20th-century American botanists 21st-century American botanists American conservationists American geneticists Botanists active in North America Botanists active in Central America Czech botanists Czech Jews Czech refugees Jewish American scientists Jewish emigrants from Nazi Germany to the United States Scientists from Brno Naturalized citizens of the United States Scientists from Madison, Wisconsin Scientists from Virginia University of Arkansas faculty University of Tennessee alumni University of Wisconsin–Madison faculty Washington University in St. Louis alumni