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The hepatic tanager (''Piranga hepatica'') is a medium-sized American
songbird A songbird is a bird belonging to the suborder Passeri of the perching birds (Passeriformes). Another name that is sometimes seen as the scientific or vernacular name is Oscines, from Latin ''oscen'', "songbird". The Passeriformes contains 5,00 ...
. Formerly placed in the
tanager The tanagers (singular ) comprise the bird family (biology), family Thraupidae, in the order Passeriformes. The family has a Neotropical distribution and is the second-largest family of birds. It represents about 4% of all avian species and 12 ...
family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
(Thraupidae), it and other members of the genus
Piranga ''Piranga'' is a genus of birds long placed in the tanager family, but now considered members of the family Cardinalidae. The genus name ''Piranga'' is from Tupi language, Tupi word ''tijepiranga'', the name for an unknown small bird. Similar i ...
are now classified in the cardinal family (
Cardinalidae Cardinalidae (sometimes referred to as "cardinal-grosbeaks" or simply "cardinals") is a family of New World-Endemism, endemic passerine birds that consists of Cardinalis, cardinals, grosbeaks, and Passerina, buntings. It also includes several ot ...
).


Etymology

The common name ''hepatic'' means "liver-coloured", namely, brownish-red.
"(adj.) Liver-coloured, dark brownish-red; as in '' hepatic aloes'', ''hepatic tanager''."


