Hippodamia Convergens
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''Hippodamia convergens'', commonly known as the convergent lady beetle, is one of the most common lady beetles in
North America North America is a continent in the Northern Hemisphere, Northern and Western Hemisphere, Western hemispheres. North America is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South Ameri ...
and is found throughout the continent. They tend to live a variety of habitats, including grasslands and forests. Female ''H. convergens'' can lay over 1000 eggs over the span of a few months during the spring or early summer. In some populations, the beetles may undergo diapause if there are limited food resources to delay reproduction. ''H. convergens'' eat soft-bodied insects, with
aphids Aphids are small sap-sucking insects in the Taxonomic rank, family Aphididae. Common names include greenfly and blackfly, although individuals within a species can vary widely in color. The group includes the fluffy white Eriosomatinae, woolly ...
being the primary food resource. Aphids are a known pest, so the ''H. convergens'' has been used as a method to control aphids by releasing the beetles to act as a predator for the aphids.


Range

Convergent lady beetles are native to North America, but have also been found in South America after they were imported from California.


Habitat

''H. convergens'' tend to be more successful and more likely to survive to adulthood in warmer temperatures. In some areas, they gather on sunflower patches, having moved from wheat fields. They use the petioles of the sunflowers to hydrate, particularly in arid summer months. They populate grasslands, forests, agricultural fields, gardens, and national parks. These beetles have been shown to have a lower temperature tolerance of 6.5°C (approximately 43°F) and an upper temperature tolerance of 50°C (approximately 122°F). They have been found to achieve optimal reproductive and survival rates at 25.12°C (approximately 77°F) with a relative humidity of 63.78%.


Life cycle

The female lady beetle lays 200 to 1000
eggs An egg is an organic vessel in which an embryo begins to develop. Egg, EGG or eggs may also refer to: Biology * Egg cell, the female reproductive cell (gamete) in oogamous organisms Food * Eggs as food Places * Egg, Austria * Egg, Switzerland ...
over several months during spring and early summer. The eggs are small and spindle-shaped and are laid near the prey in upright batches of fifteen to thirty eggs. The
larva A larva (; : larvae ) is a distinct juvenile form many animals undergo before metamorphosis into their next life stage. Animals with indirect development such as insects, some arachnids, amphibians, or cnidarians typically have a larval phase ...
e are dark and somewhat alligator-shaped. Once the larvae begin feeding, they grow quickly and
molt In biology, moulting (British English), or molting (American English), also known as sloughing, shedding, or in many invertebrates, ecdysis, is a process by which an animal casts off parts of its body to serve some beneficial purpose, either at ...
four times over a period of up to a month. Larvae generally move between plants by traveling across leaves. However, they are able to travel via soil if the leaves cannot be crossed. The pupal stage lasts about a week and mating takes place soon after adult
eclosion A pupa (; : pupae) is the life stage of some insects undergoing transformation between immature and mature stages. Insects that go through a pupal stage are holometabolous: they go through four distinct stages in their life cycle, the stages th ...
. If the food supply is abundant, the female may start laying within about a week of mating, but if it is scarce, she may wait for up to nine months. Upon reaching the adult stage, females feed on fats and proteins for a week. This increases the production of
juvenile hormone Juvenile hormones (JHs) are a group of acyclic sesquiterpenoids that regulate many aspects of insect physiology. The first discovery of a JH was by Vincent Wigglesworth. JHs regulate development, reproduction, diapause, and polyphenisms. In ...
, helping the ovaries mature. This hormone also causes a behavior that results in long distance migration. In the western United States, these beetles may spend up to nine months in
diapause In animal dormancy, diapause is the delay in development in response to regular and recurring periods of adverse environmental conditions.Tauber, M.J., Tauber, C.A., Masaki, S. (1986) ''Seasonal Adaptations of Insects''. Oxford University Press It ...
in large groups in mountain valleys. Some populations have been shown to undergo diapause when nutrients are scarce, using limited food resources to develop fat bodies and postpone the onset of reproduction until they can find a consistent and sufficient food source. During diapause, adult females are known to actively engage in flight.


