Haskap Berries
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''Lonicera caerulea'', commonly known as honeyberry or by various honeysuckle names, is a non-
climbing Climbing is the activity of using one's hands, feet, or other parts of the body to ascend a steep topographical object that can range from the world's tallest mountains (e.g. the eight thousanders) to small boulders. Climbing is done for locom ...
honeysuckle Honeysuckles are arching shrubs or Vine#Twining vines, twining vines in the genus ''Lonicera'' () of the family Caprifoliaceae. The genus includes 158 species native to northern latitudes in North America, Eurasia, and North Africa. Widely kno ...
native throughout the cool temperate
Northern Hemisphere The Northern Hemisphere is the half of Earth that is north of the equator. For other planets in the Solar System, north is defined by humans as being in the same celestial sphere, celestial hemisphere relative to the invariable plane of the Solar ...
regions of North America, Europe, and Asia. The plant or its fruit has also come to be called haskap, derived from its name in
the language "The Language" is a song by Canadian rapper Drake from his third studio album ''Nothing Was the Same'' (2013). "The Language" was produced by frequent collaborator Boi-1da, along with additional production by Allen Ritter and Vinylz. It also fea ...
of the native
Ainu people The Ainu are an Indigenous peoples, indigenous ethnic group who reside in northern Japan and southeastern Russia, including Hokkaido and the Tōhoku region of Honshu, as well as the land surrounding the Sea of Okhotsk, such as Sakhalin, the Ku ...
of
Hokkaido, Japan is the second-largest island of Japan and comprises the largest and northernmost prefecture, making up its own region. The Tsugaru Strait separates Hokkaidō from Honshu; the two islands are connected by railway via the Seikan Tunnel. The ...
.


Description

Haskap is a
deciduous In the fields of horticulture and botany, the term deciduous () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed Leaf, leaves, usually in the autumn; to the shedding of petals, aft ...
shrub A shrub or bush is a small to medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees by their multiple ...
growing to tall. The
leaves A leaf (: leaves) is a principal appendage of the stem of a vascular plant, usually borne laterally above ground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, stem, ...
are opposite, oval, long and broad, greyish green, with a slightly waxy texture. The
flower Flowers, also known as blooms and blossoms, are the reproductive structures of flowering plants ( angiosperms). Typically, they are structured in four circular levels, called whorls, around the end of a stalk. These whorls include: calyx, m ...
s are yellowish-white, 12–16 mm long, with five equal lobes; they are produced in pairs on the shoots. The
fruit In botany, a fruit is the seed-bearing structure in flowering plants (angiosperms) that is formed from the ovary after flowering. Fruits are the means by which angiosperms disseminate their seeds. Edible fruits in particular have long propaga ...
is an edible, blue
berry A berry is a small, pulpy, and often edible fruit. Typically, berries are juicy, rounded, brightly colored, sweet, sour or tart, and do not have a stone or pit although many pips or seeds may be present. Common examples of berries in the cul ...
, somewhat
cylindrical A cylinder () has traditionally been a Solid geometry, three-dimensional solid, one of the most basic of curvilinear geometric shapes. In elementary geometry, it is considered a Prism (geometry), prism with a circle as its base. A cylinder may ...
in shape weighing , and about in diameter. The plant is
winter-hardy Hardiness of plants describes their ability to survive adverse growing conditions. It is usually limited to discussions of climatic adversity. Thus a plant's ability to tolerate cold, heat, drought, flooding, or wind are typically considered measu ...
and can tolerate temperatures below . Its flowers are frost-tolerant. Fruits mature early and are high in
vitamin C Vitamin C (also known as ascorbic acid and ascorbate) is a water-soluble vitamin found in citrus and other fruits, berries and vegetables. It is also a generic prescription medication and in some countries is sold as a non-prescription di ...
. Haskap
cultivars A cultivar is a kind of cultivated plant that people have selected for desired traits and which retains those traits when propagated. Methods used to propagate cultivars include division, root and stem cuttings, offsets, grafting, tissue cult ...
can survive a large range of
soil acidity Soil pH is a measure of the acidity or basicity (alkalinity) of a soil. Soil pH is a key characteristic that can be used to make informative analysis both qualitative and quantitatively regarding soil characteristics. pH is defined as the nega ...
from 3.9-7.7 (optimum 5.5-6.5), requiring high organic matter, well drained soils, and plentiful sunlight for optimum
productivity Productivity is the efficiency of production of goods or services expressed by some measure. Measurements of productivity are often expressed as a ratio of an aggregate output to a single input or an aggregate input used in a production proce ...
. ''Lonicera caerulea'' plants are more tolerant of wet conditions than most fruit species. Chromosome count is 2n = 18.


