Harpactea
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''Harpactea'' is a
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
in the family
Dysderidae The Dysderidae, also known as woodlouse hunters, sowbug-eating spiders, and cell spiders, are a family of araneomorph spiders first described by Carl Ludwig Koch in 1837. They are found primarily in Eurasia, extending into North Africa with very ...
(woodlouse hunting spiders). ''Harpactea'' is a
replacement name In biological nomenclature, a ''nomen novum'' (Latin for "new name"), replacement name (or new replacement name, new substitute name, substitute name) is a replacement scientific name that is created when technical, nomenclatural reasons have mad ...
published by W. S. Bristowe in 1939 for the unavailable name "''Harpactes''" published by R. Templeton in 1835, which had already been used for a genus of birds. They are non-web building predators that forage on the ground and on tree trunks at night, mainly in xerothermic forests. During the day, they hide in
silk Silk is a natural fiber, natural protein fiber, some forms of which can be weaving, woven into textiles. The protein fiber of silk is composed mainly of fibroin and is most commonly produced by certain insect larvae to form cocoon (silk), c ...
retreats they build under rocks or bark. '' Harpactea sadistica'' was found to use
traumatic insemination Traumatic insemination, also known as hypodermic insemination, is the mating practice in some species of invertebrates in which the male pierces the female's abdomen with his aedeagus and injects his sperm through the wound into her abdominal ca ...
, the arthropod behavior of directly inserting its sperm into the body cavity of females. It is the first time it has ever been observed in spiders.


Description

Like all woodlouse hunters, ''Harpactea'' have six eyes. The
type species In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...
, '' H. hombergi'', can grow up to a body length of . Males and females are similar, but the female has no
epigyne The epigyne or epigynum is the external genital structure of female spiders. As the epigyne varies greatly in form in different species, even in closely related ones, it often provides the most distinctive characteristic for recognizing species. ...
. Like the rest of their family, they are
nocturnal Nocturnality is a ethology, behavior in some non-human animals characterized by being active during the night and sleeping during the day. The common adjective is "nocturnal", versus diurnality, diurnal meaning the opposite. Nocturnal creatur ...
. Unlike them, ''Harpactea'' do not specialize on hunting
woodlice Woodlice are terrestrial isopods in the suborder Oniscidea. Their name is derived from being often found in old wood, and from louse, a parasitic insect, although woodlice are neither parasitic nor insects. Woodlice evolved from marine isopods ...
. ''H. rubicunda'' also hunts ''
Drassodes ''Drassodes'' is a genus of ground spiders that was first described by Niklas Westring in 1851. They are brown, gray, and red spiders that live under rocks or bark in mostly dry habitats, and are generally long, but can reach up to in length. ...
'' and other spiders, but most ''Harpactea'' feed on insects in addition to woodlice.


Taxonomy

In 1835, the name "''Harpactes''" was published by R. Templeton for a taxon split off from the related genus ''
Dysdera ''Dysdera'' is a genus of woodlouse hunting spiders that was first described by Pierre André Latreille in 1804. They originated from Central Asia to Central Europe. The family has gained many common names from their individual species, includi ...
''. Subsequent authors used this genus name for many years, but when published, ''
Harpactes ''Harpactes'' is a genus of birds in the family Trogonidae found in forests in South and Southeast Asia, extending into southernmost China. They are strongly sexually dimorphic, with females generally being duller than males. Their back is brown ...
'' had already been used for a bird genus, so it was not available. Accordingly, in 1939, W. S. Bristowe published the
replacement name In biological nomenclature, a ''nomen novum'' (Latin for "new name"), replacement name (or new replacement name, new substitute name, substitute name) is a replacement scientific name that is created when technical, nomenclatural reasons have mad ...
''Harpactea''. The
type species In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...
is ''Dysdera latreillei'', synonym "''Harpactes''" ''latreillei'', now accepted as a synonym of ''Harpactea hombergi''. Templeton did not explain his choice of genus name, but
Greek Greek may refer to: Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group *Greek language, a branch of the Indo-European language family **Proto-Greek language, the assumed last common ancestor of all kno ...
, , means 'snatcher', 'plunderer', 'pillager' or 'thief'.


