Helicops Angulatus
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The brown-banded water snake (''Helicops angulatus'') is a
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
of aquatic
snake Snakes are elongated limbless reptiles of the suborder Serpentes (). Cladistically squamates, snakes are ectothermic, amniote vertebrates covered in overlapping scales much like other members of the group. Many species of snakes have s ...
found in tropical
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a considerably smaller portion in the Northern Hemisphere. It can also be described as the southern Subregion#Americas, subregion o ...
and
Trinidad and Tobago Trinidad and Tobago, officially the Republic of Trinidad and Tobago, is the southernmost island country in the Caribbean, comprising the main islands of Trinidad and Tobago, along with several List of islands of Trinidad and Tobago, smaller i ...
. It is also known as the water mapepire.


Description

The brown-banded water snake grows to a maximum total length (including tail) of 78 cm (31 inches).
Dorsally Standard anatomical terms of location are used to describe unambiguously the anatomy of humans and other animals. The terms, typically derived from Latin or Greek roots, describe something in its standard anatomical position. This position provi ...
, it is olive or gray-brown, with dark brown, black-edged crossbands, which narrow at the sides, and are usually confluent with the black crossbands of the belly. There is a large dark rhomboid on the nape. Ventrally, it is yellowish (in alcohol) with black crossbands or black spots. The
dorsal scales In snakes, the dorsal scales are the longitudinal series of plates that encircle the body, but do not include the ventral scales. Campbell JA, Lamar WW (2004). ''The Venomous Reptiles of the Western Hemisphere''. Ithaca and London: Comstock Publis ...
are strongly keeled, even on the occiput and nape, and are arranged in 19 rows. Ventrals are 102–130 in number, the anal scale is divided, and the 61-94
subcaudals In snakes, the subcaudal scales are the enlarged plates on the underside of the tail. Wright AH, Wright AA (1957). ''Handbook of Snakes of the United States and Canada''. Comstock Publishing Associates, a Division of Cornell University Press. (7t ...
are paired and keeled.


Habitat and diet

''H. angulatus'' lives in
fresh Fresh may refer to: Arts, entertainment, and media Films and television * ''Fresh'' (1994 film), a crime film * ''Fresh'' (2009 film), a documentary film on sustainable agriculture * ''Fresh'' (2022 film), a thriller film *''Fresh with the Aust ...
and
brackish Brackish water, sometimes termed brack water, is water occurring in a natural environment that has more salinity than freshwater, but not as much as seawater. It may result from mixing seawater (salt water) and fresh water together, as in estuari ...
water, where it feeds on
fish A fish (: fish or fishes) is an aquatic animal, aquatic, Anamniotes, anamniotic, gill-bearing vertebrate animal with swimming fish fin, fins and craniate, a hard skull, but lacking limb (anatomy), limbs with digit (anatomy), digits. Fish can ...
(such as freshwater
eel Eels are ray-finned fish belonging to the order Anguilliformes (), which consists of eight suborders, 20 families, 164 genera, and about 1000 species. Eels undergo considerable development from the early larval stage to the eventual adult stage ...
s) and possibly also
frog A frog is any member of a diverse and largely semiaquatic group of short-bodied, tailless amphibian vertebrates composing the order (biology), order Anura (coming from the Ancient Greek , literally 'without tail'). Frog species with rough ski ...
s and their
egg An egg is an organic vessel grown by an animal to carry a possibly fertilized egg cell (a zygote) and to incubate from it an embryo within the egg until the embryo has become an animal fetus that can survive on its own, at which point the ...
s,
tadpole A tadpole or polliwog (also spelled pollywog) is the Larva, larval stage in the biological life cycle of an amphibian. Most tadpoles are fully Aquatic animal, aquatic, though some species of amphibians have tadpoles that are terrestrial animal, ...
s,
lizard Lizard is the common name used for all Squamata, squamate reptiles other than snakes (and to a lesser extent amphisbaenians), encompassing over 7,000 species, ranging across all continents except Antarctica, as well as most Island#Oceanic isla ...
s,
earthworm An earthworm is a soil-dwelling terrestrial invertebrate that belongs to the phylum Annelida. The term is the common name for the largest members of the class (or subclass, depending on the author) Oligochaeta. In classical systems, they we ...
s, and
carrion Carrion (), also known as a carcass, is the decaying flesh of dead animals. Overview Carrion is an important food source for large carnivores and omnivores in most ecosystems. Examples of carrion-eaters (or scavengers) include crows, vultures ...
.


Reproduction

''H. angulatus'' has been reported to be "facultatively viviparous".


Venom

''H. angulatus'' is a non-front-fanged colubroid (NFFC), venomous snake, its venom causes rapid death in mice with an injection of 0.4 mg/kg (intraperitoneally). There is an urgent need for training of the medical team in the snake identification, clinical management of snakebite, and the existence of a human-snake conflict involving NFFC species in Bolivia.


References


Further reading

* * 1982. ''Snakes of South America''. Hong Kong: T.F.H. Publications.189 pp. . (''Helicops angulatus'', p. 99). * Linnaeus, C. 1758. ''Systema naturæ per regna tria naturæ, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Editio Decima, Reformata''. Stockholm: L. Salvius. 824 pp. (''Coluber angulatus'', new species, p. 217). (in
Latin Latin ( or ) is a classical language belonging to the Italic languages, Italic branch of the Indo-European languages. Latin was originally spoken by the Latins (Italic tribe), Latins in Latium (now known as Lazio), the lower Tiber area aroun ...
). * Helicops Reptiles of Trinidad and Tobago Reptiles of Guyana Reptiles described in 1758 Snakes of South America Taxa named by Carl Linnaeus {{Dipsadinae-stub