Helix Aperta
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''Cantareus apertus'', commonly known as the green garden snail, is a
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
of air-breathing
land snail A land snail is any of the numerous species of snail that live on land, as opposed to the sea snails and freshwater snails. ''Land snail'' is the common name for terrestrial molluscs, terrestrial gastropod mollusks that have gastropod shell, shel ...
, a
terrestrial Terrestrial refers to things related to land or the planet Earth, as opposed to extraterrestrial. Terrestrial may also refer to: * Terrestrial animal, an animal that lives on land opposed to living in water, or sometimes an animal that lives on o ...
pulmonate Pulmonata or pulmonates is an informal group (previously an order, and before that, a subclass) of snails and slugs characterized by the ability to breathe air, by virtue of having a pallial lung instead of a gill, or gills. The group inclu ...
gastropod Gastropods (), commonly known as slugs and snails, belong to a large Taxonomy (biology), taxonomic class of invertebrates within the phylum Mollusca called Gastropoda (). This class comprises snails and slugs from saltwater, freshwater, and fro ...
mollusc Mollusca is a phylum of protostome, protostomic invertebrate animals, whose members are known as molluscs or mollusks (). Around 76,000 extant taxon, extant species of molluscs are recognized, making it the second-largest animal phylum ...
in the
family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
Helicidae Helicidae is a large, diverse family of western Palaearctic, medium to large-sized, air-breathing land snails, sometimes called the "typical snails." It includes some of the largest European land snails, several species are common in anthropoge ...
, the typical snails.


Distribution

''Cantareus apertus'' is native to
Europe Europe is a continent located entirely in the Northern Hemisphere and mostly in the Eastern Hemisphere. It is bordered by the Arctic Ocean to the north, the Atlantic Ocean to the west, the Mediterranean Sea to the south, and Asia to the east ...
primarily near the
Mediterranean Sea The Mediterranean Sea ( ) is a sea connected to the Atlantic Ocean, surrounded by the Mediterranean basin and almost completely enclosed by land: on the east by the Levant in West Asia, on the north by Anatolia in West Asia and Southern Eur ...
, and also
North Africa North Africa (sometimes Northern Africa) is a region encompassing the northern portion of the African continent. There is no singularly accepted scope for the region. However, it is sometimes defined as stretching from the Atlantic shores of t ...
. Distribution of ''Cantareus apertus'' include: *
France France, officially the French Republic, is a country located primarily in Western Europe. Overseas France, Its overseas regions and territories include French Guiana in South America, Saint Pierre and Miquelon in the Atlantic Ocean#North Atlan ...
west of Rhone estuary It is protected in France, must not be collected for commercial purposes. * Islands in the
Tyrrhenian Sea The Tyrrhenian Sea (, ; or ) , , , , is part of the Mediterranean Sea off the western coast of Italy. It is named for the Tyrrhenians, Tyrrhenian people identified with the Etruscans of Italy. Geography The sea is bounded by the islands of C ...
*
Italy Italy, officially the Italian Republic, is a country in Southern Europe, Southern and Western Europe, Western Europe. It consists of Italian Peninsula, a peninsula that extends into the Mediterranean Sea, with the Alps on its northern land b ...
, Italian peninsula to
Liguria Liguria (; ; , ) is a Regions of Italy, region of north-western Italy; its Capital city, capital is Genoa. Its territory is crossed by the Alps and the Apennine Mountains, Apennines Mountain chain, mountain range and is roughly coextensive with ...
and
Romagna Romagna () is an Italian historical region that approximately corresponds to the south-eastern portion of present-day Emilia-Romagna, in northern Italy. Etymology The name ''Romagna'' originates from the Latin name ''Romania'', which originally ...
*
Ionian islands The Ionian Islands (Modern Greek: , ; Ancient Greek, Katharevousa: , ) are a archipelago, group of islands in the Ionian Sea, west of mainland Greece. They are traditionally called the Heptanese ("Seven Islands"; , ''Heptanēsa'' or , ''Heptanē ...
*
Malta Malta, officially the Republic of Malta, is an island country in Southern Europe located in the Mediterranean Sea, between Sicily and North Africa. It consists of an archipelago south of Italy, east of Tunisia, and north of Libya. The two ...
* Central
Greece Greece, officially the Hellenic Republic, is a country in Southeast Europe. Located on the southern tip of the Balkan peninsula, it shares land borders with Albania to the northwest, North Macedonia and Bulgaria to the north, and Turkey to th ...
* Aegean Islands *
Cyprus Cyprus (), officially the Republic of Cyprus, is an island country in the eastern Mediterranean Sea. Situated in West Asia, its cultural identity and geopolitical orientation are overwhelmingly Southeast European. Cyprus is the List of isl ...
(only one locality) * Mediterranean north Africa In
Salento Salento (; Salentino dialect, Salentino: ''Salentu''; Griko language, Salento Griko: ) is a Cultural area, cultural, List of historical states of Italy, historical, and geographic region at the southern end of the administrative region of Apuli ...
it is known as municeḍḍe and in
Sicily Sicily (Italian language, Italian and ), officially the Sicilian Region (), is an island in the central Mediterranean Sea, south of the Italian Peninsula in continental Europe and is one of the 20 regions of Italy, regions of Italy. With 4. ...
as attuppateḍḍu. It has also been introduced to other areas: * It has become established in
California California () is a U.S. state, state in the Western United States that lies on the West Coast of the United States, Pacific Coast. It borders Oregon to the north, Nevada and Arizona to the east, and shares Mexico–United States border, an ...
and
Louisiana Louisiana ( ; ; ) is a state in the Deep South and South Central regions of the United States. It borders Texas to the west, Arkansas to the north, and Mississippi to the east. Of the 50 U.S. states, it ranks 31st in area and 25 ...
. *
Western Australia Western Australia (WA) is the westernmost state of Australia. It is bounded by the Indian Ocean to the north and west, the Southern Ocean to the south, the Northern Territory to the north-east, and South Australia to the south-east. Western Aust ...
– nonindigenous This species is already established in the
United States The United States of America (USA), also known as the United States (U.S.) or America, is a country primarily located in North America. It is a federal republic of 50 U.S. state, states and a federal capital district, Washington, D.C. The 48 ...
, and is considered to represent a potentially serious threat as a pest, an
invasive species An invasive species is an introduced species that harms its new environment. Invasive species adversely affect habitats and bioregions, causing ecological, environmental, and/or economic damage. The term can also be used for native spec ...
which could negatively affect agriculture, natural ecosystems, human health or commerce. Therefore, it has been suggested that this species be given top national
quarantine A quarantine is a restriction on the movement of people, animals, and goods which is intended to prevent the spread of disease or pests. It is often used in connection to disease and illness, preventing the movement of those who may have bee ...
significance in the USA.Cowie R. H., Dillon R. T., Robinson D. G. & Smith J. W. (2009). "Alien non-marine snails and slugs of priority quarantine importance in the United States: A preliminary risk assessment". ''American Malacological Bulletin'' 27: 113–132
PDF
.