Distribution and habitat

The species is circumpolar, primarily found in or near wetlands of
boreal forests Taiga or tayga ( ; , ), also known as boreal forest or snow forest, is a biome characterized by coniferous forests consisting mostly of pines, spruces, and larches. The taiga, or boreal forest, is the world's largest land biome. In North Ame ...
in heavy
peat Peat is an accumulation of partially Decomposition, decayed vegetation or organic matter. It is unique to natural areas called peatlands, bogs, mires, Moorland, moors, or muskegs. ''Sphagnum'' moss, also called peat moss, is one of the most ...
soils of North America, Europe, and Asia. It also can be found in high-
calcium Calcium is a chemical element; it has symbol Ca and atomic number 20. As an alkaline earth metal, calcium is a reactive metal that forms a dark oxide-nitride layer when exposed to air. Its physical and chemical properties are most similar to it ...
soils, in mountains, and along the coasts of northeastern Asia and northwestern North America. Different
varieties Variety may refer to: Arts and entertainment Entertainment formats * Variety (radio) * Variety show, in theater and television Films * ''Variety'' (1925 film), a German silent film directed by Ewald Andre Dupont * ''Variety'' (1935 film), ...
are distributed across central and northern Canada, northern United States, northern and eastern Europe,
Siberia Siberia ( ; , ) is an extensive geographical region comprising all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has formed a part of the sovereign territory of Russia and its predecessor states ...
, middle Asia, and northeastern China.


Classification

The classification within the species is not settled. One classification uses nine botanical varieties: *''Lonicera caerulea'' var. ''altaica''. Northern Asia. *''Lonicera caerulea'' var. ''caerulea''. Europe. *''Lonicera caerulea'' var. ''cauriana''. Western North America. *''Lonicera caerulea'' var. ''dependens''. Central Asia. *''Lonicera caerulea'' var. ''edulis'', synonym: ''L. edulis''. Eastern Asia. *''Lonicera caerulea'' var. ''emphyllocalyx'' (also known as haskap). Eastern Asia. *''Lonicera caerulea'' var. ''kamtschatica''. Northeastern Asia. *''Lonicera caerulea'' var. ''pallasii''. Northern Asia, northeastern Europe. * ''Lonicera caerulea'' var. ''villosa''. Eastern North America.


Cultivated varieties

Improved cultivars include: * 'Aurora' * 'Boreal Beauty' * 'Boreal Beast' * 'Boreal Blizzard' * 'Honeybee' * 'Wojtek' * 'Berry Blue' * 'Indigo Gem' * 'Indigo Treat' * 'Indigo Yum' * 'Tundra' * 'Borealis' * 'Atlaj' * 'Nimfa' * 'Polar Jewel' * 'Sinoglaska' According to research at the
University of Saskatchewan The University of Saskatchewan (U of S, or USask) is a Universities in Canada, Canadian public university, public research university, founded on March 19, 1907, and located on the east side of the South Saskatchewan River in Saskatoon, Saskatch ...
, each variety can be distinguished by the size of berries, taste, and bush dimensions.


Common names

''Lonicera caerulea'' is known by several common names: * Haskap: name of the
Ainu language Ainu (, ), or more precisely Hokkaido Ainu (), is a language spoken by a few elderly members of the Ainu people on the northern Japanese island of Hokkaido. It is a member of the Ainu language family, itself considered a language family isola ...
in northern Japan * Blue honeysuckle: descriptive translation from Russian origin * Honeyberry: common in North America * Swamp fly honeysuckle: coined by
botanist Botany, also called plant science, is the branch of natural science and biology studying plants, especially Plant anatomy, their anatomy, Plant taxonomy, taxonomy, and Plant ecology, ecology. A botanist or plant scientist is a scientist who s ...
s who found it growing wild in
swamp A swamp is a forested wetland.Keddy, P.A. 2010. Wetland Ecology: Principles and Conservation (2nd edition). Cambridge University Press, Cambridge, UK. 497 p. Swamps are considered to be transition zones because both land and water play a role in ...
y areas of Canada


Cultivation

The indigenous peoples of eastern Russia, northern Japan and northern China have long harvested the wild berries, but cultivation efforts are relatively recent, beginning in the Soviet Union in the 1950s. Research into commercial cultivation continued in
Hokkaido, Japan is the second-largest island of Japan and comprises the largest and northernmost prefecture, making up its own region. The Tsugaru Strait separates Hokkaidō from Honshu; the two islands are connected by railway via the Seikan Tunnel. The ...
in the 1970s. The plant is mostly unknown in the Western world, even while some varieties grow in northern Canada and northern United States. Haskap variety ''edulis'' has been used frequently in breeding efforts, but other varieties have been bred with it to increase productivity and flavor. In several haskap breeding programs, the variety ''emphyllocalyx'' has been the dominant one used. In recent years, a new programme has aimed at growing honeyberries commercially in
Scotland Scotland is a Countries of the United Kingdom, country that is part of the United Kingdom. It contains nearly one-third of the United Kingdom's land area, consisting of the northern part of the island of Great Britain and more than 790 adjac ...
(with a similar climate to Japan) at farms in Duns, Angus, Tayside, Perth and Fife.


