''Haopterus'' is a
genus
Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of
pterodactyloid
Pterodactyloidea ( ; derived from the Greek words ''πτερόν'' (''pterón'', for usual ''ptéryx'') "wing", and ''δάκτυλος'' (''dáktylos'') "finger") is one of the two traditional suborders of pterosaurs ("wing lizards"), and contai ...
pterosaur
Pterosaurs are an extinct clade of flying reptiles in the order Pterosauria. They existed during most of the Mesozoic: from the Late Triassic to the end of the Cretaceous (228 million to 66 million years ago). Pterosaurs are the earli ...
from the
Barremian
The Barremian is an age in the geologic timescale (or a chronostratigraphic stage) between 125.77 Ma (million years ago) and 121.4 ± 1.0 Ma (Historically, this stage was placed at 129.4 million to approximately 125 million years ago) It is a ...
-
Aptian
The Aptian is an age (geology), age in the geologic timescale or a stage (stratigraphy), stage in the stratigraphic column. It is a subdivision of the Early Cretaceous, Early or Lower Cretaceous epoch (geology), Epoch or series (stratigraphy), S ...
-age
Lower Cretaceous
Lower may refer to:
* ''Lower'' (album), 2025 album by Benjamin Booker
* Lower (surname)
* Lower Township, New Jersey
*Lower Receiver (firearms)
* Lower Wick Gloucestershire, England
See also
* Nizhny
{{Disambiguation ...
Yixian Formation
The Yixian Formation (; formerly Romanization of Chinese, transcribed as Yihsien Formation or Yixiang Formation) is a geological formation in Jinzhou, Liaoning, People's Republic of China, that spans the Barremian stage of the Early Cretaceous. I ...
of
Liaoning
)
, image_skyline =
, image_alt =
, image_caption = Clockwise: Mukden Palace in Shenyang, Xinghai Square in Dalian, Dalian coast, Yalu River at Dandong
, image_map = Liaoning in China (+all claims hatched).svg
, ...
,
China
China, officially the People's Republic of China (PRC), is a country in East Asia. With population of China, a population exceeding 1.4 billion, it is the list of countries by population (United Nations), second-most populous country after ...
. Its fossil remains dated back 124.6 million years ago.
Discovery and naming
It was in 2001 named by
Wang Xiaolin
Wang may refer to:
Names
* Wang (surname) (王), a common Chinese surname
* Wāng (汪), a less common Chinese surname
* Titles in Chinese nobility
* A title in Korean nobility
* A title in Mongolian nobility
Places
* Wang River in Thailand ...
and
Lü Junchang
Lü Junchang (; 1965 – 9 October 2018) was a Chinese palaeontologist and professor at the Institute of Geology, Chinese Academy of Geological Sciences. An expert on Mesozoic reptiles, he described and named dozens of dinosaur and pterosaur taxa ...
. The
type species
In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...
is ''Haopterus gracilis''. The genus name honors Professor
Hao Yichun
Hao Yichun (; 1920–2001) was a Chinese paleontologist and a member of the Chinese Communist Party. She is known as a pioneer in bringing paleontology to China, co-authoring the first Chinese textbooks in paleontology and micropaleontology. She ...
and combines her name with a Latinized Greek ''pteron'', "wing". The
specific name Specific name may refer to:
* in Database management systems, a system-assigned name that is unique within a particular database
In taxonomy, either of these two meanings, each with its own set of rules:
* Specific name (botany), the two-part (bino ...
, "slender-built" in
Latin
Latin ( or ) is a classical language belonging to the Italic languages, Italic branch of the Indo-European languages. Latin was originally spoken by the Latins (Italic tribe), Latins in Latium (now known as Lazio), the lower Tiber area aroun ...
, refers to the condition of the
metatarsals
The metatarsal bones or metatarsus (: metatarsi) are a group of five long bones in the midfoot, located between the tarsal bones (which form the heel and the ankle) and the phalanges (toes). Lacking individual names, the metatarsal bones are nu ...
.
The genus is based on
holotype
A holotype (Latin: ''holotypus'') is a single physical example (or illustration) of an organism used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of s ...