Species

Almost all species of this genus appear to be
endemic Endemism is the state of a species being found only in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also foun ...
to small regions of the
Mediterranean The Mediterranean Sea ( ) is a sea connected to the Atlantic Ocean, surrounded by the Mediterranean basin and almost completely enclosed by land: on the east by the Levant in West Asia, on the north by Anatolia in West Asia and Southern ...
. , the
World Spider Catalog The World Spider Catalog (WSC) is an online searchable database concerned with spider taxonomy. It aims to list all accepted families, genera and species, as well as provide access to the related taxonomic literature. The WSC began as a series of ...
accepted 213 species from
Europe Europe is a continent located entirely in the Northern Hemisphere and mostly in the Eastern Hemisphere. It is bordered by the Arctic Ocean to the north, the Atlantic Ocean to the west, the Mediterranean Sea to the south, and Asia to the east ...
and
Northern Africa North Africa (sometimes Northern Africa) is a region encompassing the northern portion of the African continent. There is no singularly accepted scope for the region. However, it is sometimes defined as stretching from the Atlantic shores of t ...
to
Turkmenistan Turkmenistan is a landlocked country in Central Asia bordered by Kazakhstan to the northwest, Uzbekistan to the north, east and northeast, Afghanistan to the southeast, Iran to the south and southwest and the Caspian Sea to the west. Ash ...
and
Iran Iran, officially the Islamic Republic of Iran (IRI) and also known as Persia, is a country in West Asia. It borders Iraq to the west, Turkey, Azerbaijan, and Armenia to the northwest, the Caspian Sea to the north, Turkmenistan to the nort ...
: *''
Harpactea abantia ''Harpactea'' is a genus in the family Dysderidae (woodlouse hunting spiders). ''Harpactea'' is a replacement name published by W. S. Bristowe in 1939 for the unavailable name "''Harpactes''" published by R. Templeton in 1835, which had already ...
'' (Simon, 1885) – Greece *''
Harpactea abasgiana ''Harpactea'' is a genus in the family Dysderidae (woodlouse hunting spiders). ''Harpactea'' is a replacement name published by W. S. Bristowe in 1939 for the unavailable name "''Harpactes''" published by R. Templeton in 1835, which had already ...
'' Zamani & Marusik, 2024 – Georgia *''
Harpactea achsuensis ''Harpactea'' is a genus in the family Dysderidae (woodlouse hunting spiders). ''Harpactea'' is a replacement name published by W. S. Bristowe in 1939 for the unavailable name "''Harpactes''" published by R. Templeton in 1835, which had already ...
'' Dunin, 1991 – Azerbaijan *''
Harpactea acuta ''Harpactea'' is a genus in the family Dysderidae (woodlouse hunting spiders). ''Harpactea'' is a replacement name published by W. S. Bristowe in 1939 for the unavailable name "''Harpactes''" published by R. Templeton in 1835, which had already ...
