Huchen
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The huchen (''Hucho hucho'') (, from German), also known as Danube salmon or redfish (), is a large species of freshwater fish in the
family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
Salmonidae Salmonidae (, ) is a family (biology), family of ray-finned fish, the only extant member of the suborder Salmonoidei, consisting of 11 extant genera and over 200 species collectively known as "salmonids" or "salmonoids". The family includes salmo ...
native to the Danube basin in Central and
Eastern Europe Eastern Europe is a subregion of the Europe, European continent. As a largely ambiguous term, it has a wide range of geopolitical, geographical, ethnic, cultural and socio-economic connotations. Its eastern boundary is marked by the Ural Mountain ...
. It is the
type species In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...
of
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
'' Hucho'' (a.k.a. the taimens), being closely related (in the same
subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end botanical subfamily names with "-oideae", and zo ...
) to
salmon Salmon (; : salmon) are any of several list of commercially important fish species, commercially important species of euryhaline ray-finned fish from the genera ''Salmo'' and ''Oncorhynchus'' of the family (biology), family Salmonidae, native ...
, trout, char and lenoks.


Distribution

The huchen is
endemic Endemism is the state of a species being found only in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also foun ...
to the
Danube The Danube ( ; see also #Names and etymology, other names) is the List of rivers of Europe#Longest rivers, second-longest river in Europe, after the Volga in Russia. It flows through Central and Southeastern Europe, from the Black Forest sou ...
basin in Europe where the remaining population is threatened primarily by river damming, resulting in
habitat fragmentation Habitat fragmentation describes the emergence of discontinuities (fragmentation) in an organism's preferred environment (habitat), causing population fragmentation and ecosystem decay. Causes of habitat fragmentation include geological proces ...
and loss through river impoundment and disruption of the longitudinal continuity of rivers, cutting away fish from its spawning grounds, with
overfishing Overfishing is the removal of a species of fish (i.e. fishing) from a body of water at a rate greater than that the species can replenish its population naturally (i.e. the overexploitation of the fishery's existing Fish stocks, fish stock), resu ...
and fisheries mismanagement as an additional issue in many areas. Damming and all these other problems are especially visible in the
Balkans The Balkans ( , ), corresponding partially with the Balkan Peninsula, is a geographical area in southeastern Europe with various geographical and historical definitions. The region takes its name from the Balkan Mountains that stretch throug ...
. The upper reaches of the Danube basin, rivers and tributaries contain almost all of the recent population. This includes: *In
Austria Austria, formally the Republic of Austria, is a landlocked country in Central Europe, lying in the Eastern Alps. It is a federation of nine Federal states of Austria, states, of which the capital Vienna is the List of largest cities in Aust ...
: the Inn river, the upper
Drava The Drava or Drave (, ; ; ; ; ), historically known as the Dravis or Dravus, is a river in southern Central Europe.
, with the lower Gail in
Carinthia Carinthia ( ; ; ) is the southernmost and least densely populated States of Austria, Austrian state, in the Eastern Alps, and is noted for its mountains and lakes. The Lake Wolayer is a mountain lake on the Carinthian side of the Carnic Main ...
, the Pielach, in
Styria Styria ( ; ; ; ) is an Austrian Federal states of Austria, state in the southeast of the country. With an area of approximately , Styria is Austria's second largest state, after Lower Austria. It is bordered to the south by Slovenia, and cloc ...
the upper Mur. *In
Germany Germany, officially the Federal Republic of Germany, is a country in Central Europe. It lies between the Baltic Sea and the North Sea to the north and the Alps to the south. Its sixteen States of Germany, constituent states have a total popu ...
: middle and lower Inn,
Isar The Isar () is a river in Austria and in Bavaria, Germany. Its source is in the Karwendel mountain range of the Alps. The Isar river enters Germany near Mittenwald and flows through Krün, Wallgau, Bad Tölz, Munich, and Landshut before reaching ...
(especially in Munich) In the
Balkans The Balkans ( , ), corresponding partially with the Balkan Peninsula, is a geographical area in southeastern Europe with various geographical and historical definitions. The region takes its name from the Balkan Mountains that stretch throug ...