IVPP V11726, a crushed fossil found in 1998 at the
Sihetun-locality. The layer it was discovered in was
argon-dated at an age of 124.6 million years. It was the first Chinese pterosaur fossil preserving the skull. It consists of the front half of a
subadult
A juvenile is an individual organism (especially an animal) that has not yet reached its adult form, sexual maturity or size. Juveniles can look very different from the adult form, particularly in colour, and may not fill the same niche as t ...
, including a skull, lower jaws,
pectoral girdle
The shoulder girdle or pectoral girdle is the set of bones in the appendicular skeleton which connects to the arm on each side. In humans, it consists of the clavicle and scapula; in those species with three bones in the shoulder, it consists o ...
,
sternum
The sternum (: sternums or sterna) or breastbone is a long flat bone located in the central part of the chest. It connects to the ribs via cartilage and forms the front of the rib cage, thus helping to protect the heart, lungs, and major bl ...
, wings,
cervical and
dorsal
Dorsal (from Latin ''dorsum'' ‘back’) may refer to:
* Dorsal (anatomy), an anatomical term of location referring to the back or upper side of an organism or parts of an organism
* Dorsal, positioned on top of an aircraft's fuselage
The fus ...
vertebra
Each vertebra (: vertebrae) is an irregular bone with a complex structure composed of bone and some hyaline cartilage, that make up the vertebral column or spine, of vertebrates. The proportions of the vertebrae differ according to their spina ...
e, partial
pelvis
The pelvis (: pelves or pelvises) is the lower part of an Anatomy, anatomical Trunk (anatomy), trunk, between the human abdomen, abdomen and the thighs (sometimes also called pelvic region), together with its embedded skeleton (sometimes also c ...
and metatarsals.
Description

The skull of ''Haopterus'' is with a length of long and low, lacking a crest. The snout is pointed but rounded. The
maxilla
In vertebrates, the maxilla (: maxillae ) is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. The two maxil ...
and
praemaxilla
The premaxilla (or praemaxilla) is one of a pair of small cranial bones at the very tip of the upper jaw of many animals, usually, but not always, bearing teeth. In humans, they are fused with the maxilla. The "premaxilla" of therian mammals has ...
are completely fused with no visible
suture. The nasopraeorbital skull opening is elongated and elliptical with a length of , 27,6 percent of the total skull length. The lower jaws have a length of . On the front two thirds of their length teeth are present. There are twelve pairs of teeth in both the upper and the lower jaws. The
teeth
A tooth (: teeth) is a hard, calcified structure found in the jaws (or mouths) of many vertebrates and used to break down food. Some animals, particularly carnivores and omnivores, also use teeth to help with capturing or wounding prey, tear ...
are robust, sharp, pointed, and curving backwards. To the front they gradually increase in length and point more to the front. The first three pairs in the praemaxilla are very small though; the describers assumed these were replacement teeth, recently erupted.
The back of the skull and the cervical vertebrae are strongly crushed, obscuring most details. Eight dorsal vertebrae are preserved, with a total length of . The sternum was fan-shaped with a prominent keel. The wings are robustly built; the
ulna
The ulna or ulnar bone (: ulnae or ulnas) is a long bone in the forearm stretching from the elbow to the wrist. It is on the same side of the forearm as the little finger, running parallel to the Radius (bone), radius, the forearm's other long ...
is with longer than the wing metacarpal which has a length of . The
wingspan
The wingspan (or just span) of a bird or an airplane is the distance from one wingtip to the opposite wingtip. For example, the Boeing 777–200 has a wingspan of , and a wandering albatross (''Diomedea exulans'') caught in 1965 had a wingsp ...
of the type individual was estimated at 1.35 meter (4.43 ft). In comparison, the hindlimbs must have been weakly built, the metatarsal having a length of just .
Phylogeny
''Haopterus'' was by Wang classified as a member of the
Pterodactylidae
Pterodactylidae is a controversial group of pterosaurs. During the 2000s and 2010s, several competing definitions for the various Jurassic pterodactyloid groups were proposed. Pereda-Suberbiola ''et al.'' (2012) used Fabien Knoll's (2000) defini ...
, mainly because of the combination of robust teeth with the lack of a skull crest. In 2006 however, a
cladistic
Cladistics ( ; from Ancient Greek 'branch') is an approach to biological classification in which organisms are categorized in groups ("clades") based on hypotheses of most recent common ancestry. The evidence for hypothesized relationships is ...
analysis by Lü showed it was a basal member of the
Ornithocheiroidea
Pterodactyloidea ( ; derived from the Greek words ''πτερόν'' (''pterón'', for usual ''ptéryx'') "wing", and ''δάκτυλος'' (''dáktylos'') "finger") is one of the two traditional suborders of pterosaurs ("wing lizards"), and contai ...
