Heterometrus Longimanus
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''Heterometrus'', whose members are also known by the collective
vernacular name Vernacular is the ordinary, informal, spoken form of language, particularly when perceived as having lower social status or less prestige than standard language, which is more codified, institutionally promoted, literary, or formal. More n ...
Asian Forest Scorpion, is a
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of
scorpions Scorpions are predatory arachnids of the Order (biology), order Scorpiones. They have eight legs and are easily recognized by a pair of Chela (organ), grasping pincers and a narrow, segmented tail, often carried in a characteristic forward cur ...
belonging to the family
Scorpionidae Scorpionidae is a family of burrowing scorpions or pale-legged scorpions in the Taxonomic rank, superfamily Scorpionoidea. The family was established by Pierre André Latreille, 1802. Genera Scorpionidae contains the following genera: * ''Aops ...
. It is distributed widely across
tropical The tropics are the regions of Earth surrounding the equator, where the sun may shine directly overhead. This contrasts with the temperate or polar regions of Earth, where the Sun can never be directly overhead. This is because of Earth's ax ...
and
subtropical The subtropical zones or subtropics are geographical zone, geographical and Köppen climate classification, climate zones immediately to the Northern Hemisphere, north and Southern Hemisphere, south of the tropics. Geographically part of the Ge ...
southeastern Asia Southeast Asia is the geographical southeastern region of Asia, consisting of the regions that are situated south of China, east of the Indian subcontinent, and northwest of the Australian mainland, which is part of Oceania. Southeast Asia is ...
, including
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,
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,
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,
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,
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,
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,
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,
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,
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,
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,
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(
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,
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), and
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(
Hainan Hainan is an island provinces of China, province and the southernmost province of China. It consists of the eponymous Hainan Island and various smaller islands in the South China Sea under the province's administration. The name literally mean ...
). It is notable for containing some of the largest living
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
of scorpions.


Taxonomy

The genus was introduced by C.G. Ehrenberg (in Hemprich & Ehrenberg, 1828), originally as a
subgenus In biology, a subgenus ( subgenera) is a taxonomic rank directly below genus. In the International Code of Zoological Nomenclature, a subgeneric name can be used independently or included in a species name, in parentheses, placed between the ge ...
of the genus ''
Buthus ''Buthus'' is a genus of scorpion belonging and being eponymous to the family Buthidae. It is distributed widely across northern Africa, including Morocco, Mauritania, Algeria, Tunisia, Libya, Egypt, Senegal, Guinea-Bissau, Nigeria, Sudan, Somali ...
''. It was elevated to genus rank by F. Karsch in 1879. H.W.C. Couzijn (1978, 1981) subdivided the genus into several
subgenera In biology, a subgenus ( subgenera) is a taxonomic rank directly below genus. In the International Code of Zoological Nomenclature, a subgeneric name can be used independently or included in a species name, in parentheses, placed between the ge ...
, but F. Kovařík (2004)
synonym A synonym is a word, morpheme, or phrase that means precisely or nearly the same as another word, morpheme, or phrase in a given language. For example, in the English language, the words ''begin'', ''start'', ''commence'', and ''initiate'' are a ...
ized these subgenera with the nominal genus. In 2020 the genus was reviewed by L. Prendini & S. F. Loria, three of the former subgenera were revalidated and elevated to genera and one valid subgenus was elevated to genus rank, species were transferred to appropriate genera, resulting in 28 new combinations.


Diversity

The content of this genus may vary, depending on the authority. Eight species are known, many of which are quite similar in appearance: *'' Heterometrus glaucus'' (Thorell, 1876) *'' Heterometrus laevigatus'' (Thorell, 1876) *'' Heterometrus laoticus'' Couzijn, 1981 *'' Heterometrus longimanus'' (Herbst, 1800) *'' Heterometrus petersii'' (Thorell, 1876) *'' Heterometrus silenus'' (Simon, 1884) *''
Heterometrus spinifer ''Heterometrus spinifer'', or its common name, Malaysian forest scorpion or in Japan Malaysian Cancer Scorpion (マレーキャンサースコーピオン), is a species of scorpions belonging to the family Scorpionidae. Description ''H. spini ...
'' (Ehrenberg, 1828) *'' Heterometrus thorellii'' (Pocock, 1897)