Description

The
shell Shell may refer to: Architecture and design * Shell (structure), a thin structure ** Concrete shell, a thin shell of concrete, usually with no interior columns or exterior buttresses Science Biology * Seashell, a hard outer layer of a marine ani ...
has a
periostracum The periostracum ( ) is a thin, organic coating (or "skin") that is the outermost layer of the shell of many shelled animals, including molluscs and brachiopods. Among molluscs, it is primarily seen in snails and clams, i.e. in gastropods an ...
which is olive green in colour. The
last whorl The body whorl is part of the morphology of the shell in those gastropod mollusks that possess a coiled shell. The term is also sometimes used in a similar way to describe the shell of a cephalopod mollusk. In gastropods In gastropods, the b ...
is much larger than the others. The width of the shell is 22–28 mm.; the height of the shell is also 22–28 mm.


Ecology

''Cantareus apertus'' inhabits Mediterranean shrublands, near cultivated fields, gardens. In
Gavdos Gavdos ( ) is the southernmost Greek island, located to the south of its much larger neighbour, Crete, of which it is administratively a part, in the regional unit of Chania. It forms a community with surrounding islets and was part of the forme ...
(Greece) also in woodland spreading on recently abandoned cultivated fields, more rarely in natural habitats. In
Crete Crete ( ; , Modern Greek, Modern: , Ancient Greek, Ancient: ) is the largest and most populous of the Greek islands, the List of islands by area, 88th largest island in the world and the List of islands in the Mediterranean#By area, fifth la ...
this species is active for 3–4 months after the first rainfalls in October. It aestivates buried relatively deep in the soil. In hot, dry weather, it burrows three to six inches into the ground and becomes dormant until rain softens the soil. A white convex
epiphragm An epiphragm (from the Ancient Greek ἐπί, '' epi '' " upon, on, over " and φράγμα, '' -phrágma '' "fence") is a temporary structure which can be created by many species of shelled, air-breathing land snails, terrestrial pulmonate gas ...
is created for
aestivation Aestivation ( (summer); also spelled estivation in American English) is a state of animal dormancy, similar to hibernation, although taking place in the summer rather than the winter. Aestivation is characterized by inactivity and a lowered m ...
. This species of snail makes and uses
love darts A love dart (also known as a gypsobelum, shooting darts, or just as darts) is a sharp, calcium carbonate, calcareous or chitinous Dart (missile), dart which some hermaphroditic land snails and slugs create. Love darts are both formed and stor ...
.


References

This article incorporates public domain text from the reference."Species summary for ''Helix apertus''"
AnimalBase AnimalBase is a project brought to life in 2004 and is maintained by the University of Göttingen, Germany. The goal of the AnimalBase project is to digitize early zoological literature, provide copyright-free open access to zoological works, and p ...
, last modified 24 January 2010, accessed 13 October 2010.


External links

* {{Taxonbar, from=Q1130456 Gastropods described in 1778 Taxa named by Ignaz von Born Helicidae