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''Helcogramma'' is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial n ...
in the
triplefin Threefin or triplefin blennies are blenniiforms, small percomorph marine fish of the family Tripterygiidae. Found in tropical and temperate waters of the Atlantic, Pacific and Indian Oceans, the family contains about 150 species in 30 genera ...
family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
Tripterygiidae. The blennies in the genus ''Helcogramma'' are found throughout the
Indo-Pacific The Indo-Pacific is a vast biogeographic region of Earth. In a narrow sense, sometimes known as the Indo-West Pacific or Indo-Pacific Asia, it comprises the tropical waters of the Indian Ocean, the western and central Pacific Ocean, and the ...
and in the South Atlantic Ocean off the islands of
St Helena Saint Helena () is a British overseas territory located in the South Atlantic Ocean. It is a remote volcanic tropical island west of the coast of south-western Africa, and east of Rio de Janeiro in South America. It is one of three constitu ...
and
Ascension Ascension or ascending may refer to: Religion * "Ascension", "Assumption", or "Translation", the belief in some religions that some individuals have ascended into Heaven without dying first * Ascension of Jesus * Feast of the Ascension (Ascen ...
.


Characteristics

The fishes in the genus ''Helcogramma'' are characterised by the possession of a continuous lateral line which bends down towards the operculum, this lateral line comprises 7–39 tubular pored scales. There are three spines in the first
dorsal fin A dorsal fin is a fin located on the back of most marine and freshwater vertebrates within various taxa of the animal kingdom. Many species of animals possessing dorsal fins are not particularly closely related to each other, though through c ...
and the
anal fin Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as s ...
has one spine, the
pelvic fin Pelvic fins or ventral fins are paired fins located on the ventral surface of fish. The paired pelvic fins are homologous to the hindlimbs of tetrapods. Structure and function Structure In actinopterygians, the pelvic fin consists of two e ...
has a single spine and two rays. The head is scaleless and normally the bases of the dorsal and anal fins have a scaleless strip. There are five bones in the joint between the
caudal fin Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as s ...
and the
vertebrae The spinal column, a defining synapomorphy shared by nearly all vertebrates, Hagfish are believed to have secondarily lost their spinal column is a moderately flexible series of vertebrae (singular vertebra), each constituting a characteristi ...
, the hypurals.


