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Hohlenstein-Stadel is a cave located in the Hohlenstein cliff (not to be confused with the Hohle Fels) at the southern rim of the Lonetal (valley of the Lone) in the
Swabian Jura The Swabian Jura ( , more rarely ), sometimes also named Swabian Alps in English, is a mountain range in Baden-Württemberg, Germany, extending from southwest to northeast and in width. It is named after the region of Swabia. It is part of th ...
in
Germany Germany, officially the Federal Republic of Germany, is a country in Central Europe. It lies between the Baltic Sea and the North Sea to the north and the Alps to the south. Its sixteen States of Germany, constituent states have a total popu ...
. While first excavations were started after the second half of the 19th century, the significance of some of the findings was not realized until 1969. The most significant finding was a small ivory statue called the '' Löwenmensch'', which is one of the oldest pieces of figurative art ever found. The Hohlenstein cliffs are made of limestone which was hollowed out by natural causes to create caves. The Stadel (meaning "barn") is one of three caves in the area that are of important paleontological and
archaeological Archaeology or archeology is the study of human activity through the recovery and analysis of material culture. The archaeological record consists of Artifact (archaeology), artifacts, architecture, biofact (archaeology), biofacts or ecofacts, ...
significance. The other two are the Kleine Scheuer ("little barn") and the Bärenhöhle ("bears' cave"). In 2017 the site became part of the UNESCO World Heritage Site
Caves and Ice Age Art in the Swabian Jura The Caves and Ice Age Art in the Swabian Jura are a collection of six caves in southern Germany which were used by Last Glacial Period, Ice Age humans for shelter about 33,000 to 43,000 years ago. Within the caves were found the oldest non-stati ...
.


Excavations

The first excavations at Hohlenstein were made in 1861 by Oskar Fraas, geologist and paleontologist, who was searching for
cave bear The cave bear (''Ursus spelaeus'') is a prehistoric species of bear that lived in Europe and Asia during the Pleistocene and became extinct about 24,000 years ago during the Last Glacial Maximum. Both the word ''cave'' and the scientific name '' ...
bones at the Bärenhöhle and the Stadel. He returned in 1866, realizing the archeological importance of the site. In 1935, archaeologists returned to excavate the cave. An initial trial dig under Robert Wetzel was followed by further work in 1936. Between 1937 and 1939 further excavations were conducted directed by Wetzel and Otto Völzing, a geologist. On the last day of digging, on 25 August 1939, Völzing found a large number of broken pieces of ivory. They were little noted and went into storage at the Museum Ulm. It was not until 1969 that Joachim Hahn came across the more than 200 pieces and assembled them into a 31-cm-tall figurine of a humanoid with a lion's head. This is now known as the '' Löwenmensch''. At 35,000 to 40,000 years old, it qualifies as one of the oldest pieces of figurative art ever discovered. Although the significance of figurines such as this is still unknown, they may have been effigies of a primitive religion. Further excavations followed in 1956 and 1957, and between 1959 and 1961. The stratigraphy includes layers from the
Neolithic The Neolithic or New Stone Age (from Ancient Greek, Greek 'new' and 'stone') is an archaeological period, the final division of the Stone Age in Mesopotamia, Asia, Europe and Africa (c. 10,000 BCE to c. 2,000 BCE). It saw the Neolithic Revo ...
,
Mesolithic The Mesolithic (Ancient Greek language, Greek: μέσος, ''mesos'' 'middle' + λίθος, ''lithos'' 'stone') or Middle Stone Age is the Old World archaeological period between the Upper Paleolithic and the Neolithic. The term Epipaleolithic i ...
,
Upper Paleolithic The Upper Paleolithic (or Upper Palaeolithic) is the third and last subdivision of the Paleolithic or Old Stone Age. Very broadly, it dates to between 50,000 and 12,000 years ago (the beginning of the Holocene), according to some theories ...
, notably the
Magdalenian Magdalenian cultures (also Madelenian; ) are later cultures of the Upper Paleolithic and Mesolithic in western Europe. They date from around 17,000 to 12,000 years before present. It is named after the type site of Abri de la Madeleine, a ro ...
and the
Aurignacian The Aurignacian () is an archaeological industry of the Upper Paleolithic associated with Cro-Magnon, Early European modern humans (EEMH) lasting from 43,000 to 26,000 years ago. The Upper Paleolithic developed in Europe some time after the L ...
periods and finally the
Middle Paleolithic The Middle Paleolithic (or Middle Palaeolithic) is the second subdivision of the Paleolithic or Old Stone Age as it is understood in Europe, Africa and Asia. The term Middle Stone Age is used as an equivalent or a synonym for the Middle P ...
. More work followed in 1996 and 1997, when a dig led by Nicholas Conard, Michael Bolus and Andrew Kandel was conducted in the valley in front of the caves. Interior excavation was resumed in 2008 to 2013 by the (state office for monument protection). This work, led by Thomas Beutelspacher and C.J. Kind managed to locate the exact spot in which the Lion-man had been discovered and to find numerous additional ivory splinters that were found to fit onto the figurine. Dating of bones found immediately next to them yielded an age of 35,000 to 41,000 years. In addition to the Lion-man figurine, pendants carved from mammoth ivory and perforated animal teeth dating from the Aurignacian have been uncovered at the cave.