huchen still appears in following river systems: *In
Slovenia Slovenia, officially the Republic of Slovenia, is a country in Central Europe. It borders Italy to the west, Austria to the north, Hungary to the northeast, Croatia to the south and southeast, and a short (46.6 km) coastline within the Adriati ...
: in the headwaters of
Sava The Sava, is a river in Central Europe, Central and Southeast Europe, a right-bank and the longest tributary of the Danube. From its source in Slovenia it flows through Croatia and along its border with Bosnia and Herzegovina, and finally reac ...
and its tributaries. *In
Croatia Croatia, officially the Republic of Croatia, is a country in Central Europe, Central and Southeast Europe, on the coast of the Adriatic Sea. It borders Slovenia to the northwest, Hungary to the northeast, Serbia to the east, Bosnia and Herze ...
: in the upper and middle course of
Kupa The Kupa () or Kolpa ( or ; from in Roman Empire, Roman times; ) river, a right tributary of the Sava, forms a natural border between north-west Croatia and southeast Slovenia. It is long, with a length of serving as the border between Croati ...
. *In
Bosnia and Herzegovina Bosnia and Herzegovina, sometimes known as Bosnia-Herzegovina and informally as Bosnia, is a country in Southeast Europe. Situated on the Balkans, Balkan Peninsula, it borders Serbia to the east, Montenegro to the southeast, and Croatia to th ...
: in the upper and middle Una, with the upper and middle Sana; the upper and middle Vrbas, with the lower and middle Vrbanja; the main channel of the Bosna lost its population due to degradation of its waters, its banks and riverbed, but its four main tributaries, the Krivaja, the Fojnička, the
Lašva The Lašva ( sr-cyrl, Лашва) is a river in Central Bosnia, Bosnia and Herzegovina. It is a left tributary of the Bosna.Vojnogeografski institut, Ed. (1955): Travnik (List karte 1:100.000, Izohipse na 20 m). Vojnogeografski institut, Beograd / ...
and the Željeznica, are still important habitats; one of the most important huchen habitats in the Balkans and Europe is the Drina, with its numerous tributaries, especially the Lim, the Tara and the Ćehotina, and smaller ones, the Bistrica, the Prača, and the Drinjača; * In
Serbia , image_flag = Flag of Serbia.svg , national_motto = , image_coat = Coat of arms of Serbia.svg , national_anthem = () , image_map = , map_caption = Location of Serbia (gree ...
: the Lim. * In
Montenegro , image_flag = Flag of Montenegro.svg , image_coat = Coat of arms of Montenegro.svg , coa_size = 80 , national_motto = , national_anthem = () , image_map = Europe-Mont ...
: the Lim, with the Lake Plav, the Ćehotina, and the Tara. Some of these habitats, especially in the Balkans, are endangered with planned, or already implemented plans for construction of new dams and
hydroelectric power plant Hydroelectricity, or hydroelectric power, is Electricity generation, electricity generated from hydropower (water power). Hydropower supplies 15% of the world's electricity, almost 4,210 TWh in 2023, which is more than all other Renewable energ ...
s, such as on the Lim in Serbia, or like in case of the river Piva in Montenegro and Bosnia and Herzegovina, whose respective populations were completely wiped out since mid 20th century. In some cases rehabilitation of parts of the habitat is attempted, through restoration of the river course. Such positive example is the part of the course of the river Inn, with some -long stretch around the Bavarian town of Mühldorf was rebuilt and renaturalized and the huchen has returned in recent years. It has been introduced to other major river basins elsewhere on the continent and even North Africa, to rivers in
Morocco Morocco, officially the Kingdom of Morocco, is a country in the Maghreb region of North Africa. It has coastlines on the Mediterranean Sea to the north and the Atlantic Ocean to the west, and has land borders with Algeria to Algeria–Morocc ...
, but these populations are not self-sustaining. Some evidence suggest that in historic times the huchen has also been found in the neighboring
Dniester The Dniester ( ) is a transboundary river in Eastern Europe. It runs first through Ukraine and then through Moldova (from which it more or less separates the breakaway territory of Transnistria), finally discharging into the Black Sea on Uk ...
basin. Huchen sometimes successfully establish in accessible natural lakes, like glacial Lake Plav which is the source of the Lim river in Montenegro, one of the major huchen habitats in the Balkans. However, the species sometimes survives even when cut off from the rest of the population in big dam reservoirs on mountain rivers, such as reservoirs on the Drina in Bosnia, or Lake Czorsztyn in Poland, as long as competitive and/or allochthonous predator species are not introduced into the newly created lakes.