'' Beladjal & Bosmans, 1997 – Algeria *''
Harpactea adicensis ''Harpactea'' is a genus in the family Dysderidae (woodlouse hunting spiders). ''Harpactea'' is a replacement name published by W. S. Bristowe in 1939 for the unavailable name "''Harpactes''" published by R. Templeton in 1835, which had already ...
'' Řezáč, 2023 – Portugal *'' Harpactea aeoliensis'' Alicata, 1973 – Italy *'' Harpactea agnolettii'' Brignoli, 1978 – Turkey *'' Harpactea alanyana'' Özkütük, Elverici, Marusik & Kunt, 2015 – Turkey *'' Harpactea albanica'' (Caporiacco, 1949) – Albania *'' Harpactea alexandrae'' Lazarov, 2006 – Bulgaria, Romania, Ukraine, Russia (Caucasus) *'' Harpactea algarvensis'' Ferrández, 1990 – Portugal *'' Harpactea alicatai'' Brignoli, 1979 – Italy (Sardinia) *'' Harpactea angustata'' (Lucas, 1846) – Algeria *'' Harpactea antoni'' Bosmans, 2009 – Greece (Lesbos) *'' Harpactea apollinea'' Brignoli, 1979 – Greece *'' Harpactea arguta'' (Simon, 1907) – France, Italy *'' Harpactea armenica'' Dunin, 1989 – Armenia, Iran *'' Harpactea arnedoi'' Kunt, Elverici, Özkütük & Yağmur, 2011 – Turkey *'' Harpactea asparuhi'' Lazarov, 2008 – Bulgaria *'' Harpactea auresensis'' Bosmans & Beladjal, 1991 – Algeria *'' Harpactea auriga'' (Simon, 1911) – Algeria *'' Harpactea aurigoides'' Bosmans & Beladjal, 1991 – Algeria *'' Harpactea azerbajdzhanica'' Dunin, 1991 – Azerbaijan *'' Harpactea azowensis'' Charitonov, 1956 – Ukraine, Russia (Europe) *'' Harpactea babori'' (Nosek, 1905) – Bulgaria, Greece, Turkey *'' Harpactea ballarini'' Kunt, Özkütük & Elverici, 2013 – Turkey *'' Harpactea bilecenoglui'' Kunt & Özkütük, 2023 – Turkey *'' Harpactea bilgenur'' Kunt & Özkütük, 2023 – Turkey *'' Harpactea blasi'' Ribera & Ferrández, 1986 – Spain *'' Harpactea brachati'' Wunderlich, 2020 – Turkey *'' Harpactea buchari'' Dunin, 1991 – Azerbaijan *'' Harpactea bulgarica'' Lazarov & Naumova, 2010 – North Macedonia, Bulgaria *'' Harpactea caligata'' Beladjal & Bosmans, 1997 – Algeria *'' Harpactea carusoi'' Alicata, 1974 – Italy, Tunisia *'' Harpactea catholica'' (Brignoli, 1984) – Greece (Crete) *'' Harpactea caucasia'' (Kulczyński, 1895) – Russia (Caucasus), Georgia *'' Harpactea cecconii'' (Kulczyński, 1908) – Cyprus, Greece? *'' Harpactea cesari'' Van Keer, 2009 – Greece (Lesbos) *'' Harpactea chaniaensis'' Bosmans, 2023 – Greece (Crete) *'' Harpactea chreensis'' Bosmans & Beladjal, 1989 – Algeria *'' Harpactea christae'' Bosmans & Beladjal, 1991 – Algeria *'' Harpactea christodeltshevi'' Bayram, Kunt & Yağmur, 2009 – Turkey *'' Harpactea clementi'' Bosmans, 2009 – Greece, Turkey *'' Harpactea coccifera'' Brignoli, 1984 – North Macedonia, Greece (Crete) *'' Harpactea colchidis'' Brignoli, 1978 – Turkey *'' Harpactea complicata'' Deltshev, 2011 – Serbia *'' Harpactea corinthia'' Brignoli, 1984 – Greece *'' Harpactea