''sensu'' Unwin (2003). Later, an analysis by Kellner ''et al.'', 2019 showed ''Haopterus'' to be a close relative of ''
Mimodactylus
''Mimodactylus'' is a genus of istiodactyliform pterosaur that lived in what is now Lebanon during the Late Cretaceous, 95 million years ago. The only known specimen was discovered in a limestone quarry near the town of Hjoula, belonging to ...
''. Together they form the family
Mimodactylidae
Mimodactylidae is a group or family of pterosaurs known from Cretaceous deposits in East Asia and the Middle East. They are phylogenetically more closely related to the Istiodactylidae than to the Ornithocheiridae or the Anhangueridae, forming th ...
, which is the sister group to the
Istiodactylidae
Istiodactylidae is a small Family (biology), family of pterosaurs. This family was named in 2001 after the type genus ''Istiodactylus'' was discovered not to be a member of the genus ''Ornithodesmus''.
Systematics and distribution
Remains of t ...
.
The
cladogram
A cladogram (from Greek language, Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an Phylogenetic tree, evolutionary tree because it does not s ...
below follows Witton's 2012 analysis:

The cladogram below is a topology recovered by Kellner ''et al.'' (2019). In the analyses, they recovered ''Haopterus'' as the sister taxon of ''
Mimodactylus
''Mimodactylus'' is a genus of istiodactyliform pterosaur that lived in what is now Lebanon during the Late Cretaceous, 95 million years ago. The only known specimen was discovered in a limestone quarry near the town of Hjoula, belonging to ...
'' within the family
Mimodactylidae
Mimodactylidae is a group or family of pterosaurs known from Cretaceous deposits in East Asia and the Middle East. They are phylogenetically more closely related to the Istiodactylidae than to the Ornithocheiridae or the Anhangueridae, forming th ...
, and placed within the more inclusive group
Istiodactyliformes
Ornithocheiromorpha (from Ancient Greek, meaning "bird hand form") is a group of pterosaurs within the suborder Pterodactyloidea. Fossil remains of this group date back from the Early Cretaceous, Early to Late Cretaceous Period (geology), period ...
.
Paleobiology
The authors concluded that its slender hindfeet meant that it was forced to move
quadruped
Quadrupedalism is a form of locomotion in which animals have four legs that are used to bear weight and move around. An animal or machine that usually maintains a four-legged posture and moves using all four legs is said to be a quadruped (fr ...
ally on land, suggesting a
piscivore
A piscivore () is a carnivorous animal that primarily eats fish. Fish were the diet of early tetrapod evolution (via water-bound amphibians during the Devonian period); insectivory came next; then in time, the more terrestrially adapted repti ...
lifestyle as a specialised soarer.
[Wang X. and Lü J. (2001). Discovery of a pterodactyloid pterosaur from the Yixian Formation of western Liaoning, China. ''Chinese Science Bulletin'' 45(12):447–454.] Although its close relative ''
Mimodactylus
''Mimodactylus'' is a genus of istiodactyliform pterosaur that lived in what is now Lebanon during the Late Cretaceous, 95 million years ago. The only known specimen was discovered in a limestone quarry near the town of Hjoula, belonging to ...
'' was considered to be a faunivore feeding on crustaceans, suggested by Kellner ''et al.'', 2019.
See also
*
List of pterosaur genera
This list of pterosaurs is a comprehensive listing of all Genus, genera that have ever been included in the order Pterosauria, excluding purely vernacular terms. The list includes all commonly accepted genera, but also genera that are now considere ...
*
Timeline of pterosaur research
This timeline of pterosaur research is a chronologically ordered list of important fossil discoveries, controversies of interpretation, and Biological taxonomy, taxonomic revisions of pterosaurs, the famed flying reptiles of the Mesozoic Era (ge ...
References
External links
''Haopterus''in The Pterosauria
{{Portal bar, Paleontology, China
Pteranodontoidea
Early Cretaceous pterosaurs of Asia
Fossil taxa described in 2001
Yixian fauna
Taxa named by Lü Junchang
Pterosaur genera