Guanaco
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The guanaco ( ; ''Lama guanicoe'') is a
camelid Camelids are members of the biological family (biology), family Camelidae, the only currently living family in the suborder Tylopoda. The seven extant taxon, extant members of this group are: dromedary, dromedary camels, Bactrian camels, wild Bac ...
native to
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a considerably smaller portion in the Northern Hemisphere. It can also be described as the southern Subregion#Americas, subregion o ...
, closely related to the
llama The llama (; or ) (''Lama glama'') is a domesticated South American camelid, widely used as a List of meat animals, meat and pack animal by Inca empire, Andean cultures since the pre-Columbian era. Llamas are social animals and live with ...
. Guanacos are one of two wild South American camelids; the other species is the
vicuña The vicuña (''Lama vicugna'') or vicuna (both , very rarely spelled ''vicugna'', Vicugna, its former genus name) is one of the two wild South American camelids, which live in the high alpine tundra, alpine areas of the Andes; the other cameli ...
, which lives at higher elevations.


Etymology

The guanaco gets its name from the Quechua word ''wanaku''. Young guanacos are called ''chulengos'' or "guanaquitos".


Characteristics

Guanacos stand between at the
shoulder The human shoulder is made up of three bones: the clavicle (collarbone), the scapula (shoulder blade), and the humerus (upper arm bone) as well as associated muscles, ligaments and tendons. The articulations between the bones of the shoulder m ...
, body length of , and weigh . Their color varies very little (unlike the domestic
llama The llama (; or ) (''Lama glama'') is a domesticated South American camelid, widely used as a List of meat animals, meat and pack animal by Inca empire, Andean cultures since the pre-Columbian era. Llamas are social animals and live with ...
), ranging from a light brown to dark cinnamon and shading to white underneath. Guanacos have grey faces and small, straight ears. The lifespan of a guanaco can be as long as 28 years. Guanacos are one of the largest terrestrial
mammal A mammal () is a vertebrate animal of the Class (biology), class Mammalia (). Mammals are characterised by the presence of milk-producing mammary glands for feeding their young, a broad neocortex region of the brain, fur or hair, and three ...
s native to South America today.San Diego Zoo's Animal Bytes
Other terrestrial mammalian
megafauna In zoology, megafauna (from Ancient Greek, Greek μέγας ''megas'' "large" and Neo-Latin ''fauna'' "animal life") are large animals. The precise definition of the term varies widely, though a common threshold is approximately , this lower en ...
weighing as much or more than the guanaco include the
tapir Tapirs ( ) are large, herbivorous mammals belonging to the family Tapiridae. They are similar in shape to a Suidae, pig, with a short, prehensile nose trunk (proboscis). Tapirs inhabit jungle and forest regions of South America, South and Centr ...
s, the marsh deer, the
white-tailed deer The white-tailed deer (''Odocoileus virginianus''), also known Common name, commonly as the whitetail and the Virginia deer, is a medium-sized species of deer native to North America, North, Central America, Central and South America. It is the ...
, the spectacled bear, and the
jaguar The jaguar (''Panthera onca'') is a large felidae, cat species and the only extant taxon, living member of the genus ''Panthera'' that is native to the Americas. With a body length of up to and a weight of up to , it is the biggest cat spe ...
. Guanacos have thick skin on their necks, a trait also found in their domestic counterparts, the
llama The llama (; or ) (''Lama glama'') is a domesticated South American camelid, widely used as a List of meat animals, meat and pack animal by Inca empire, Andean cultures since the pre-Columbian era. Llamas are social animals and live with ...
, and their relatives, the wild
vicuña The vicuña (''Lama vicugna'') or vicuna (both , very rarely spelled ''vicugna'', Vicugna, its former genus name) is one of the two wild South American camelids, which live in the high alpine tundra, alpine areas of the Andes; the other cameli ...
and domesticated
alpaca The alpaca (''Lama pacos'') is a species of South American camelid mammal. Traditionally, alpacas were kept in herds that grazed on the level heights of the Andes of Southern Peru, Western Bolivia, Ecuador, and Northern Chile. More recentl ...
. This protects their necks from predator attacks. Bolivians use the neck skin of these animals to make
shoe A shoe is an item of footwear intended to protect and comfort the human foot. Though the human foot can adapt to varied terrains and climate conditions, it is vulnerable, and shoes provide protection. Form was originally tied to function, but ...
s, flattening and pounding the skin to be used for the soles. In Chile, hunting is allowed only in
Tierra del Fuego Tierra del Fuego (, ; Spanish for "Land of Fire", rarely also Fireland in English) is an archipelago off the southernmost tip of the South America, South American mainland, across the Strait of Magellan. The archipelago consists of the main is ...
, where the only population not classified as endangered in the country resides. Between 2007 and 2012, 13,200 guanacos were legally hunted in Tierra del Fuego.