Taxonomy

The IOC World Bird List recognises five subspecies of ''Piranga hepatica'', found in North and Central America. * ''Piranga hepatica hepatica'' (
California California () is a U.S. state, state in the Western United States that lies on the West Coast of the United States, Pacific Coast. It borders Oregon to the north, Nevada and Arizona to the east, and shares Mexico–United States border, an ...
,
Arizona Arizona is a U.S. state, state in the Southwestern United States, Southwestern region of the United States, sharing the Four Corners region of the western United States with Colorado, New Mexico, and Utah. It also borders Nevada to the nort ...
, and
New Mexico New Mexico is a state in the Southwestern United States, Southwestern region of the United States. It is one of the Mountain States of the southern Rocky Mountains, sharing the Four Corners region with Utah, Colorado, and Arizona. It also ...
in the United States and
Oaxaca Oaxaca, officially the Free and Sovereign State of Oaxaca, is one of the 32 states that compose the political divisions of Mexico, Federative Entities of the Mexico, United Mexican States. It is divided into municipalities of Oaxaca, 570 munici ...
in western Mexico) * ''Piranga hepatica dextra'' (New Mexico and western
Texas Texas ( , ; or ) is the most populous U.S. state, state in the South Central United States, South Central region of the United States. It borders Louisiana to the east, Arkansas to the northeast, Oklahoma to the north, New Mexico to the we ...
in the United States,
Chiapas Chiapas, officially the Free and Sovereign State of Chiapas, is one of the states that make up the Political divisions of Mexico, 32 federal entities of Mexico. It comprises Municipalities of Chiapas, 124 municipalities and its capital and large ...
in eastern Mexico (to Chiapas), and western Guatemala) * ''Piranga hepatica figlina'' (eastern
Guatemala Guatemala, officially the Republic of Guatemala, is a country in Central America. It is bordered to the north and west by Mexico, to the northeast by Belize, to the east by Honduras, and to the southeast by El Salvador. It is hydrologically b ...
and
Belize Belize is a country on the north-eastern coast of Central America. It is bordered by Mexico to the north, the Caribbean Sea to the east, and Guatemala to the west and south. It also shares a maritime boundary with Honduras to the southeast. P ...
* ''Piranga hepatica savannarum'' (eastern
Honduras Honduras, officially the Republic of Honduras, is a country in Central America. It is bordered to the west by Guatemala, to the southwest by El Salvador, to the southeast by Nicaragua, to the south by the Pacific Ocean at the Gulf of Fonseca, ...
and northeast
Nicaragua Nicaragua, officially the Republic of Nicaragua, is the geographically largest Sovereign state, country in Central America, comprising . With a population of 7,142,529 as of 2024, it is the third-most populous country in Central America aft ...
) * ''Piranga hepatica albifacies'' (highlands of eastern Guatemala and Nicaragua) Other taxonomic authorities don't recognise ''Piranga hepatica'' as a distinct species and include it in ''Piranga flava'', a more broadly defined hepatic tanager, with a range from the United States to Argentina. These authorities recognise three
subspecies In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...
groups, which are recognised as three species by the IOC. According to this taxonomy the three subspecies groups and their subspecies are as follows: * the ''hepatica'' group (northern hepatic tanager), breeding from
Nicaragua Nicaragua, officially the Republic of Nicaragua, is the geographically largest Sovereign state, country in Central America, comprising . With a population of 7,142,529 as of 2024, it is the third-most populous country in Central America aft ...
northwards to the United States, in
pine A pine is any conifer tree or shrub in the genus ''Pinus'' () of the family Pinaceae. ''Pinus'' is the sole genus in the subfamily Pinoideae. ''World Flora Online'' accepts 134 species-rank taxa (119 species and 15 nothospecies) of pines as cu ...
and pine-
oak An oak is a hardwood tree or shrub in the genus ''Quercus'' of the beech family. They have spirally arranged leaves, often with lobed edges, and a nut called an acorn, borne within a cup. The genus is widely distributed in the Northern Hemisp ...
forests and partially migratory. This is the species ''Piranga hepatica'' as recognised by the IOC, with the five subspecies listed above. * the ''lutea'' group (highland hepatic tanager), resident from
Costa Rica Costa Rica, officially the Republic of Costa Rica, is a country in Central America. It borders Nicaragua to the north, the Caribbean Sea to the northeast, Panama to the southeast, and the Pacific Ocean to the southwest, as well as Maritime bo ...
to northern and western
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a considerably smaller portion in the Northern Hemisphere. It can also be described as the southern Subregion#Americas, subregion o ...
in highland forest edges. The IOC recognises these subspecies as the species ''Piranga lutea'' (the tooth-billed tanager). ** ''Piranga hepatica testacea'' (highlands of northern
Costa Rica Costa Rica, officially the Republic of Costa Rica, is a country in Central America. It borders Nicaragua to the north, the Caribbean Sea to the northeast, Panama to the southeast, and the Pacific Ocean to the southwest, as well as Maritime bo ...
and eastern
Panama Panama, officially the Republic of Panama, is a country in Latin America at the southern end of Central America, bordering South America. It is bordered by Costa Rica to the west, Colombia to the southeast, the Caribbean Sea to the north, and ...
) ** ''Piranga hepatica faceta'' ( Santa Marta Mountains in northern
Colombia Colombia, officially the Republic of Colombia, is a country primarily located in South America with Insular region of Colombia, insular regions in North America. The Colombian mainland is bordered by the Caribbean Sea to the north, Venezuel ...
, mountains of northern
Venezuela Venezuela, officially the Bolivarian Republic of Venezuela, is a country on the northern coast of South America, consisting of a continental landmass and many Federal Dependencies of Venezuela, islands and islets in the Caribbean Sea. It com ...
and
Trinidad Trinidad is the larger, more populous island of the Republic of Trinidad and Tobago, the country. The island lies off the northeastern coast of Venezuela and sits on the continental shelf of South America. It is the southernmost island in ...
) ** ''Piranga hepatica haemalea'' ( Amazonas and Bolívar in southern Venezuela, the
Guianas The Guianas, also spelled Guyanas or Guayanas, are a geographical region in north-eastern South America. Strictly, the term refers to the three Guianas: Guyana, Suriname, and French Guiana, formerly British Guiana, British, Surinam (Dutch colo ...
, Serra Imeri in northern
Brazil Brazil, officially the Federative Republic of Brazil, is the largest country in South America. It is the world's List of countries and dependencies by area, fifth-largest country by area and the List of countries and dependencies by population ...
) ** ''Piranga hepatica toddi'' (mountains of central Colombia) ** ''Piranga hepatica desidiosa'' (southwestern Colombia) ** ''Piranga hepatica lutea'' (southwestern Colombia and northwestern
Bolivia Bolivia, officially the Plurinational State of Bolivia, is a landlocked country located in central South America. The country features diverse geography, including vast Amazonian plains, tropical lowlands, mountains, the Gran Chaco Province, w ...
) * the ''flava'' group (lowland hepatic tanager), resident in open woods elsewhere in South America. The IOC recognises these subspecies as the species ''Piranga flava'' (the red tanager). ** ''Piranga hepatica macconnelli'' (southern
Guyana Guyana, officially the Co-operative Republic of Guyana, is a country on the northern coast of South America, part of the historic British West Indies. entry "Guyana" Georgetown, Guyana, Georgetown is the capital of Guyana and is also the co ...
, southern
Suriname Suriname, officially the Republic of Suriname, is a country in northern South America, also considered as part of the Caribbean and the West Indies. It is a developing country with a Human Development Index, high level of human development; i ...
and northern Brazil) ** ''Piranga hepatica rosacea'' (eastern Bolivia) ** ''Piranga hepatica saira'' (eastern and southern Brazil) ** ''Piranga hepatica flava'' (southeastern Bolivia,
Paraguay Paraguay, officially the Republic of Paraguay, is a landlocked country in South America. It is bordered by Argentina to the Argentina–Paraguay border, south and southwest, Brazil to the Brazil–Paraguay border, east and northeast, and Boli ...
, northern Argentina and
Uruguay Uruguay, officially the Oriental Republic of Uruguay, is a country in South America. It shares borders with Argentina to its west and southwest and Brazil to its north and northeast, while bordering the Río de la Plata to the south and the A ...
The IUCN follows another taxonomy, recognising ''Piranga hepatica'' (hepatic tanager) and ''Piranga flava'' (red tanager), with the former including the subspecies includes in the ''hepatica'' and ''lutea'' groups.


