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''Hepatica'' (hepatica, liverleaf, or liverwort) is a
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of
herbaceous Herbaceous plants are vascular plants that have no persistent woody stems above ground. This broad category of plants includes many perennials, and nearly all annuals and biennials. Definitions of "herb" and "herbaceous" The fourth edition of ...
perennials In horticulture, the term perennial ('' per-'' + '' -ennial'', "through the year") is used to differentiate a plant from shorter-lived annuals and biennials. It has thus been defined as a plant that lives more than 2 years. The term is also ...
in the
buttercup family Ranunculaceae (, buttercup or crowfoot family; Latin "little frog", from "frog") is a family of over 2,000 known species of flowering plants in 43 genera, distributed worldwide. The largest genera are ''Ranunculus'' (600 species), ''Delphinium ...
, native to central and northern
Europe Europe is a continent located entirely in the Northern Hemisphere and mostly in the Eastern Hemisphere. It is bordered by the Arctic Ocean to the north, the Atlantic Ocean to the west, the Mediterranean Sea to the south, and Asia to the east ...
,
Asia Asia ( , ) is the largest continent in the world by both land area and population. It covers an area of more than 44 million square kilometres, about 30% of Earth's total land area and 8% of Earth's total surface area. The continent, which ...
and eastern
North America North America is a continent in the Northern Hemisphere, Northern and Western Hemisphere, Western hemispheres. North America is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South Ameri ...
. Some botanists include ''Hepatica'' within a wider interpretation of ''
Anemone ''Anemone'' () is a genus of flowering plants in the buttercup family Ranunculaceae. Plants of the genus are commonly called windflowers. They are native to the temperate and subtropical regions of all regions except Australia, New Zealand, and ...
''.


Description

Bisexual Bisexuality is romantic attraction, sexual attraction, or sexual behavior toward both males and females. It may also be defined as the attraction to more than one gender, to people of both the same and different gender, or the attraction t ...
flower Flowers, also known as blooms and blossoms, are the reproductive structures of flowering plants ( angiosperms). Typically, they are structured in four circular levels, called whorls, around the end of a stalk. These whorls include: calyx, m ...
s with pink, purple, blue, or white
sepal A sepal () is a part of the flower of angiosperms (flowering plants). Usually green, sepals typically function as protection for the flower in bud, and often as support for the petals when in bloom., p. 106 Etymology The term ''sepalum'' ...
s and three green
bract In botany, a bract is a modified or specialized leaf, associated with a reproductive structure such as a flower, inflorescence axis or cone scale. Bracts are usually different from foliage leaves in size, color, shape or texture. They also lo ...
s appear singly on hairy
stem Stem or STEM most commonly refers to: * Plant stem, a structural axis of a vascular plant * Stem group * Science, technology, engineering, and mathematics Stem or STEM can also refer to: Language and writing * Word stem, part of a word respon ...
s from late
winter Winter is the coldest and darkest season of the year in temperate and polar climates. It occurs after autumn and before spring. The tilt of Earth's axis causes seasons; winter occurs when a hemisphere is oriented away from the Sun. Dif ...
to
spring Spring(s) may refer to: Common uses * Spring (season), a season of the year * Spring (device), a mechanical device that stores energy * Spring (hydrology), a natural source of water * Spring (mathematics), a geometric surface in the shape of a he ...
.
Butterflies Butterflies are winged insects from the lepidopteran superfamily Papilionoidea, characterized by large, often brightly coloured wings that often fold together when at rest, and a conspicuous, fluttering flight. The oldest butterfly fossi ...
,
moth Moths are a group of insects that includes all members of the order Lepidoptera that are not Butterfly, butterflies. They were previously classified as suborder Heterocera, but the group is Paraphyly, paraphyletic with respect to butterflies (s ...
s,
bee Bees are winged insects closely related to wasps and ants, known for their roles in pollination and, in the case of the best-known bee species, the western honey bee, for producing honey. Bees are a monophyletic lineage within the superfamil ...
s,
flies Flies are insects of the Order (biology), order Diptera, the name being derived from the Ancient Greek, Greek δι- ''di-'' "two", and πτερόν ''pteron'' "wing". Insects of this order use only a single pair of wings to fly, the hindwin ...
and
beetle Beetles are insects that form the Taxonomic rank, order Coleoptera (), in the superorder Holometabola. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 40 ...
s are known
pollinator A pollinator is an animal that moves pollen from the male anther of a flower to the female carpel, stigma of a flower. This helps to bring about fertilization of the ovules in the flower by the male gametes from the pollen grains. Insects are ...
s. The
leaves A leaf (: leaves) is a principal appendage of the stem of a vascular plant, usually borne laterally above ground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, stem, ...
are basal, leathery, and usually three-lobed, remaining over
winter Winter is the coldest and darkest season of the year in temperate and polar climates. It occurs after autumn and before spring. The tilt of Earth's axis causes seasons; winter occurs when a hemisphere is oriented away from the Sun. Dif ...
.