corticalis'' (Simon, 1882) – France, Italy *'' Harpactea crespoi'' Řezáč, 2023 – Portugal *'' Harpactea cressa'' Brignoli, 1984 – Greece (Crete) *'' Harpactea cruriformis'' Bosmans, 2011 – Greece (Chios) *'' Harpactea damini'' Pavlek & Arnedo, 2020 – Croatia *'' Harpactea dashdamirovi'' Dunin, 1993 – Azerbaijan *'' Harpactea decebali'' Nae, 2021 – Romania *'' Harpactea deelemanae'' Dunin, 1989 – Armenia *'' Harpactea deltshevi'' Dimitrov & Lazarov, 1999 – Bulgaria *'' Harpactea dianae'' Gavalas & Bosmans, 2023 – Greece (Iraklia) *'' Harpactea digiovannii'' Gasparo, 2014 – Greece *'' Harpactea diraoi'' Brignoli, 1978 – Turkey *'' Harpactea dobati'' Alicata, 1974 – Turkey *'' Harpactea doblikae'' (Thorell, 1875) – Bulgaria, Ukraine *''
Harpactea dolanskyi ''Harpactea'' is a genus in the family Dysderidae (woodlouse hunting spiders). ''Harpactea'' is a Nomen novum, replacement name published by W. S. Bristowe in 1939 for the unavailable name "''Harpactes''" published by R. Templeton in 1835, which ...
'' Řezáč, 2023 – Portugal *'' Harpactea dufouri'' (Thorell, 1873) – Spain *'' Harpactea dumonti'' Bosmans & Beladjal, 1991 – Algeria *''
Harpactea dunini ''Harpactea'' is a genus in the family Dysderidae (woodlouse hunting spiders). ''Harpactea'' is a replacement name published by W. S. Bristowe in 1939 for the unavailable name "''Harpactes''" published by R. Templeton in 1835, which had already ...
'' Zamani & Marusik, 2024 – Azerbaijan *'' Harpactea elvericii'' Kunt & Özkütük, 2023 – Turkey *'' Harpactea eskovi'' Dunin, 1989 – Georgia, Armenia *'' Harpactea fageli'' Brignoli, 1980 – Portugal, Spain *'' Harpactea fontetareja'' Wunderlich, 2024 – Portugal *'' Harpactea forceps'' Varol & Danışman, 2018 – Turkey *'' Harpactea forcipifera'' (Simon, 1911) – Algeria *'' Harpactea gaditana'' Pesarini, 1988 – Portugal, Spain *'' Harpactea galatica'' Brignoli, 1978 – Turkey *'' Harpactea gennargentu'' Wunderlich, 1995 – Italy (Sardinia) *'' Harpactea globifera'' (Simon, 1911) – Algeria *'' Harpactea gobustanica'' Nuruyeva & Huseynov, 2022 – Azerbaijan *'' Harpactea golovatchi'' Dunin, 1989 – Armenia *'' Harpactea gridellii'' (Caporiacco, 1951) – Italy *'' Harpactea grisea'' (Canestrini, 1868) – Switzerland, Austria, Italy, Slovenia *'' Harpactea gunselorum'' Gücel, Fuller, Göçmen & Kunt, 2018 – Cyprus *'' Harpactea hamidae'' Beladjal & Abrous, 2023 – Algeria *'' Harpactea hauseri'' Brignoli, 1976 – Greece *'' Harpactea heizerensis'' Bosmans & Beladjal, 1991 – Algeria *'' Harpactea heliconia'' Brignoli, 1984 – Greece *'' Harpactea henriquesi'' Řezáč, 2023 – Portugal *'' Harpactea henschi'' (Kulczyński, 1915) – Bosnia and Herzegovina *'' Harpactea herodis'' Brignoli, 1978 – Israel *'' Harpactea hispana'' (Simon, 1882) – Spain, France *''
Harpactea hombergi ''Harpactea hombergi'' is genus of spiders in the family Dysderidae. It is native to Europe. Taxonomy In 1763, Giovanni Antonio Scopoli described a spider under the name ''Aranea hombergii''. In 1830, Charles Walckenaer transferred the species t ...