Description

The species's plumage and vocalizations are similar to other members of the cardinal family. The hepatic tanager is a strongly built songbird up to 20 cm long with a short and quite thick beak and long wings and tail. The male is greyish brick-red above, reddish below with greyish ear-coverts. The female is olive yellow above, yellowish below with soot colored ear coverts. Both sexes have dark beaks and dark legs. The song is a three to four second long beautiful and thrush-like series, but also reminiscent of a black-capped cardinal. The most common sound is rendered in English literature as a clipped "tchup", while in flight it sometimes emits a soft, scream-like "wenk".


Habitat

Hepatic tanager is found in open mountain forests with pine or pine and oak. There it is seen jumping slowly upwards in trees and bushes in search of food, but can also catch flying insects by lunging. It is often found in pairs or small groups, probably families. The nest is placed in a cleft far out on a branch, about 15 to 30 meters above the ground. There she lays three to five eggs.Hepatic Tanager @ allaboutbirds.org
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Behavior

The habits of the hepatic tanager are similar to those of the
western tanager The western tanager (''Piranga ludoviciana''), is a medium-sized Americas, American songbird. Formerly placed in the tanager family (biology), family (Thraupidae), it and other members of its genus are classified in the cardinal family (Cardinali ...
. Its call is a low, dry ''chup'' like the
hermit thrush The hermit thrush (''Catharus guttatus'') is a medium-sized North American thrush. Taxonomy It is not very closely related to the other North American migrant species of ''Catharus'', but rather to the Mexican russet nightingale-thrush. T ...
. Its song is clearer than Thraupidae tanagers and far more similar to the song of the black-headed grosbeak, another member of the Cardinalidae. The flight call is a husky and rising ''weet''.


Feeding

It looks for food in the foliage of trees, moving slowly and methodically; different individuals use different strategies. In summer, the northern form largely eats insects, spiders and some fruit. In Mexico, it has been observed to eat nectar. From
Oaxaca Oaxaca, officially the Free and Sovereign State of Oaxaca, is one of the 32 states that compose the political divisions of Mexico, Federative Entities of the Mexico, United Mexican States. It is divided into municipalities of Oaxaca, 570 munici ...
south, it follows swarms of
army ants The name army ant (or legionary ant or ''marabunta'') is applied to over 200 ant species in different lineages. Because of their aggressive predatory foraging groups, known as "raids", a huge number of ants forage simultaneously over a limited ...
.


References


External links


Hepatic tanager Species Account
– Cornell Lab of Ornithology * from
Paraguay Paraguay, officially the Republic of Paraguay, is a landlocked country in South America. It is bordered by Argentina to the Argentina–Paraguay border, south and southwest, Brazil to the Brazil–Paraguay border, east and northeast, and Boli ...
at * * * * {{Taxonbar, from1=Q3389102 Piranga Birds of Central America Native birds of the Southwestern United States Birds of the Rio Grande valleys Meso-American montane bird species Birds described in 1827 Taxa named by William Swainson