Hederelloids
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Hederellids are
extinct Extinction is the termination of an organism by the death of its Endling, last member. A taxon may become Functional extinction, functionally extinct before the death of its last member if it loses the capacity to Reproduction, reproduce and ...
colonial
animal Animals are multicellular, eukaryotic organisms in the Biology, biological Kingdom (biology), kingdom Animalia (). With few exceptions, animals heterotroph, consume organic material, Cellular respiration#Aerobic respiration, breathe oxygen, ...
s with calcitic tubular branching
exoskeleton An exoskeleton () . is a skeleton that is on the exterior of an animal in the form of hardened integument, which both supports the body's shape and protects the internal organs, in contrast to an internal endoskeleton (e.g. human skeleton, that ...
s. They range from the
Silurian The Silurian ( ) is a geologic period and system spanning 23.5 million years from the end of the Ordovician Period, at million years ago ( Mya), to the beginning of the Devonian Period, Mya. The Silurian is the third and shortest period of t ...
to the
Permian The Permian ( ) is a geologic period and System (stratigraphy), stratigraphic system which spans 47 million years, from the end of the Carboniferous Period million years ago (Mya), to the beginning of the Triassic Period 251.902 Mya. It is the s ...
and were most common in the
Devonian The Devonian ( ) is a period (geology), geologic period and system (stratigraphy), system of the Paleozoic era (geology), era during the Phanerozoic eon (geology), eon, spanning 60.3 million years from the end of the preceding Silurian per ...
period. They are more properly known as hederelloids because they were originally defined as a suborder by Bassler, who described about 130 species. Although they have traditionally been considered
bryozoans Bryozoa (also known as the Polyzoa, Ectoprocta or commonly as moss animals) are a phylum of simple, aquatic animal, aquatic invertebrate animals, nearly all living in sedentary Colony (biology), colonies. Typically about long, they have a spe ...
, they are clearly not because of their branching patterns, lack of an astogenetic gradient, skeletal microstructure, and wide range in tube diameters. Work continues on assessing the true affinities of hederelloids, but they appear to be most closely related to
phoronid Phoronids (Taxonomy (biology), taxonomic name Phoronida, sometimes called horseshoe worms) are a small phylum of marine animals that Filter feeder, filter-feed with a lophophore (a "crown" of tentacles), and build upright tubes of chitin to suppo ...
s and other
lophophorate The Lophophorata (also called Tentaculata; not to be confused with Tentaculata Eschscholtz 1825, a class within the Ctenophora) are a Lophotrochozoan clade consisting of the Brachiozoa and the Bryozoa. They have a lophophore. Molecular phyloge ...
s.


Classification

*Family Hederellidae **Genus '' Diversipora'' **Genus '' Hederella'' *Family Reptariidae **Genus '' Cystoporella'' **Genus '' Hederopsis'' **Genus '' Hernodia'' File:HederellaOH3.jpg, Hederelloids encrusting a
brachiopod Brachiopods (), phylum (biology), phylum Brachiopoda, are a phylum of animals that have hard "valves" (shells) on the upper and lower surfaces, unlike the left and right arrangement in bivalve molluscs. Brachiopod valves are hinged at the rear e ...
from the Devonian of Ohio File:HederelloidSEM.jpg,
SEM SEM or Sem can refer to: Computing * Search engine marketing, promoting websites by increasing their visibility in search engine results * Security event manager, a security log tool used on data networks Economics and management * Stock Ex ...
image of a hederelloid from the Devonian of Michigan (largest tube diameter is 0.75 mm)


References


Further reading

* * Enigmatic prehistoric animal taxa Carboniferous invertebrates Devonian animals Late Devonian animals Permian animals Permian extinctions Enigmatic protostome taxa Silurian animals Silurian first appearances {{paleo-invertebrate-stub