'' (Scopoli, 1763) (
type species In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...
) – Europe *'' Harpactea hyrcanica'' Dunin, 1991 – Azerbaijan *'' Harpactea iasoni'' Gavalas & Bosmans, 2023 – Greece (Iraklia) *'' Harpactea ice'' Komnenov & Chatzaki, 2016 – Greece *'' Harpactea incerta'' Brignoli, 1979 – Greece *'' Harpactea incurvata'' Bosmans & Beladjal, 1991 – Algeria *'' Harpactea indistincta'' Dunin, 1991 – Russia (Caucasus), Azerbaijan *'' Harpactea innupta'' Beladjal & Bosmans, 1997 – Algeria *'' Harpactea isaurica'' Brignoli, 1978 – Turkey *'' Harpactea johannitica'' Brignoli, 1976 – Greece *'' Harpactea kalaensis'' Beladjal & Bosmans, 1997 – Algeria *'' Harpactea kalavachiana'' Gücel, Charalambidou, Göçmen & Kunt, 2019 – Cyprus *'' Harpactea kankilicorum'' Kunt & Özkütük, 2023 – Turkey *'' Harpactea karabachica'' Dunin, 1991 – Azerbaijan *'' Harpactea karaschkhan'' Kunt, Özkütük, Elverici, Marusik & Karakaş, 2016 – Turkey *'' Harpactea kareli'' Bosmans & Beladjal, 1991 – Algeria *'' Harpactea kencei'' Kunt, Elverici, Özkütük & Yağmur, 2011 – Turkey *'' Harpactea konradi'' Lazarov, 2009 – Bulgaria *'' Harpactea korenkoi'' Řezáč, 2023 – Portugal *'' Harpactea korgei'' Brignoli, 1979 – Turkey *'' Harpactea krejcii'' Řezáč, 2023 – Portugal *'' Harpactea krueperi'' (Simon, 1885) – Greece *'' Harpactea krumi'' Lazarov, 2010 – Bulgaria *'' Harpactea kubrati'' Lazarov, 2008 – Bulgaria *'' Harpactea kulczynskii'' Brignoli, 1976 – Albania, Greece *'' Harpactea lazarovi'' Deltshev, 2011 – Bulgaria *'' Harpactea lazonum'' Brignoli, 1978 – Turkey *'' Harpactea lepida'' (C. L. Koch, 1838) – Europe *'' Harpactea loebli'' Brignoli, 1974 – Greece *'' Harpactea logunovi'' Dunin, 1992 – Russia (Caucasus), Georgia *'' Harpactea longitarsa'' Alicata, 1974 – Algeria, Tunisia *''
Harpactea longobarda ''Harpactea'' is a genus in the family Dysderidae (woodlouse hunting spiders). ''Harpactea'' is a replacement name published by W. S. Bristowe in 1939 for the unavailable name "''Harpactes''" published by R. Templeton in 1835, which had already ...
'' Pesarini, 2001 – Spain (Balearic Is.), Italy, Ukraine *'' Harpactea lucdevosi'' Bosmans, 2023 – Greece (Iraklia) *'' Harpactea maelfaiti'' Beladjal & Bosmans, 1997 – Algeria *'' Harpactea magnibulbi'' Machado & Ferrández, 1991 – Portugal *'' Harpactea major'' (Simon, 1911) – Algeria *'' Harpactea mariae'' Komnenov, 2014 – North Macedonia *'' Harpactea martensi'' Dunin, 1991 – Azerbaijan *'' Harpactea mateparlovi'' Pavlek & Arnedo, 2020 – Croatia *'' Harpactea mcheidzeae'' Dunin, 1992 – Georgia *'' Harpactea medeae'' Brignoli, 1978 – Turkey *'' Harpactea mehennii'' Bosmans & Beladjal, 1989 – Algeria *'' Harpactea mentor'' Lazarov & Naumova, 2010 – North Macedonia, Bulgaria *'' Harpactea