Archaeogenetics

On August 27, 1937, excavators discovered the right femur
diaphysis The diaphysis (: diaphyses) is the main or midsection (shaft) of a long bone. It is made up of cortical bone and usually contains bone marrow and adipose tissue (fat). It is a middle tubular part composed of compact bone which surrounds a centr ...
, measuring around in length, of an archaic hominin in the cave. The femur came from a layer associated with Middle Paleolithic
Mousterian The Mousterian (or Mode III) is an Industry (archaeology), archaeological industry of Lithic technology, stone tools, associated primarily with the Neanderthals in Europe, and with the earliest anatomically modern humans in North Africa and We ...
artefacts. This femur represents the only archaic hominin fossil found in a Mousterian context within the entire Swabian Jura region. Attempts to radiocarbon date the femur have yielded inconsistent results; however, molecular dating suggests that the femur is roughly 124,000 years old. In 2017, researchers successfully sequenced the full
mtDNA Mitochondrial DNA (mtDNA and mDNA) is the DNA located in the mitochondria organelles in a eukaryotic cell that converts chemical energy from food into adenosine triphosphate (ATP). Mitochondrial DNA is a small portion of the DNA contained in ...
genome from the femur. The results confirmed that the femur belonged to a
Neanderthal Neanderthals ( ; ''Homo neanderthalensis'' or sometimes ''H. sapiens neanderthalensis'') are an extinction, extinct group of archaic humans who inhabited Europe and Western and Central Asia during the Middle Pleistocene, Middle to Late Plei ...
. The mtDNA from the Hohlenstein-Stadel sample is highly divergent from those of other available Neanderthal samples. The addition of this mtDNA sample itself results in a near doubling of the genetic diversity of available Neanderthal mtDNA using Watterson's estimator theta; this suggests that Neanderthal mtDNA diversity was higher than previously presumed. Researchers estimate that the Hohlenstein-Stadel mtDNA diverged from other Neanderthal lineages around 270,000 years ago. This article contains quotations from this source, which is available under th
Creative Commons Attribution 4.0 International (CC BY 4.0)
license.


UNESCO World Heritage Site

In January 2016, the federal government of Germany applied for the status of
World Heritage Site World Heritage Sites are landmarks and areas with legal protection under an treaty, international treaty administered by UNESCO for having cultural, historical, or scientific significance. The sites are judged to contain "cultural and natural ...
for two valleys with six caves named ("caves with the oldest Ice Age art"). The site would encompass areas in the Lonetal (valley of the Lone) and the Achtal (valley of the Ach), both in the southern Swabian Jura. The former includes the caves ''Hohlenstein-Stadel'', '' Vogelherd'' and ''Bocksteinhöhle'', the latter '' Geissenklösterle'', '' Hohle Fels'' and '' Sirgenstein Cave''. Each valley would contain a core area of around length, surrounded by a buffer zone of a least width. In the argument why these sites deserve recognition as a part of the universal human heritage, the area is described as the source of the currently oldest (non-stationary) works of human art in the form of carved animal and humanoid figurines as well as the oldest musical instruments. Their creators lived, were inspired, and worked in and around these caves. The caves also served as the repositories of the figurines which may have been used in a religious context. In addition, they were the venue where performers used the excavated musical instruments and where the social groups lived from which the artists sprang. The committee awarded the status of WHS in July 2017.


References


External links

*
Hohlenstein-Stadel at a Lonetal tourism website
{{Coord, 48, 32, 57.57, N, 10, 10, 20.75, E, region:DE_type:landmark, display=no World Heritage Sites in Germany Caves of Germany Archaeological sites in Germany Geography of Baden-Württemberg Landforms of Baden-Württemberg Stone Age Germany Upper Paleolithic sites in Europe Magdalenian Aurignacian Middle Paleolithic Mesolithic sites of Europe Neolithic sites of Europe Cave bear