Hacrobia
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The cryptomonads-haptophytes assemblage is a proposed but disputed monophyletic grouping of unicellular
eukaryote The eukaryotes ( ) constitute the Domain (biology), domain of Eukaryota or Eukarya, organisms whose Cell (biology), cells have a membrane-bound cell nucleus, nucleus. All animals, plants, Fungus, fungi, seaweeds, and many unicellular organisms ...
s that are not included in the SAR supergroup. Several alternative names have been used for the group, including Hacrobia (derived from "ha-" referring to Haptophyta, "-cr-" referring to cryptomonads, and "-bia" as a general suffix referring to life); CCTH (standing for Cryptophyta, Centrohelida, Telonemia and Haptophyta); and "Eukaryomonadae". , it is unclear whether this group is
monophyletic In biological cladistics for the classification of organisms, monophyly is the condition of a taxonomic grouping being a clade – that is, a grouping of organisms which meets these criteria: # the grouping contains its own most recent co ...
or not; results of phylogenetic studies are "often dependent on the selection of taxa and gene data set". Two 2012 studies produced opposite results.


Members

In the past,
heterokont The stramenopiles, also called heterokonts, are protists distinguished by the presence of stiff tripartite external hairs. In most species, the hairs are attached to flagella, in some they are attached to other areas of the cellular surface, an ...
s, haptophytes, and cryptomonads have sometimes been grouped together in a group known as chromists. Though the heterokonts are now split out, Cryptophyta and Haptophyta are considered in some studies to be closely related (and are sometimes simply referred to as the "Cryptophyta+Haptophyta" group). A 2009 paper suggested that the
Telonemia Telonemia is a phylum of microscopic eukaryotes commonly known as telonemids. They are unicellular free-living flagellates with a unique combination of Cell (biology), cell structures, including a highly complex cytoskeleton unseen in other eu ...
and centrohelids may form a clade with the cryptophytes and haptophytes. The
picobiliphyte Picozoa, Picobiliphyta, Picobiliphytes, or Biliphytes are protists of a phylum of marine unicellular heterotrophic eukaryotes with a size of less than about 3 micrometers. They were formerly treated as Eukaryote, eukaryotic algae and the smallest ...
s may belong in this group but are too poorly known to be classified with confidence. Several recent studies have concluded that Haptophyta and Cryptophyta do not form a monophyletic group. The former are a sister group to the SAR group, the latter cluster with the Archaeplastida (plants in the broad sense). , it remains unclear whether the Hacrobia forms a monophyletic group. Another study suggested the following arrangement: centrohelids are related to haptophytes and form the clade
Haptista Haptista is a proposed group of protists made up of centrohelids and haptophytes. Phylogenomic studies indicate that Haptista, together with ''Ancoracysta twista'', forms a sister clade to the SAR supergroup#Internal phylogeny, TSAR Supergroup (b ...
; Haptista is the sister group to SAR; Cryptista are related to Archaeplastida; and Haptista + SAR is the sister clade to Cryptista + Archaeplastida.


Phylogeny

Based on work done by Silar 2016.


References


External links


Tree of Life: Hacrobia
{{Taxonbar, from=Q1044732 Subkingdoms