mertensi'' Bosmans & Beladjal, 1991 – Algeria *'' Harpactea minoccii'' Ferrández, 1982 – Portugal, Spain *'' Harpactea minuta'' Alicata, 1974 – Tunisia *'' Harpactea mithridatis'' Brignoli, 1979 – Turkey, Georgia *'' Harpactea mitidjae'' Bosmans & Beladjal, 1991 – Algeria *'' Harpactea modesta'' Dunin, 1991 – Caucasus (Russia, Georgia, Azerbaijan) *'' Harpactea monicae'' Bosmans & Beladjal, 1991 – Algeria *'' Harpactea mouzaiensis'' Bosmans & Beladjal, 1989 – Algeria *'' Harpactea muscicola'' (Simon, 1882) – France (Corsica) *'' Harpactea nachitschevanica'' Dunin, 1991 – Azerbaijan *'' Harpactea nausicaae'' Brignoli, 1976 – Albania, Kosovo, North Macedonia, Greece *'' Harpactea nenilini'' Dunin, 1989 – Armenia *'' Harpactea nuragica'' Alicata, 1966 – Italy (Sardinia) *'' Harpactea oglasana'' Gasparo, 1992 – Italy *''
Harpactea oranensis ''Harpactea'' is a genus in the family Dysderidae (woodlouse hunting spiders). ''Harpactea'' is a Nomen novum, replacement name published by W. S. Bristowe in 1939 for the unavailable name "''Harpactes''" published by R. Templeton in 1835, which ...
'' Bosmans & Beladjal, 1991 – Algeria *'' Harpactea ortegai'' Ribera & De Mas, 2003 – Spain *'' Harpactea osellai'' Brignoli, 1978 – Turkey *'' Harpactea ouarsenensis'' Bosmans & Beladjal, 1991 – Algeria *'' Harpactea ovata'' Beladjal & Bosmans, 1997 – Algeria *'' Harpactea paradoxa'' Dunin, 1992 – Georgia *'' Harpactea parthica'' Brignoli, 1980 – Iran, Turkmenistan? *'' Harpactea pekari'' Řezáč, 2023 – Portugal *'' Harpactea persephone'' Gasparo, 2011 – Greece (Crete) *'' Harpactea petrovi'' Lazarov & Dimitrov, 2018 – Bulgaria *'' Harpactea piligera'' (Thorell, 1875) – Italy *'' Harpactea pisidica'' Brignoli, 1978 – Turkey *''
Harpactea planembolus ''Harpactea'' is a genus in the family Dysderidae (woodlouse hunting spiders). ''Harpactea'' is a replacement name published by W. S. Bristowe in 1939 for the unavailable name "''Harpactes''" published by R. Templeton in 1835, which had already ...
'' Wunderlich, 2024 – Portugal *'' Harpactea popovi'' Dimitrov, Deltshev & Lazarov, 2019 – Bulgaria *'' Harpactea proxima'' Ferrández, 1990 – Portugal *'' Harpactea pugio'' Varol & Akpınar, 2016 – Turkey *'' Harpactea punica'' Alicata, 1974 – Algeria, Tunisia *'' Harpactea reniformis'' Beladjal & Bosmans, 1997 – Algeria *'' Harpactea rubicunda'' (C. L. Koch, 1838) – Europe, Caucasus (Russia, Georgia) *'' Harpactea rucnerorum'' Polenec & Thaler, 1975 – Croatia *'' Harpactea ruffoi'' Alicata, 1974 – Tunisia *'' Harpactea rugichelis'' Denis, 1955 – Lebanon *'' Harpactea sadistica'' Řezáč, 2008 – Israel *'' Harpactea saeva'' (Herman, 1879) – Slovakia, Hungary, south-eastern Europe to Ukraine *'' Harpactea salvatorei'' Platania & Arnedo, 2020 – Italy *'' Harpactea samuili'' Lazarov, 2006 – North Macedonia, Bulgaria, Greece *'' Harpactea sanctaeinsulae'' Brignoli, 1978 – Turkey *'' Harpactea