Disease

This plant is not affected by many pests and diseases.
Powdery mildew Powdery mildew is a fungus, fungal disease that affects a wide range of plants. Powdery mildew diseases are caused by many different species of Ascomycota, ascomycete fungi in the order Erysiphales. Powdery mildew is one of the easier plant disea ...
is one disease documented to affect ''Lonicera caerulea'', usually after fruit maturity in mid– to late summer. When the plant is affected, it is common for the leaves to turn white, with brown patches eventually developing.


Harvest and uses

Honeysuckle is harvested in late spring or early summer two weeks before strawberries for Russian type varieties, with Japanese types ripening at a similar time to
strawberries The garden strawberry (or simply strawberry; ''Fragaria × ananassa'') is a widely grown hybrid plant cultivated worldwide for its fruit. The genus ''Fragaria'', the strawberries, is in the rose family, Rosaceae. The fruit is appreciated f ...
. The berries are ready to harvest when the inner layer is dark purple or blue. The outer layer is dark blue and looks ripened, but the inner layer may be green with a sour flavor. Two compatible varieties are needed for
cross pollination Pollination is the transfer of pollen from an anther of a plant to the stigma of a plant, later enabling fertilisation and the production of seeds. Pollinating agents can be animals such as insects, for example bees, beetles or butterflies; birds ...
and fruit set. In North America, most Russian varieties are adapted to
hardiness zone A hardiness zone is a geographic area defined as having a certain average annual minimum temperature, a factor relevant to the survival of many plants. In some systems other statistics are included in the calculations. The original and most widely ...
s 1 to 4. The plants may take three or four years to produce an abundant harvest. Average production on a good bush is about , and bushes can maintain productivity for 30 years. Honeysuckle can be used in various processed products, such as
pastries Pastry refers to a variety of doughs (often enriched with fat or eggs), as well as the sweet and savoury baked goods made from them. The dough may be accordingly called pastry dough for clarity. Sweetened pastries are often described as '' bak ...
,
jam Fruit preserves are preparations of fruits whose main preserving agent is sugar and sometimes acid, often stored in glass jars and used as a condiment or spread. There are many varieties of fruit preserves globally, distinguished by the meth ...
s,
juice Juice is a drink made from the extraction or Cold-pressed juice, pressing of the natural liquid contained in fruit and vegetables. It can also refer to liquids that are flavored with concentrate or other biological food sources, such as meat ...
,
ice cream Ice cream is a frozen dessert typically made from milk or cream that has been flavoured with a sweetener, either sugar or an alternative, and a spice, such as Chocolate, cocoa or vanilla, or with fruit, such as strawberries or peaches. Food ...
,
yogurt Yogurt (; , from , ; also spelled yoghurt, yogourt or yoghourt) is a food produced by bacterial Fermentation (food), fermentation of milk. Fermentation of sugars in the milk by these bacteria produces lactic acid, which acts on milk protein to ...
,
sauces In cooking, a sauce is a liquid, cream, or semi- solid food, served on or used in preparing other foods. Most sauces are not normally consumed by themselves; they add flavour, texture, and visual appeal to a dish. ''Sauce'' is a French wor ...
, candies and a wine similar in color and flavor to red grape or cherry wine.


Phytochemicals

As a blue
pigment A pigment is a powder used to add or alter color or change visual appearance. Pigments are completely or nearly solubility, insoluble and reactivity (chemistry), chemically unreactive in water or another medium; in contrast, dyes are colored sub ...
ed fruit, ''Lonicera caerulea'' contains
polyphenol Polyphenols () are a large family of naturally occurring phenols. They are abundant in plants and structurally diverse. Polyphenols include phenolic acids, flavonoids, tannic acid, and ellagitannin, some of which have been used historically as ...
compounds, including cyanidin 3-glucoside, cyanidin 3-rutinoside, peonidin 3-glucoside, proanthocyanidins and
organic acids An organic acid is an organic compound with acidic properties. The most common organic acids are the carboxylic acids, whose acidity is associated with their carboxyl group –COOH. Sulfonic acids, containing the group –SO2OH, are r ...
, including
citric acid Citric acid is an organic compound with the formula . It is a Transparency and translucency, colorless Weak acid, weak organic acid. It occurs naturally in Citrus, citrus fruits. In biochemistry, it is an intermediate in the citric acid cycle, ...
.


Traditional medicine

Over centuries in East Asian countries, ''Lonicera caerulea'' has been used for supposed therapeutic applications in
traditional medicine Traditional medicine (also known as indigenous medicine or folk medicine) refers to the knowledge, skills, and practices rooted in the cultural beliefs of various societies, especially Indigenous groups, used for maintaining health and treatin ...
.


References

{{Taxonbar, from=Q157699
caerulea ''Caerulea'' is a genus of butterflies in the family Lycaenidae. It is a small genus with only two species. *'' Caerulea coeligena'' (Oberthür, 1876) western China, central China. *'' Caerulea coelestis'' (Alphéraky, 1897) Tibet, western Chi ...
Plants described in 1753 Taxa named by Carl Linnaeus Edible fruits Flora of Korea Plants used in Native American cuisine