Taxonomy

''Hepatica'' was described by the English
botanist Botany, also called plant science, is the branch of natural science and biology studying plants, especially Plant anatomy, their anatomy, Plant taxonomy, taxonomy, and Plant ecology, ecology. A botanist or plant scientist is a scientist who s ...
Philip Miller Philip Miller Royal Society, FRS (1691 – 18 December 1771) was an English botany, botanist and gardener of Scottish descent. Miller was chief gardener at the Chelsea Physic Garden for nearly 50 years from 1722, and wrote the highly popular ...
in 1754. It was proposed as a
subgenus In biology, a subgenus ( subgenera) is a taxonomic rank directly below genus. In the International Code of Zoological Nomenclature, a subgeneric name can be used independently or included in a species name, in parentheses, placed between the ge ...
of ''
Anemone ''Anemone'' () is a genus of flowering plants in the buttercup family Ranunculaceae. Plants of the genus are commonly called windflowers. They are native to the temperate and subtropical regions of all regions except Australia, New Zealand, and ...
'' in 1836, but later segregated into genus ''Hepatica''.


Taxa

,
Kew Kew () is a district in the London Borough of Richmond upon Thames. Its population at the 2011 census was 11,436. Kew is the location of the Royal Botanic Gardens ("Kew Gardens"), now a World Heritage Site, which includes Kew Palace. Kew is ...
's
Plants of the World Online Plants of the World Online (POWO) is an online taxonomic database published by the Royal Botanic Gardens, Kew. History Following the Convention on Biological Diversity, the Royal Botanic Gardens in Kew launched Plants of the World Online i ...
(POWO) accepts 7 
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
and one
hybrid Hybrid may refer to: Science * Hybrid (biology), an offspring resulting from cross-breeding ** Hybrid grape, grape varieties produced by cross-breeding two ''Vitis'' species ** Hybridity, the property of a hybrid plant which is a union of two diff ...
in the genus ''Hepatica'': One
infraspecific In botany, an infraspecific name is the scientific name for any taxon below the rank of species, i.e. an infraspecific taxon or infraspecies. The scientific names of botanical taxa are regulated by the ''International Code of Nomenclature for alg ...
taxon In biology, a taxon (back-formation from ''taxonomy''; : taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular name and ...
is also recognized by POWO: * ''Hepatica nobilis'' var. ''japonica'' ** Synonym: ''Hepatica asiatica'' ** Synonym: ''Hepatica insularis'' ''Hepatica'' can be divided into two
series Series may refer to: People with the name * Caroline Series (born 1951), English mathematician, daughter of George Series * George Series (1920–1995), English physicist Arts, entertainment, and media Music * Series, the ordered sets used i ...
with respect to leaf edge:


Series ''Triloba''