Ecology


Appearance

The huchen reaches about in length and more than in weight. The average length is between . The huchen has a slender body that is nearly round in cross-section. On the reddish brown back are several dark patches in an X or crescent shape, but most distinctive feature is its head, which is larger than in other salmonides (longer and wider), with large elongated mouth.


Diet

As the largest fish found within many European rivers, Huchen like many other salmonoids under an ontogenic change as they grow older. Smaller fish feed on the larvae of water insects or on insects dropped into the water. As they get older, larger individuals prey on other species of fish and small vertebrates. From a study in Slovakia, older angler caught individuals primarily seemed to prey on Spirlin ''Alburnoides bipunctatus,'' Chub ''Leuciscus cephalus'', Brown trout ''Salmo trutta'', and Barbel ''Barbus barbus.''


Historical occurrence

There is no international convention nor single agreed body concerned with verification of records and mandated with the task, however there are many modern and historical records for the largest huchen caught. Among these recorded is a fish weighing , caught in February 1985 in Spittal an der Drau in
Austria Austria, formally the Republic of Austria, is a landlocked country in Central Europe, lying in the Eastern Alps. It is a federation of nine Federal states of Austria, states, of which the capital Vienna is the List of largest cities in Aust ...
,; also one weighing , which is stuffed and was probably grown on fish-farm in Austria for breeding. Other storied examples of recorded specimens with existing photographs and media reports include , caught in January 1938 by Bosnian railway worker and angler Halil Sofradžija at the Dragojevića Rapids on the Drina river, near the town of Ustikolina in Bosnia and Herzegovina. This case was recorded on photograph and reportage was later published in prime Serbian daily
Politika ( sr-Cyrl, Политика, lit=Politics) is a Serbian daily newspaper, published in Belgrade. Founded in 1904 by Vladislav F. Ribnikar, it is the oldest daily newspaper still in circulation in the Balkans. Publishing and ownership is publ ...
on 23 February 1938. Dževat Šarkinović, angler from Plav, caught Danube salmon weighing , long, in late spring of 1985 in lake Plav, and gave it to Hotel "Plavsko jezero" who presented it at '' International Fair of Hunting and Fishing'' in Novi Sad where it was noted as a "world record". Today, this specimen stands as stuffed exhibit at the wall of hotel's foyer in Plav. Another recorded trophy from the Drina include huchen caught by angler Remzija Krkalić from
Foča Foča ( sr-Cyrl, Фоча, ) is a town and municipality in Republika Srpska, Bosnia and Herzegovina, located in the south-east on the banks of Drina river. As of 2013, the town has a population of 12,234 inhabitants, while the municipality has 1 ...
in 1940, weighing .


Reproduction

This permanent freshwater salmonid spawns in April, once water reaches a temperature of . For spawning, the huchen migrates up the river and enters small and shallow affluents, where females excavate depressions in the gravel into which they deposit their eggs, then a male releases a cloud of sperm and the female then covers the eggs with sand. Larvae hatch 30 to 35 days after fertilization.


Commercial breeding

There is a considerable effort to produce huchen fry commercially and to reintroduce the species into the wild with moderate to good success. This requires catching the adults just before spawning and keeping them in special tanks. Fry are released in appropriate places once they have reached .


References

{{Authority control Hucho Freshwater fish of Europe Fish described in 1758 Taxa named by Carl Linnaeus Habitats Directive species