Gulf Menhaden
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The Gulf menhaden (''Brevoortia patronus'') is a small marine filter-feeding fish belonging to the family
Alosidae The Alosidae, or the shads, are a family of clupeiform fishes. The family currently comprises four genera worldwide, and about 32 species. The shads are pelagic (open water) schooling fish, of which many are anadromous or even landlocked. Seve ...
. The range of Gulf menhaden encompasses the entirety of the
Gulf of Mexico The Gulf of Mexico () is an oceanic basin and a marginal sea of the Atlantic Ocean, mostly surrounded by the North American continent. It is bounded on the northeast, north, and northwest by the Gulf Coast of the United States; on the southw ...
nearshore waters, with the exception of the extreme eastern Yucatan and western
Cuba Cuba, officially the Republic of Cuba, is an island country, comprising the island of Cuba (largest island), Isla de la Juventud, and List of islands of Cuba, 4,195 islands, islets and cays surrounding the main island. It is located where the ...
.FAO 2002. The living marine resources of the western central Atlantic. ASIH special publication No. 5, Kent E. Carpenter, ed. ISSN 1020-6868. Evidence from morphology Dahlberg, M.D. 1970. Atlantic and Gulf of Mexico menhadens, genus Brevoortia (Pisces:Clupeidae). Bulletin of the Florida State Museum, Biological Sciences 15:91-162. and DNA analyses suggest that the Gulf menhaden is the Gulf of Mexico complement to the Atlantic menhaden (''Brevoortia tyrannus''). Both species support large commercial reduction fisheries, with Gulf menhaden supporting the second largest fishery, by weight, in the United States.Pritchard, E.S. 2005. Fisheries of the United States 2004. Silver Spring, MD: National Marine Fisheries Service, Office of Science and Technology, pp. 1–19.


Range and distribution

The Gulf menhaden resides throughout the Gulf of Mexico, but its distribution is patchy. The center of distribution of the species appears to be the northwest/northcentral Gulf, particularly in
Louisiana Louisiana ( ; ; ) is a state in the Deep South and South Central regions of the United States. It borders Texas to the west, Arkansas to the north, and Mississippi to the east. Of the 50 U.S. states, it ranks 31st in area and 25 ...
and
Texas Texas ( , ; or ) is the most populous U.S. state, state in the South Central United States, South Central region of the United States. It borders Louisiana to the east, Arkansas to the northeast, Oklahoma to the north, New Mexico to the we ...
where populations are very large and numerous. In the southern Gulf of Mexico, the range of Gulf menhaden overlaps that of the closely related finescale menhaden (''Brevoortia gunteri''), and there is evidence for resource partitioning (a process whereby closely related species occurring in close proximity results in subtle differences in ecological niches) between these species.Castillo-Revera, M., A. Kobelkowsky and V. Zamayoa. 1996. Food resource partitioning and trophic morphology of Brevoortia gunteri and B. patronus. Journal of Fish Biology 49:1102-1111. In the eastern Gulf, the range of Gulf menhaden overlaps that of the yellowfin menhaden (''Brevoortia smithi''), and hybridization between these species has been demonstrated using morphological and DNA evidence.Anderson, J.D. and W.J. Karel. 2007. Genetic evidence for asymmetric hybridization between menhadens (Brevoortia spp.) from peninsular Florida. Journal of Fish Biology 71b:235-249. Gulf menhaden also may have a presence on the southern Atlantic coast of Florida, although this finding is based primarily upon DNA evidence.


General biology

Gulf menhaden are commonly eight inches in length but can reach 12 inches. Gulf menhaden are a dull silver with a greenish back. Like their Atlantic counterpart, Gulf menhaden have a prominent black spot found behind the gill cover followed by a row of smaller spots.