The leaves of the series ''Triloba'' Ulbr.Ulbrich, O.E.: Über die systematische Gliederung und geographische Verbreitung der Gattung Anemone L. Bot. Jahrb. Syst. (1905) 37: 172 - 257, 38: 257 - 334. Tamura:Tamura, M.: Morphology, ecology and phylogeny of the Ranunculaceae” VII. Science reports of South College, North College of Osaka University, Japan 16:21-43, 1968. are three-lobed with a smooth leaf edge. * ''Hepatica acutiloba'':
Eastern Canada Eastern Canada (, also the Eastern provinces, Canadian East or the East) is generally considered to be the region of Canada south of Hudson Bay/ Hudson Strait and east of Manitoba, consisting of the following provinces (from east to west): Newf ...
,
Midwestern United States The Midwestern United States (also referred to as the Midwest, the Heartland or the American Midwest) is one of the four census regions defined by the United States Census Bureau. It occupies the northern central part of the United States. It ...
,
Eastern United States The Eastern United States, often abbreviated as simply the East, is a macroregion of the United States located to the east of the Mississippi River. It includes 17–26 states and Washington, D.C., the national capital. As of 2011, the Eastern ...
* ''Hepatica americana'':
Central Canada Central Canada (, sometimes the ''Central Provinces'') is a Canadian region consisting of Ontario and Quebec, the largest and most populous provinces of the country. Geographically, they are not at the centre of Canada but instead overlap wi ...
, Eastern Canada, Midwestern United States, Eastern United States * ''Hepatica maxima'': endemic to
Ulleung-do Ulleungdo (), also spelled Ulreungdo, is a South Korean island east of the Korean Peninsula in the Sea of Japan. It was formerly known as Dagelet Island or Argonaut Island in Europe. Volcanic in origin, the rocky steep-sided island is the top o ...
island (South Korea) * ''Hepatica nobilis'': ** ''Hepatica nobilis'' var. ''nobilis'':
Europe Europe is a continent located entirely in the Northern Hemisphere and mostly in the Eastern Hemisphere. It is bordered by the Arctic Ocean to the north, the Atlantic Ocean to the west, the Mediterranean Sea to the south, and Asia to the east ...
to
Western Siberia Western Siberia or West Siberia ( rus, Западная Сибирь, p=ˈzapədnəjə sʲɪˈbʲirʲ; , ) is a region in North Asia. It is part of the wider region of Siberia that is mostly located in the Russia, Russian Federation, with a Sout ...
** ''Hepatica nobilis'' var. ''japonica'':
Russian Far East The Russian Far East ( rus, Дальний Восток России, p=ˈdalʲnʲɪj vɐˈstok rɐˈsʲiɪ) is a region in North Asia. It is the easternmost part of Russia and the Asia, Asian continent, and is coextensive with the Far Easte ...
,
North China North China () is a list of regions of China, geographical region of the People's Republic of China, consisting of five province-level divisions of China, provincial-level administrative divisions, namely the direct-administered municipalities ...
,
Central China Central China () is a List of regions of China, region in China. It mainly includes the provinces of China, provinces of Henan, Hubei and Hunan. Jiangxi is sometimes also regarded to be part of this region. Central China is now officially par ...
,
East China East China () is a geographical region in the People’s Republic of China, mainly consisting of seven province-level administrative divisions, namely the provinces (from north to south) Shandong, Jiangsu, Anhui, Zhejiang, Jiangxi, Fujian, ...
,
Korea Korea is a peninsular region in East Asia consisting of the Korean Peninsula, Jeju Island, and smaller islands. Since the end of World War II in 1945, it has been politically Division of Korea, divided at or near the 38th parallel north, 3 ...
, and Japan (
Honshu , historically known as , is the largest of the four main islands of Japan. It lies between the Pacific Ocean (east) and the Sea of Japan (west). It is the list of islands by area, seventh-largest island in the world, and the list of islands by ...
,
Shikoku is the smallest of the List of islands of Japan#Main islands, four main islands of Japan. It is long and between at its widest. It has a population of 3.8 million, the least populated of Japan's four main islands. It is south of Honshu ...
, north of
Kyushu is the third-largest island of Japan's Japanese archipelago, four main islands and the most southerly of the four largest islands (i.e. excluding Okinawa Island, Okinawa and the other Ryukyu Islands, Ryukyu (''Nansei'') Ryukyu Islands, Islands ...
island)


Series ''Angulosa''