General characteristics

Members of ''Heterometrus'' are generally large-sized
scorpion Scorpions are predatory arachnids of the Order (biology), order Scorpiones. They have eight legs and are easily recognized by a pair of Chela (organ), grasping pincers and a narrow, segmented tail, often carried in a characteristic forward cur ...
s (100–200 mm or about 4-8  in total length). Coloration is dark in most species, often uniformly brown or black, sometimes with a greenish shine, with brighter-colored
telson The telson () is the hindmost division of the body of an arthropod. Depending on the definition, the telson is either considered to be the final segment (biology), segment of the arthropod body, or an additional division that is not a true segm ...
, walking legs, and/or
pedipalp Pedipalps (commonly shortened to palps or palpi) are the secondary pair of forward appendages among Chelicerata, chelicerates – a group of arthropods including spiders, scorpions, horseshoe crabs, and sea spiders. The pedipalps are lateral to ...
pincers in some species. The scorpions are heavily built with especially powerful and globose pedipalp pionkes, broad
mesosoma The mesosoma is the middle part of the body, or tagma, of arthropods whose body is composed of three parts, the other two being the prosoma and the metasoma. It bears the legs, and, in the case of winged insects, the wings. Wasps, bees and a ...
l
tergites A ''tergum'' (Latin for "the back"; : ''terga'', associated adjective tergal) is the dorsal ('upper') portion of an arthropod segment other than the head. The anterior edge is called the 'base' and posterior edge is called the 'apex' or 'margin'. ...
and a proportionally slender and thin
metasoma The metasoma is the posterior part of the body, or tagma (biology), tagma, of arthropods whose body is composed of three parts, the other two being the prosoma and the mesosoma. In insects, it contains most of the digestive tract, respiratory sy ...
. The telson is proportionally small and the stinger is often shorter than the vesicle. The
cephalothorax The cephalothorax, also called prosoma in some groups, is a tagma of various arthropods, comprising the head and the thorax fused together, as distinct from the abdomen behind. (The terms ''prosoma'' and ''opisthosoma'' are equivalent to ''cepha ...
and mesosoma are largely devoid of carinae and granulation and the median eyes are situated in a small, lenticular depression on the cephalothorax. Some species are parthenogenic.


Toxicity

As in other genera of the Scorpionidae, the symptoms from ''Heterometrus'' envenomations are rather mild and no human fatalities are known. The sting causes local pain,
inflammation Inflammation (from ) is part of the biological response of body tissues to harmful stimuli, such as pathogens, damaged cells, or irritants. The five cardinal signs are heat, pain, redness, swelling, and loss of function (Latin ''calor'', '' ...
,
oedema Edema (American English), also spelled oedema (British English), and also known as fluid retention, swelling, dropsy and hydropsy, is the build-up of fluid in the body's tissue. Most commonly, the legs or arms are affected. Symptoms may inclu ...
, swelling, and redness of the skin, lasting for hours to a few days. Plant extracts known in traditional Thai medicine as natural scorpion venom antidotes are effective as symptomatic treatment of ''H. laoticus'' stings. The
protein Proteins are large biomolecules and macromolecules that comprise one or more long chains of amino acid residue (biochemistry), residues. Proteins perform a vast array of functions within organisms, including Enzyme catalysis, catalysing metab ...
heteroscorpine-1 was found the major component of the venom in ''H. laoticus''.


Habitat for the scorpion

Species of ''Heterometrus'' live in vegetated, often forested,
humid Humidity is the concentration of water vapor present in the air. Water vapor, the gaseous state of water, is generally invisible to the human eye. Humidity indicates the likelihood for precipitation, dew, or fog to be present. Humidity depe ...
regions with
subtropical The subtropical zones or subtropics are geographical zone, geographical and Köppen climate classification, climate zones immediately to the Northern Hemisphere, north and Southern Hemisphere, south of the tropics. Geographically part of the Ge ...
to
tropical The tropics are the regions of Earth surrounding the equator, where the sun may shine directly overhead. This contrasts with the temperate or polar regions of Earth, where the Sun can never be directly overhead. This is because of Earth's ax ...
climate Climate is the long-term weather pattern in a region, typically averaged over 30 years. More rigorously, it is the mean and variability of meteorological variables over a time spanning from months to millions of years. Some of the meteoro ...
s. As most scorpions, they are predominantly
nocturnal Nocturnality is a ethology, behavior in some non-human animals characterized by being active during the night and sleeping during the day. The common adjective is "nocturnal", versus diurnality, diurnal meaning the opposite. Nocturnal creatur ...
and hide in
burrow file:Chipmunk-burrow (exits).jpg, An eastern chipmunk at the entrance of its burrow A burrow is a hole or tunnel excavated into the ground by an animal to construct a space suitable for habitation or temporary refuge, or as a byproduct of Animal lo ...
s, below logs, and in
leaf litter Plant litter (also leaf litter, tree litter, soil litter, litterfall, or duff) is dead plant material (such as leaves, bark, needles, twigs, and cladodes) that has fallen to the ground. This detritus or dead organic material and its constituen ...
.


In captivity

Due to their impressive size, low toxicity, and docile behavior, species of ''Heterometrus'' are popular pet scorpions. Unlike many other scorpions, they can be kept in pairs or small groups.


References


External links

* Keeping instructions for ''H. spinifer'' and ''H. laoticus''
Exotic Pets
* Images and descriptions of various species of ''Heterometrus''
Mes Scorpions
{{Authority control Scorpionidae Scorpion genera Taxa named by Christian Gottfried Ehrenberg