Diet

Both larvae and adult ''H. convergens'' primarily consume
aphids Aphids are small sap-sucking insects in the Taxonomic rank, family Aphididae. Common names include greenfly and blackfly, although individuals within a species can vary widely in color. The group includes the fluffy white Eriosomatinae, woolly ...
. They are active hunters, meaning that they mobilize and travel to hunt for their prey. This also means that the larvae do not rely on helpers at the very least to gather food, meaning that the larvae will start searching for prey almost immediately upon hatching. The first larvae that hatch in each batch may start by eating the unhatched eggs. This may provide energy for the larvae before they find any aphids. Fourth-
instar An instar (, from the Latin '' īnstar'' 'form, likeness') is a developmental stage of arthropods, such as insects, which occurs between each moult (''ecdysis'') until sexual maturity is reached. Arthropods must shed the exoskeleton in order to ...
larvae may consume about fifty aphids per day and adults may eat about twenty. When aphids are scarce, the adults can eat honeydew,
nectar Nectar is a viscous, sugar-rich liquid produced by Plant, plants in glands called nectaries, either within the flowers with which it attracts pollination, pollinating animals, or by extrafloral nectaries, which provide a nutrient source to an ...
and
pollen Pollen is a powdery substance produced by most types of flowers of seed plants for the purpose of sexual reproduction. It consists of pollen grains (highly reduced Gametophyte#Heterospory, microgametophytes), which produce male gametes (sperm ...
or even petals and other soft parts of plants. ''H. convergens'' feed on other soft-bodied insects such as
scales Scale or scales may refer to: Mathematics * Scale (descriptive set theory), an object defined on a set of points * Scale (ratio), the ratio of a linear dimension of a model to the corresponding dimension of the original * Scale factor, a number ...
and
thrips Thrips (Order (biology) , order Thysanoptera) are minute (mostly long or less), slender insects with fringed wings and unique asymmetrical mouthparts. Entomologists have species description , described approximately 7,700 species. They fly on ...
. They are also known to exhibit cannibalistic behaviors when food is especially scarce.


Biological control

Convergent lady beetles have been used for augmentative biological control to temporarily increase predator numbers to control aphids. Because of the overwintering habits of non-reproductive adults, released beetles tend to quickly disperse from their release site. Adults released in enclosed settings such as greenhouses can contribute to lower aphid numbers. However, they tend to disperse before mating and laying eggs, so eggs are not left behind to hatch and continue the cycle of controlling the aphid population. This occurs even when live prey is still present. Beetles of this species used for biological control that are in a state of diapause have been known to not consume prey. This species was not included in the list of predatory insects usable for population control in the 2021 guidelines issued by the University of Florida, Institute of Food and Agricultural Sciences.


Natural enemies

Entomopathogenic fungi Entomopathogenic fungi are parasitic unicellular or multicellular microorganisms belonging to the kingdom of Fungi, that can infect and seriously disable or kill insects. Pathogenicity for insects is widely distributed in the kingdom of fungi and ...
used as biopesticides such as ''
Metarhizium anisopliae ''Metarhizium anisopliae'' is a fungus, the type species in the ''Metarhizium'' genus. It grows naturally in soils throughout the world and causes disease in various insects by acting as a parasitoid. Ilya Ilyich Mechnikov, Ilya I. Mechnikov nam ...
'', '' Paecilomyces fumosoroseus'', and ''
Beauveria bassiana ''Beauveria bassiana'' is a fungus that grows naturally in soils throughout the world and acts as a parasite on various arthropod species, causing white muscardine disease; it thus belongs to the group of entomopathogenic fungi. It is used as a ...
'' can infect larvae. Infection by ''Beauveria bassiana'' in particular has been shown to affect the temperatures these beetles will tolerate. ''
Geocoris bullatus ''Geocoris bullatus'', the large big-eyed bug, is a species of big-eyed bug in the family Geocoridae. It is found in North America. Subspecies These three subspecies belong to the species ''Geocoris bullatus'': * ''Geocoris bullatus borealis'' ...
'' and '' Nabis alternatus'' prey on ''H. convergens'' eggs. ''H. convergens'' may be a host for different invertebrate parasites such as '' Dinocampus coccinellae'', '' Homalotylus terminalis'', and '' Tetrapolipus hippodamiae''.


References


External links


Natural Enemies Gallery , Convergent Lady Beetle
''
University of California The University of California (UC) is a public university, public Land-grant university, land-grant research university, research university system in the U.S. state of California. Headquartered in Oakland, California, Oakland, the system is co ...
Statewide Integrated Pest Management Program''.
Biological Control: Lady Beetles , (''Coleoptera: Coccinellidae'')
''
Cornell University Cornell University is a Private university, private Ivy League research university based in Ithaca, New York, United States. The university was co-founded by American philanthropist Ezra Cornell and historian and educator Andrew Dickson W ...
College of Agricaulture and Life Sciences''. Archived fro
the original
on 27 November 2011.
OSU Agricultural Extension Fact Sheet - ENT-45

Species Hippodamia convergens - Convergent Lady Beetle
''Bugs Guides''.
Convergent Lady Beetle Video on Vimeo
Macro videography of Convergent Lady Beetle colony {{Taxonbar, from=Q1938912 Coccinellidae Insects used as insect pest control agents Biological pest control beetles Beetles of North America Beetles described in 1842