The leaves of series ''Angulosa'' (Ulbr.) Tamura are three- to five-lobed with a
crenate A leaf (: leaves) is a principal appendage of the stem of a vascular plant, usually borne laterally above ground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, stem, fl ...
leaf edge. * ''Hepatica falconeri'' — Mountain forests of Central Asia; India: Northwest Himalayas (
Himachal Pradesh Himachal Pradesh (; Sanskrit: ''himācāl prādes;'' "Snow-laden Mountain Province") is a States and union territories of India, state in the northern part of India. Situated in the Western Himalayas, it is one of the thirteen Indian Himalayan ...
,
Jammu and Kashmir Jammu and Kashmir may refer to: * Jammu and Kashmir (union territory), a region administered by India as a union territory since 2019 * Jammu and Kashmir (state), a region administered by India as a state from 1952 to 2019 * Jammu and Kashmir (prin ...
); Northwest China ( Tienshan); Kyrgyzstan and Tajikistan (
Pamir-Alai The Pamir-Alay is a mountain system in Tajikistan, Kyrgyzstan and Uzbekistan, encompassing four main mountain ranges extending west from the Tian Shan Mountains, and located north of the main range of Pamir. They are variously considered part of ...
); North Pakistan, Kazakhstan (North Tienshan) * ''Hepatica henryi'': North Central China,
South Central China South Central China, South-Central China or Central-South China ( zh, c = 中南, p = Zhōngnán, l = Central-South), is a List of regions of China, region of China. It consists of eight provincial administrative regions, namely Henan, Hubei, ...
(western
Hubei Hubei is a province of China, province in Central China. It has the List of Chinese provincial-level divisions by GDP, seventh-largest economy among Chinese provinces, the second-largest within Central China, and the third-largest among inland ...
, northern
Hunan Hunan is an inland Provinces of China, province in Central China. Located in the middle reaches of the Yangtze watershed, it borders the Administrative divisions of China, province-level divisions of Hubei to the north, Jiangxi to the east, Gu ...
,
Sichuan Sichuan is a province in Southwestern China, occupying the Sichuan Basin and Tibetan Plateau—between the Jinsha River to the west, the Daba Mountains to the north, and the Yunnan–Guizhou Plateau to the south. Its capital city is Cheng ...
,
Shaanxi Shaanxi is a Provinces of China, province in north Northwestern China. It borders the province-level divisions of Inner Mongolia to the north; Shanxi and Henan to the east; Hubei, Chongqing, and Sichuan to the south; and Gansu and Ningxia to t ...
) * ''Hepatica × media'':
Romania Romania is a country located at the crossroads of Central Europe, Central, Eastern Europe, Eastern and Southeast Europe. It borders Ukraine to the north and east, Hungary to the west, Serbia to the southwest, Bulgaria to the south, Moldova to ...
* ''Hepatica transsilvanica'': Romania (
Carpathian Mountains The Carpathian Mountains or Carpathians () are a range of mountains forming an arc across Central Europe and Southeast Europe. Roughly long, it is the third-longest European mountain range after the Ural Mountains, Urals at and the Scandinav ...
,
Transylvania Transylvania ( or ; ; or ; Transylvanian Saxon dialect, Transylvanian Saxon: ''Siweberjen'') is a List of historical regions of Central Europe, historical and cultural region in Central Europe, encompassing central Romania. To the east and ...
)


Etymology

The word ''hepatica'' derives from the
Greek Greek may refer to: Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group *Greek language, a branch of the Indo-European language family **Proto-Greek language, the assumed last common ancestor of all kno ...
, from 'liver', because its three-lobed leaf was thought to resemble the human
liver The liver is a major metabolic organ (anatomy), organ exclusively found in vertebrates, which performs many essential biological Function (biology), functions such as detoxification of the organism, and the Protein biosynthesis, synthesis of var ...
.


Distribution

Plants of genus ''Hepatica'' are
native Native may refer to: People * '' Jus sanguinis'', nationality by blood * '' Jus soli'', nationality by location of birth * Indigenous peoples, peoples with a set of specific rights based on their historical ties to a particular territory ** Nat ...
to
Europe Europe is a continent located entirely in the Northern Hemisphere and mostly in the Eastern Hemisphere. It is bordered by the Arctic Ocean to the north, the Atlantic Ocean to the west, the Mediterranean Sea to the south, and Asia to the east ...
,
Asia Asia ( , ) is the largest continent in the world by both land area and population. It covers an area of more than 44 million square kilometres, about 30% of Earth's total land area and 8% of Earth's total surface area. The continent, which ...
, and
North America North America is a continent in the Northern Hemisphere, Northern and Western Hemisphere, Western hemispheres. North America is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South Ameri ...
. * Europe: Albania, Austria, the
Baltic states The Baltic states or the Baltic countries is a geopolitical term encompassing Estonia, Latvia, and Lithuania. All three countries are members of NATO, the European Union, the Eurozone, and the OECD. The three sovereign states on the eastern co ...
, Belarus, Bulgaria, Corsica, Czechoslovakia, Denmark,
European Russia European Russia is the western and most populated part of the Russia, Russian Federation. It is geographically situated in Europe, as opposed to the country's sparsely populated and vastly larger eastern part, Siberia, which is situated in Asia ...
, Finland, France, Germany, Greece, Hungary, Italy, Norway, Poland, Romania, Spain, Sweden, Switzerland, Ukraine, Yugoslavia * Central Asia: Kazakhstan, Kyrgyzstan, Tajikistan, Western Siberia * Eastern Asia:
North China North China () is a list of regions of China, geographical region of the People's Republic of China, consisting of five province-level divisions of China, provincial-level administrative divisions, namely the direct-administered municipalities ...
,
South Central China South Central China, South-Central China or Central-South China ( zh, c = 中南, p = Zhōngnán, l = Central-South), is a List of regions of China, region of China. It consists of eight provincial administrative regions, namely Henan, Hubei, ...
,
East China East China () is a geographical region in the People’s Republic of China, mainly consisting of seven province-level administrative divisions, namely the provinces (from north to south) Shandong, Jiangsu, Anhui, Zhejiang, Jiangxi, Fujian, ...
, Japan, Korea, Manchuria, Primorsky Krai * South Asia: Pakistan, Western Himalaya * Canada: Manitoba, New Brunswick, Nova Scotia, Ontario, Québec * United States: Alabama, Arkansas, Connecticut, Delaware, District of Columbia, Florida, Georgia, Illinois, Indiana, Iowa, Kentucky, Maine, Maryland, Massachusetts, Michigan, Minnesota, Mississippi, Missouri, New Hampshire, New Jersey, New York, North Carolina, Ohio, Pennsylvania, Rhode Island, South Carolina, Tennessee, Vermont, Virginia, West Virginia, Wisconsin Plants of the genus have been introduced to Belgium.