Diet

Gulf menhaden are filter feeders, meaning that they collect food by filtering water through modifications of the branchial apparatus (gill or
branchial arch Branchial arches or gill arches are a series of paired bony/ cartilaginous "loops" behind the throat ( pharyngeal cavity) of fish, which support the fish gills. As chordates, all vertebrate embryos develop pharyngeal arches, though the eve ...
es and
gill raker Gill rakers in fish are bony or cartilaginous processes that project from the branchial arch (gill arch) and are involved with suspension feeding tiny prey. They are not to be confused with the gill filaments that compose the fleshy part of th ...
s). Like
Atlantic menhaden The Atlantic menhaden (''Brevoortia tyrannus'') is a North American species of fish in the herring family, Alosidae. Atlantic menhaden are found in North Atlantic coastal and estuarine waters from Nova Scotia south to northern Florida. They ar ...
, Gulf menhaden's diet depends on the size of their gill rakers, which change as menhaden age. Larval gulf menhaden feed primary on zooplankton because the rakers are not well developed. The juveniles, which generally correspond to when they are under the age of one, feed on
phytoplankton Phytoplankton () are the autotrophic (self-feeding) components of the plankton community and a key part of ocean and freshwater Aquatic ecosystem, ecosystems. The name comes from the Greek language, Greek words (), meaning 'plant', and (), mea ...
. As they age and their gill rakers fully develop, menhaden shift their diet to primarily consume
zooplankton Zooplankton are the heterotrophic component of the planktonic community (the " zoo-" prefix comes from ), having to consume other organisms to thrive. Plankton are aquatic organisms that are unable to swim effectively against currents. Consequent ...
."Menhaden Facts." Gulf States Marine Fisheries Commission.


Life cycle

Spawning occurs offshore in winter (October–March). Eggs and larvae are
pelagic The pelagic zone consists of the water column of the open ocean and can be further divided into regions by depth. The word ''pelagic'' is derived . The pelagic zone can be thought of as an imaginary cylinder or water column between the sur ...
and are carried into estuarine nursery areas via prevailing currents. As a result, migration at this stage can be lengthy, and populations of Gulf menhaden throughout the Gulf of Mexico are generally thought to comprise a single genetic stock.


Relationship with humans


Commercial fishery

The Gulf menhaden fishery is one of the largest in the United States. In 2013, the fishery supported four of the nation's top ten ports by volume of landings. Gulf menhaden are harvested primarily for fish meal and fish oil based products. A much smaller number of menhaden are caught for use as bait. In addition to being one of the largest fisheries in the US, the Gulf menhaden fishery has also been recognized internationally for its sustainability. The fishery's sustainability has also been certified by independent organizations.
Friend of the Sea Friend of the Sea is a project of the World Sustainability Organization for the certification and promotion of seafood from sustainable fisheries and sustainable aquaculture. It is the only certification scheme which, with the same logo, certifies ...
, an international seafood sustainability certification program, has recognized both the
Atlantic menhaden The Atlantic menhaden (''Brevoortia tyrannus'') is a North American species of fish in the herring family, Alosidae. Atlantic menhaden are found in North Atlantic coastal and estuarine waters from Nova Scotia south to northern Florida. They ar ...
and Gulf menhaden fisheries as sustainable. This is both due to the healthy status of the stock as well as the fishery's low levels of bycatch, which it achieves with the use of purse seine nets.


Recreational fishery

The Gulf menhaden is an important bait fish for recreational anglers.


Management

Gulf menhaden are managed by an interstate compact called the Gulf States Marine Fisheries Commission (GSMFC). According to the most recent 2013 stock assessment by the GSMFC, Gulf menhaden are "neither overfished nor experiencing overfishing." In addition, according to the GSMFC, "the Gulf menhaden fishery is probably the most closely monitored and managed fishery in the Gulf of Mexico." An example of the fishery's monitoring is the Gulf of Mexico purse seine fishery's participation in NOAA's Southeast Fisheries Observer Program since 2011. Specifically, the Menhaden Advisory Committee (MAC) is the GSMFC subcommittee that oversees menhaden management.


Dead zones

According to the GSMFC: "Menhaden do not have the capacity to reduce unwanted phytoplankton blooms that arise from manmade sources, primarily because they eat mostly zooplankton. In addition, menhaden excrete large amounts of ammonia (a nitrogenous product), contributing to an already high nitrogen load." Also, the commercial menhaden fishery only targets adult menhaden, which consume zooplankton, not juvenile menhaden, which do consume phytoplankton NOAA, "Brief History of the Gulf Menhaden Purse-Seine Fishery"
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Sources


External links


Gulf States Marine Fisheries Commission
menhaden page {{Authority control Gulf menhaden Fish of the Gulf of Mexico Gulf menhaden