Species

There are currently 41 recognized species in this genus: * ''
Helcogramma albimacula ''Helcogramma albimacula'' is a species of triplefin blenny in the genus ''Helcogramma ''Helcogramma'' is a genus in the triplefin family (biology), family Tripterygiidae. The blennies in the genus ''Helcogramma'' are found throughout the Indo ...
''
J. T. Williams J. T. Williams was an American football coach. He was the fifth head football coach at Kentucky State University in Frankfort, Kentucky Frankfort is the capital city of the Commonwealth of Kentucky, United States, and the seat of Frank ...
&
Howe Howe may refer to: People and fictional characters * Howe (surname), including a list of people and fictional characters * Howe Browne, 2nd Marquess of Sligo (1788–1845), Irish peer and colonial governor Titles * Earl Howe, two titles, an exti ...
, 2003
(Whitespot triplefin) * ''
Helcogramma alkamr ''Helcogramma alkamr'' is a species of triplefin blenny in the genus ''Helcogramma''. It was described by Wouter Holleman in 2007. This species is found in the western Indian Ocean from the Comoros to the Seychelles, Mauritius and St Brandon. Th ...
''
Holleman Holleman is a surname. Notable people with the surname include: *Boyce Holleman (1924–2003), American war veteran, attorney, politician, and actor * Frederik David Holleman (1887–1958), Dutch and South African professor, ethnologist, and legal ...
, 2007
* ''
Helcogramma aquila ''Helcogramma aquila'', the darktail triplefin or blacktail triplefin, is a species of triplefin blenny in the genus ''Helcogramma''. It was described by Jeffrey T. Williams and Carolyn J. McCormick in 1990. This species occurs in the western Pa ...
''
J. T. Williams J. T. Williams was an American football coach. He was the fifth head football coach at Kentucky State University in Frankfort, Kentucky Frankfort is the capital city of the Commonwealth of Kentucky, United States, and the seat of Frank ...
& C. J. McCormick, 1990
(Darktail triplefin) * ''
Helcogramma ascensionis ''Helcogramma ascensionis'', the Ascension triplefin, is a species of triplefin blenny in the genus ''Helcogramma''. it was described by Roger Lubbock in 1980. It has been recorded at depths of in tidal pools and rocky reefs around the islands o ...
''
Lubbock Lubbock ( ) is the List of cities in Texas by population, 10th-most populous city in the U.S. state of Texas and the county seat, seat of government of Lubbock County, Texas, Lubbock County. With a population of 260,993 in 2021, the city is also ...
, 1980
(Ascension triplefin) * ''
Helcogramma atauroensis ''Helcogramma atauroensis'', the red-anal triplefin, is a species of triplefin blenny in the genus ''Helcogramma ''Helcogramma'' is a genus in the triplefin family (biology), family Tripterygiidae. The blennies in the genus ''Helcogramma'' are ...
'' Fricke & Erdmann, 2017 * ''
Helcogramma billi ''Helcogramma billi'' is a species of triplefin blenny in the genus ''Helcogramma''. It was described by Patricia E. Hadley Hansen in 1986, and the specific name honours the American ichthyologist Bill Smith-Vaniz, who collected the types Hadl ...
'' P. E. Hadley, 1986 * ''
Helcogramma capidata The hooded triplefin (''Helcogramma capidata'') is a species of triplefin blenny in the genus ''Helcogramma''. It was described by Richard Rosenblatt in 1960. This species is widespread in the western Pacific Ocean from the Mariana Islands Th ...
''
Rosenblatt Rosenblatt is a surname of German and Jewish origin, meaning " rose leaf". People with this surname include: *Albert Rosenblatt (born 1936), New York Court of Appeals judge *Dana Rosenblatt, known as "Dangerous" (born 1972), American boxer *Elie R ...
, 1960
(Hooded triplefin) * ''
Helcogramma cerasina ''Helcogramma cerasina'' is a species of triplefin blenny in the genus ''Helcogramma ''Helcogramma'' is a genus in the triplefin family (biology), family Tripterygiidae. The blennies in the genus ''Helcogramma'' are found throughout the Indo-P ...
''
J. T. Williams J. T. Williams was an American football coach. He was the fifth head football coach at Kentucky State University in Frankfort, Kentucky Frankfort is the capital city of the Commonwealth of Kentucky, United States, and the seat of Frank ...
&
Howe Howe may refer to: People and fictional characters * Howe (surname), including a list of people and fictional characters * Howe Browne, 2nd Marquess of Sligo (1788–1845), Irish peer and colonial governor Titles * Earl Howe, two titles, an exti ...
, 2003
* ''
Helcogramma chica The little hooded triplefin (''Helcogramma chica'') is a species of triplefin blenny in the genus ''Helcogramma''. It was Species description, described by Richard Rosenblatt in 1960. This species is found in the Indo-Pacific from the Cocos (Keel ...
''
Rosenblatt Rosenblatt is a surname of German and Jewish origin, meaning " rose leaf". People with this surname include: *Albert Rosenblatt (born 1936), New York Court of Appeals judge *Dana Rosenblatt, known as "Dangerous" (born 1972), American boxer *Elie R ...
, 1960
(Little hooded triplefin) * '' Helcogramma decurrens''
McCulloch McCulloch is a Scottish surname. It's a variation of the Northern Irish surname McCullough. It's commonly found in Galloway. Notable people with the surname include: *Alan McCulloch (politician), New Zealand politician *Alan McLeod McCulloch (1 ...
& Waite, 1918
(Black-throated triplefin) * '' Helcogramma desa''
J. T. Williams J. T. Williams was an American football coach. He was the fifth head football coach at Kentucky State University in Frankfort, Kentucky Frankfort is the capital city of the Commonwealth of Kentucky, United States, and the seat of Frank ...
&
Howe Howe may refer to: People and fictional characters * Howe (surname), including a list of people and fictional characters * Howe Browne, 2nd Marquess of Sligo (1788–1845), Irish peer and colonial governor Titles * Earl Howe, two titles, an exti ...
, 2003