Cultivation

''Hepatica'' cultivation has been popular in Japan since the 18th century (mid-
Edo period The , also known as the , is the period between 1600 or 1603 and 1868 in the history of Japan, when the country was under the rule of the Tokugawa shogunate and some 300 regional ''daimyo'', or feudal lords. Emerging from the chaos of the Sengok ...
), where flowers with doubled petals and a range of colour patterns have been developed. Noted for its tolerance of
alkaline In chemistry, an alkali (; from the Arabic word , ) is a basic salt of an alkali metal or an alkaline earth metal. An alkali can also be defined as a base that dissolves in water. A solution of a soluble base has a pH greater than 7.0. The ...
limestone Limestone is a type of carbonate rock, carbonate sedimentary rock which is the main source of the material Lime (material), lime. It is composed mostly of the minerals calcite and aragonite, which are different Polymorphism (materials science) ...
-derived soils, ''Hepatica'' may grow in a wide range of conditions; it can be found either in deeply shaded deciduous (especially
beech Beech (genus ''Fagus'') is a genus of deciduous trees in the family Fagaceae, native to subtropical (accessory forest element) and temperate (as dominant element of Mesophyte, mesophytic forests) Eurasia and North America. There are 14 accepted ...
)
woodland A woodland () is, in the broad sense, land covered with woody plants (trees and shrubs), or in a narrow sense, synonymous with wood (or in the U.S., the '' plurale tantum'' woods), a low-density forest forming open habitats with plenty of sunli ...
and scrub or
grassland A grassland is an area where the vegetation is dominance (ecology), dominated by grasses (Poaceae). However, sedge (Cyperaceae) and rush (Juncaceae) can also be found along with variable proportions of legumes such as clover, and other Herbaceo ...
in full sun. ''Hepatica'' will also grow in both sandy and clay-rich substrates, being associated with
limestone Limestone is a type of carbonate rock, carbonate sedimentary rock which is the main source of the material Lime (material), lime. It is composed mostly of the minerals calcite and aragonite, which are different Polymorphism (materials science) ...
. Moist soil and winter snowfall are required; ''Hepatica'' is tolerant of winter snow cover, but less so of dry frost. Propagation is done by seeds or by dividing vigorous clumps in spring. However, seedlings take several years to reach bloom size, and divided plants are slow to thicken.


Uses

''Hepatica'' was once used as a
medicinal herb Medicinal plants, also called medicinal herbs, have been discovered and used in traditional medicine practices since prehistoric times. Plants synthesize hundreds of chemical compounds for various functions, including Plant defense against h ...
. Owing to the
doctrine of signatures The doctrine of signatures, also known as the doctrine of correspondences, states that herbs or animals have physical or behavioral traits that mirror the ailment it can successfully treat. Theological justifications, such as that of botanist Wil ...
, the plant was once thought to be an effective treatment for liver disorders. Although poisonous in large doses, the leaves and flowers may be used as an
astringent An astringent (sometimes called adstringent) is a chemical that shrinks or constricts body tissues. The word derives from the Latin '' adstringere'', which means "to bind fast". Astringency, the dry, puckering or numbing mouthfeel caused by t ...
, as a
demulcent A demulcent (derived from the "caress") is a mucilaginous or oleaginous preparation that forms a soothing protective film over a mucous membrane, relieving minor pain and inflammation of the membrane. However, they generally help for less than 3 ...
for slow-healing injuries, and as a
diuretic A diuretic () is any substance that promotes diuresis, the increased production of urine. This includes forced diuresis. A diuretic tablet is sometimes colloquially called a water tablet. There are several categories of diuretics. All diuretics ...
.


References


External links

* * * {{Taxonbar, from=Q157505 Ranunculaceae genera Medicinal plants of Asia Medicinal plants of Europe Medicinal plants of North America Taxa named by Philip Miller