Diet

Like all camels, Guanacos are herbivores, grazing on grasses, shrubs, herbs, lichens, fungi, cacti, and flowers. The food is swallowed with little chewing and first enters the forestomach to be digested finally after rumination. This process is similar to that of ruminants, to which camels are not zoologically related. The camels' digestive system is likely to have developed independently of ruminants, which is evidenced by the fact that the forestomachs are equipped with glands.


Blood

Guanacos are often found at altitudes up to above sea level, except in
Patagonia Patagonia () is a geographical region that includes parts of Argentina and Chile at the southern end of South America. The region includes the southern section of the Andes mountain chain with lakes, fjords, temperate rainforests, and glaciers ...
, where the southerly latitude means ice covers the vegetation at these altitudes. Their blood is rich in
red blood cell Red blood cells (RBCs), referred to as erythrocytes (, with -''cyte'' translated as 'cell' in modern usage) in academia and medical publishing, also known as red cells, erythroid cells, and rarely haematids, are the most common type of blood cel ...
s, enabling them to survive in the low oxygen levels found at these high altitudes. A teaspoon of guanaco blood contains about 68 million red blood cells, four times that of a human.


Guanaco fiber

Guanaco fiber is particularly prized for its soft, warm feel and is found in luxury
fabric Textile is an umbrella term that includes various fiber-based materials, including fibers, yarns, filaments, threads, and different types of fabric. At first, the word "textiles" only referred to woven fabrics. However, weaving is no ...
. In South America, the guanaco's soft wool is valued second only to that of
vicuña wool Vicuña wool refers to the hair of the South American vicuña, a camelid related to llamas and alpacas. The wool has, after shahtoosh, the second smallest fiber diameter of all animal hair and is the most expensive legal wool. Properties The do ...
. The pelts, particularly from the calves, are sometimes used as a substitute for
red fox The red fox (''Vulpes vulpes'') is the largest of the true foxes and one of the most widely distributed members of the order Carnivora, being present across the entire Northern Hemisphere including most of North America, Europe and Asia, plus ...
pelts, because the texture is difficult to differentiate. Like their domestic descendant, the llama, the guanaco is double-coated with coarse
guard hair Guard hair or overhair is the outer layer of hair of most mammals, which overlay the fur. Guard hairs are long and coarse and protect the rest of the pelage (fur) from abrasion and frequently from moisture. They are visible on the surface of the ...
s and a soft undercoat, the hairs of which are about 16–18 μm in
diameter In geometry, a diameter of a circle is any straight line segment that passes through the centre of the circle and whose endpoints lie on the circle. It can also be defined as the longest Chord (geometry), chord of the circle. Both definitions a ...
and comparable to cashmere.


Subspecies

*''Lama guanicoe guanicoe'' from the north *''Lama guanicoe cacsilensis'' from the south


Population and distribution

Guanacos inhabit the steppes, scrublands and mountainous regions of
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a considerably smaller portion in the Northern Hemisphere. It can also be described as the southern Subregion#Americas, subregion o ...
. They are found in the
altiplano The Altiplano (Spanish language, Spanish for "high plain"), Collao (Quechuan languages, Quechua and Aymara language, Aymara: Qullaw, meaning "place of the Qulla people, Qulla") or Andean Plateau, in west-central South America, is the most extens ...
of
Peru Peru, officially the Republic of Peru, is a country in western South America. It is bordered in the north by Ecuador and Colombia, in the east by Brazil, in the southeast by Bolivia, in the south by Chile, and in the south and west by the Pac ...
,
Bolivia Bolivia, officially the Plurinational State of Bolivia, is a landlocked country located in central South America. The country features diverse geography, including vast Amazonian plains, tropical lowlands, mountains, the Gran Chaco Province, w ...
and
Chile Chile, officially the Republic of Chile, is a country in western South America. It is the southernmost country in the world and the closest to Antarctica, stretching along a narrow strip of land between the Andes, Andes Mountains and the Paci ...
, and in
Patagonia Patagonia () is a geographical region that includes parts of Argentina and Chile at the southern end of South America. The region includes the southern section of the Andes mountain chain with lakes, fjords, temperate rainforests, and glaciers ...
, with a small population in
Paraguay Paraguay, officially the Republic of Paraguay, is a landlocked country in South America. It is bordered by Argentina to the Argentina–Paraguay border, south and southwest, Brazil to the Brazil–Paraguay border, east and northeast, and Boli ...
. In
Argentina Argentina, officially the Argentine Republic, is a country in the southern half of South America. It covers an area of , making it the List of South American countries by area, second-largest country in South America after Brazil, the fourt ...
they are more numerous in Patagonian regions, as well as in places such as Isla Grande de Tierra del Fuego. In these areas, they have more robust populations, since grazing competition from livestock is limited. Guanaco respond to forage availability, occupying zones with low to intermediate food availability in the breeding season and those with the highest availability in the non-breeding season. Estimates, as of 2016, place their numbers around 1.5 to 2 million animals: 1,225,000–1,890,000 in Argentina, 270,000–299,000 in Chile, 3,000 in Peru, 150–200 in Bolivia and 20–100 in Paraguay. This is only 3–7% of the guanaco population before the arrival of the Spanish conquistadors in South America. A small population introduced by John Hamilton exists on Staats Island in the Falkland Islands (Malvinas), with a population of around 400 as of 2003. In Torres del Paine National Park, the numbers of guanacos increased from 175 in 1975 to 3,000 in 1993. Guanacos live in herds composed of females, their young, and a dominant male. Bachelor males form separate herds. While reproductive groups tend to remain small, often containing no more than 10 adults, bachelor herds may contain as many as 50 males. They can run at per hour, often over steep and rocky terrain. They are also excellent swimmers. A guanaco's typical lifespan is 20 to 25 years. In Bolivia, the habitat of Guanacos is found to be threatened by woody plant encroachment.