sanctidomini'' Gasparo, 1997 – Italy *'' Harpactea sardoa'' Alicata, 1966 – Italy *'' Harpactea sbordonii'' Brignoli, 1978 – Turkey *'' Harpactea sciakyi'' Pesarini, 1988 – Spain *'' Harpactea secunda'' Dunin, 1989 – Armenia *'' Harpactea senalbensis'' Beladjal & Bosmans, 1997 – Algeria *'' Harpactea serena'' (Simon, 1907) – Spain, France *'' Harpactea sicula'' Alicata, 1966 – Italy (Sicily), Malta *'' Harpactea simovi'' Deltshev & Lazarov, 2018 – Bulgaria *'' Harpactea sinuata'' Beladjal & Bosmans, 1997 – Algeria *'' Harpactea spasskyi'' Dunin, 1992 – Ukraine (Crimea), Caucasus (Russia, Georgia) *'' Harpactea spilioioannis'' Bosmans & Gavalas, 2023 – Greece (Iraklia) *'' Harpactea spirembolus'' Russell-Smith, 2011 – Greece (Chios) *'' Harpactea srednagora'' Dimitrov & Lazarov, 1999 – Albania, North Macedonia, Bulgaria *'' Harpactea stalitoides'' Ribera, 1993 – Portugal *'' Harpactea stoevi'' Deltshev & Lazarov, 2018 – Bulgaria *'' Harpactea strandi'' (Caporiacco, 1939) – Italy *'' Harpactea strandjica'' Dimitrov, 1997 – Bulgaria, Turkey *'' Harpactea strinatii'' Brignoli, 1979 – Greece *'' Harpactea sturanyi'' (Nosek, 1905) – Greece, Turkey, Georgia *'' Harpactea subiasi'' Ferrández, 1990 – Portugal *'' Harpactea talyschica'' Dunin, 1991 – Azerbaijan *'' Harpactea tavirensis'' Wunderlich, 2020 – Portugal *'' Harpactea tenuiemboli'' Deltshev, 2011 – Serbia *'' Harpactea tergestina'' Gasparo, 2014 – Italy *'' Harpactea terveli'' Lazarov, 2009 – Bulgaria, Turkey *'' Harpactea thaleri'' Alicata, 1966 – Switzerland, Italy *'' Harpactea umay'' Kunt, Yağmur & Özkütük, 2023 – Turkey *'' Harpactea undosa'' Beladjal & Bosmans, 1997 – Algeria *'' Harpactea vagabunda'' Dunin, 1991 – Azerbaijan *'' Harpactea vignai'' Brignoli, 1978 – Turkey *'' Harpactea villehardouini'' Brignoli, 1979 – Greece *''
Harpactea walterdebucki ''Harpactea'' is a genus in the family Dysderidae (woodlouse hunting spiders). ''Harpactea'' is a replacement name published by W. S. Bristowe in 1939 for the unavailable name "''Harpactes''" published by R. Templeton in 1835, which had already ...
'' Bosmans, 2023 – Greece (Iraklia) *'' Harpactea wolfgangi'' Komnenov & Chatzaki, 2016 – Greece *'' Harpactea yakourensis'' Beladjal & Bosmans, 1997 – Algeria *'' Harpactea yanardagi'' Kunt & Özkütük, 2023 – Turkey *'' Harpactea zagros'' Zamani & Marusik, 2023 – Iran *'' Harpactea zaitzevi'' Charitonov, 1956 – Georgia, Azerbaijan, Armenia *'' Harpactea zannonensis'' Alicata, 1966 – Italy *'' Harpactea zjuzini'' Dunin, 1991 – Azerbaijan *'' Harpactea zoiai'' Gasparo, 1999 – Greece


References


Further reading

* * *


External links

* BBC News (April 30, 2009)
Spider sex violent but effective
* YouTube
Mating of ''H. sadistica'': Video
* Uni Bonn

* The Essex Field Club
Picture of ''H. rubicunda''

Picture of ''H. hombergi''
{{Taxonbar, from=Q3007977 Dysderidae genera Dysderidae Spiders of Africa Spiders of Asia