''Gwyneddosaurus'' is a possibly invalid

genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...

extinct Extinction is the termination of an organism by the death of its Endling, last member. A taxon may become Functional extinction, functionally extinct before the death of its last member if it loses the capacity to Reproduction, reproduce and ...

aquatic

tanystropheid Tanystropheidae is an extinct family (biology), family of archosauromorph reptiles that lived throughout the Triassic Period, often considered to be "protorosaurs". They are characterized by their long, stiff necks formed from elongated cervical ...

reptile. The

type species In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...

, ''G. erici'' was described in 1945 by Wilhelm Bock, who identified it as a

coelurosauria Coelurosauria (; from Greek, meaning "hollow-tailed lizards") is the clade containing all theropod dinosaurs more closely related to birds than to carnosaurs. Coelurosauria is a subgroup of theropod dinosaurs that includes compsognathids, tyra ...

dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic Geological period, period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the #Evolutio ...

related to ''

Podokesaurus ''Podokesaurus'' is a genus of coelophysoid dinosaur that lived in what is now the eastern United States during the Early Jurassic Period. The first fossil was discovered by the geologist Mignon Talbot near Mount Holyoke, Massachusetts, in 19 ...

'' (at the time, "podokesaurids" were thought to be coelurosaurians). Its remains were found in the

Upper Triassic The Late Triassic is the third and final epoch of the Triassic Period in the geologic time scale, spanning the time between Ma and Ma (million years ago). It is preceded by the Middle Triassic Epoch and followed by the Early Jurassic Epoch. T ...

Lockatong Formation The Triassic Lockatong Formation is a mapped bedrock unit in Pennsylvania, New Jersey, and New York. It is named after the Lockatong Creek in Hunterdon County, New Jersey. Description The Lockatong is defined as a light to dark gray, greenish-g ...

of Montgomery County, eastern

Pennsylvania Pennsylvania, officially the Commonwealth of Pennsylvania, is a U.S. state, state spanning the Mid-Atlantic (United States), Mid-Atlantic, Northeastern United States, Northeastern, Appalachian, and Great Lakes region, Great Lakes regions o ...

, and the holotype includes skull fragments, several

vertebra Each vertebra (: vertebrae) is an irregular bone with a complex structure composed of bone and some hyaline cartilage, that make up the vertebral column or spine, of vertebrates. The proportions of the vertebrae differ according to their spina ...

, ribs,

gastralia Gastralia (: gastralium) are dermal bones found in the ventral body wall of modern crocodilians and tuatara, and many prehistoric tetrapods. They are found between the sternum and pelvis, and do not articulate with the vertebrae. In these reptil ...

, partial

shoulder The human shoulder is made up of three bones: the clavicle (collarbone), the scapula (shoulder blade), and the humerus (upper arm bone) as well as associated muscles, ligaments and tendons. The articulations between the bones of the shoulder m ...

and hip bones, and several forelimb and hindlimb elements found in soft

shale Shale is a fine-grained, clastic sedimentary rock formed from mud that is a mix of flakes of Clay mineral, clay minerals (hydrous aluminium phyllosilicates, e.g., Kaolinite, kaolin, aluminium, Al2Silicon, Si2Oxygen, O5(hydroxide, OH)4) and tiny f ...

, while the paratype includes a femur and a tibia. The

type specimen In biology, a type is a particular wikt:en:specimen, specimen (or in some cases a group of specimens) of an organism to which the scientific name of that organism is formally associated. In other words, a type is an example that serves to ancho ...

is ANSP 15072 and it was discovered by Bock's four-year-old son while the paratype is only listed as ?(ASNP coll.). It was not a large animal; the type skeleton was estimated by Bock as long, and its

thigh bone The femur (; : femurs or femora ), or thigh bone is the only bone in the thigh — the region of the lower limb between the hip and the knee. In many four-legged animals the femur is the upper bone of the hindleg. The top of the femur fits in ...

was only 23 millimeters long (0.91 in).

Friedrich von Huene Baron Friedrich Richard von Hoyningen-Huene (22 March 1875 – 4 April 1969) was a German nobleman paleontologist who described a large number of dinosaurs, more than anyone else in 20th-century Europe. He studied a range of Permo-Carbonife ...

assigned the animal to

Protorosauria Protorosauria is an extinct, likely paraphyletic group of basal archosauromorph reptiles from the latest Middle Permian ( Capitanian stage) to the end of the Late Triassic ( Rhaetian stage) of Asia, Europe and North America. It was named by t ...

in 1948 as the smallest known member of the group, finding it to be most like ''

Macrocnemus ''Macrocnemus'' is an extinct genus of archosauromorph reptile known from the Middle Triassic (Late Anisian to Ladinian) of Europe and China. ''Macrocnemus'' is a member of the Tanystropheidae family and includes three species''. Macrocnemus bas ...

''. Steel (1970) classified it as a

theropod Theropoda (; from ancient Greek , (''therion'') "wild beast"; , (''pous, podos'') "foot"">wiktionary:ποδός"> (''pous, podos'') "foot" is one of the three major groups (clades) of dinosaurs, alongside Ornithischia and Sauropodom ...

, whereas Olsen and Baird (1986) identified it as a

chimera Chimera, Chimaera, or Chimaira (Greek for " she-goat") originally referred to: * Chimera (mythology), a fire-breathing monster of ancient Lycia said to combine parts from multiple animals * Mount Chimaera, a fire-spewing region of Lycia or Cilicia ...

of mixed remains from a

coelacanth Coelacanths ( ) are an ancient group of lobe-finned fish (Sarcopterygii) in the class Actinistia. As sarcopterygians, they are more closely related to lungfish and tetrapods (the terrestrial vertebrates including living amphibians, reptiles, bi ...

and possibly the tanystropheid ''

Tanytrachelos ''Tanytrachelos'' is an extinct genus of tanystropheid archosauromorph reptile from the Late Triassic of the eastern United States. It contains a single species, ''Tanytrachelos ahynis'', which is known from several hundred fossil specimens pres ...

''; Olsen and Flynn (1989) later modified this interpretation, describing the

of ''Gwyneddosaurus'' as a "gastric ejection" (regurgitation) composed of ''Tanytrachelos'' bones and possibly scraps of a coelacanth. They noted that this would make ''Gwyneddosaurus'' a

senior synonym In taxonomy, the scientific classification of living organisms, a synonym is an alternative scientific name for the accepted scientific name of a taxon. The botanical and zoological codes of nomenclature treat the concept of synonymy differently. ...

of ''Tanytrachelos'', and recommended conserving the younger but better-represented genus.

References

''Gwyneddosaurus'' at the Paleobiology Database
{{Taxonbar, from=Q5623838 Tanystropheidae Prehistoric reptile genera Norian genera Late Triassic reptiles of North America Triassic geology of Pennsylvania Paleontology in Pennsylvania Aquatic reptiles Fossil taxa described in 1945