(Neglected triplefin) * '' Helcogramma ellioti'' ( Herre, 1944) * '' Helcogramma ememes''
Holleman Holleman is a surname. Notable people with the surname include: *Boyce Holleman (1924–2003), American war veteran, attorney, politician, and actor * Frederik David Holleman (1887–1958), Dutch and South African professor, ethnologist, and legal ...
, 2007
* '' Helcogramma fuscipectoris'' ( Fowler, 1946) (Fourspot triplefin) * '' Helcogramma fuscopinna''
Holleman Holleman is a surname. Notable people with the surname include: *Boyce Holleman (1924–2003), American war veteran, attorney, politician, and actor * Frederik David Holleman (1887–1958), Dutch and South African professor, ethnologist, and legal ...
, 1982
(Blackfin triplefin) * '' Helcogramma gymnauchen'' ( M. C. W. Weber, 1909) (Red-finned triplefin) * '' Helcogramma hudsoni'' ( D. S. Jordan & Seale, 1906) (Hudson's triplefin) * '' Helcogramma inclinata'' ( Fowler, 1946) (Triangle triplefin) * '' Helcogramma ishigakiensis'' Aoyagi, 1954 * '' Helcogramma kranos'' R. Fricke, 1997 (Helmet triplefin) * ''
Helcogramma lacuna ''Helcogramma lacuna'', the cavern triplefin, is a species of triplefin blenny in the genus ''Helcogramma''. It was described by Jeffrey T. Williams and Jeffery C. Howe in 2003. This species has only been recorded in the Similan Islands in the ...
''
J. T. Williams J. T. Williams was an American football coach. He was the fifth head football coach at Kentucky State University in Frankfort, Kentucky Frankfort is the capital city of the Commonwealth of Kentucky, United States, and the seat of Frank ...
&
Howe Howe may refer to: People and fictional characters * Howe (surname), including a list of people and fictional characters * Howe Browne, 2nd Marquess of Sligo (1788–1845), Irish peer and colonial governor Titles * Earl Howe, two titles, an exti ...
, 2003
* '' Helcogramma larvata'' R. Fricke & J. E. Randall, 1992 * '' Helcogramma maldivensis'' R. Fricke & J. E. Randall, 1992 * '' Helcogramma microstigma''
Holleman Holleman is a surname. Notable people with the surname include: *Boyce Holleman (1924–2003), American war veteran, attorney, politician, and actor * Frederik David Holleman (1887–1958), Dutch and South African professor, ethnologist, and legal ...
, 2006
* '' Helcogramma nesion''
J. T. Williams J. T. Williams was an American football coach. He was the fifth head football coach at Kentucky State University in Frankfort, Kentucky Frankfort is the capital city of the Commonwealth of Kentucky, United States, and the seat of Frank ...
&
Howe Howe may refer to: People and fictional characters * Howe (surname), including a list of people and fictional characters * Howe Browne, 2nd Marquess of Sligo (1788–1845), Irish peer and colonial governor Titles * Earl Howe, two titles, an exti ...
, 2003
* '' Helcogramma nigra''
J. T. Williams J. T. Williams was an American football coach. He was the fifth head football coach at Kentucky State University in Frankfort, Kentucky Frankfort is the capital city of the Commonwealth of Kentucky, United States, and the seat of Frank ...
&
Howe Howe may refer to: People and fictional characters * Howe (surname), including a list of people and fictional characters * Howe Browne, 2nd Marquess of Sligo (1788–1845), Irish peer and colonial governor Titles * Earl Howe, two titles, an exti ...
, 2003
(Rotuma triplefin) * '' Helcogramma novaecaledoniae'' R. Fricke, 1994 (New Caledonian triplefin) * ''
Helcogramma obtusirostris ''Helcogramma obtusirostris'', the hotlips triplefin or shortsnout triplefin, is a triplefin blenny of the family Tripterygiidae, found from the Red Sea south to South Africa and east to the western Pacific, and the southeast Atlantic, aroun ...
'' ( Klunzinger, 1871) (Hotlips triplefin) * '' Helcogramma randalli''
J. T. Williams J. T. Williams was an American football coach. He was the fifth head football coach at Kentucky State University in Frankfort, Kentucky Frankfort is the capital city of the Commonwealth of Kentucky, United States, and the seat of Frank ...
&
Howe Howe may refer to: People and fictional characters * Howe (surname), including a list of people and fictional characters * Howe Browne, 2nd Marquess of Sligo (1788–1845), Irish peer and colonial governor Titles * Earl Howe, two titles, an exti ...
, 2003
(Randall's triplefin) * '' Helcogramma rharhabe''
Holleman Holleman is a surname. Notable people with the surname include: *Boyce Holleman (1924–2003), American war veteran, attorney, politician, and actor * Frederik David Holleman (1887–1958), Dutch and South African professor, ethnologist, and legal ...
, 2007
* '' Helcogramma rhinoceros'' P. E. Hadley, 1986 (Rhinocerus triplefin) * '' Helcogramma rosea''
Holleman Holleman is a surname. Notable people with the surname include: *Boyce Holleman (1924–2003), American war veteran, attorney, politician, and actor * Frederik David Holleman (1887–1958), Dutch and South African professor, ethnologist, and legal ...
, 2006
(Rosy triplefin) * '' Helcogramma serendip''
Holleman Holleman is a surname. Notable people with the surname include: *Boyce Holleman (1924–2003), American war veteran, attorney, politician, and actor * Frederik David Holleman (1887–1958), Dutch and South African professor, ethnologist, and legal ...
, 2007
* '' Helcogramma shinglensis'' Lal Mohan, 1971 * '' Helcogramma solorensis'' R. Fricke, 1997 (Solor triplefin) * '' Helcogramma springeri'' P. E. Hadley, 1986 (Springer's triplefin) * '' Helcogramma steinitzi'' E. Clark, 1980 (Red triplefin) * '' Helcogramma striata'' P. E. Hadley, 1986 (Tropical striped triplefin) * '' Helcogramma trigloides'' (
Bleeker Bleeker is a Dutch occupational surname. Bleeker is an old spelling of ''(linnen)bleker'' ("linen bleacher").Helcogramma vulcana'' J. E. Randall & E. Clark, 1993 (Volcano triplefin) * '' Helcogramma williamsi'' M. C. Chiang & I. S. Chen, 2012


References

{{Taxonbar, from=Q628986 Tripterygiidae Taxa named by Allan Riverstone McCulloch Taxa named by Edgar Ravenswood Waite