Atacama Desert

Some guanacos live in the
Atacama Desert The Atacama Desert () is a desert plateau located on the Pacific Ocean, Pacific coast of South America, in the north of Chile. Stretching over a strip of land west of the Andes Mountains, it covers an area of , which increases to if the barre ...
, where in some areas it has not rained for over 50 years. A mountainous coastline running parallel to the desert enables them to survive in what are called "fog oases" or lomas. Where the cool water touches the hotter land, the air above the desert is cooled, creating a fog and thus water vapor. Winds carry the fog across the desert, where cacti catch the water droplets and
lichen A lichen ( , ) is a hybrid colony (biology), colony of algae or cyanobacteria living symbiotically among hypha, filaments of multiple fungus species, along with yeasts and bacteria embedded in the cortex or "skin", in a mutualism (biology), m ...
s that cling to the cacti soak it in like a sponge. Guanacos then eat the cactus flowers and the lichens. File:Guanacos, Torres del Paine.jpg, Guanacos in Torres del Paine National Park in
Patagonia Patagonia () is a geographical region that includes parts of Argentina and Chile at the southern end of South America. The region includes the southern section of the Andes mountain chain with lakes, fjords, temperate rainforests, and glaciers ...
File:The Guanacos of Atacama.jpg, alt=Guanacos near the La Silla Observatory, 2400 meters above sea level., Guanacos near the La Silla Observatory, 2400 meters above sea level File:GuanacoBonesInAtacama.jpg, The remains of a guanaco scattered in the Atacama Desert, southwest of Cerro Paranal: The only intact section of skin is the thicker skin around the neck File:Lama guanicoe in Parque Nacional Torres del Paine in Patagonia, Chile.jpg, Guanacos near Torres del Paine,
Chile Chile, officially the Republic of Chile, is a country in western South America. It is the southernmost country in the world and the closest to Antarctica, stretching along a narrow strip of land between the Andes, Andes Mountains and the Paci ...


Ecology

The guanaco is a diurnal animal. It lives in small herds consisting of one male and several females with their young. When the male detects danger, he warns the group by bleating. The guanaco can run up to . This speed is important for the survival of guanacos because they cannot easily hide in the open grasslands of the Altiplano. Natural predators of the guanaco include pumas and the culpeo or Andean fox. Fox predation was unknown until 2007 when predators began to be observed in the Karukinka Reserve in Tierra del Fuego. Scientists attribute this to the unfavourable climatic conditions on the island, which are causing food to become scarce, weakening the animals. The absence of pumas on Tierra del Fuego is also believed to be a factor that allows the fox to occupy their ecological niche. Finally, it is believed that this behaviour is not new, as the fox is nocturnal, which makes any predation challenging to observe. Faced with the threat of the fox, guanacos resort to cooperative strategies to protect their young with a shield formation, a circle around the vulnerable. If they are successful, they chase the fox away, which would be impossible with a puma. When threatened, the guanaco alerts the rest of the herd with a high-pitched bleating sound, which sounds similar to a short, sharp laugh. The male usually runs behind the herd to defend them. Though typically mild-mannered, guanacos often spit when threatened, and can do so up to a distance of six feet.


Mating season

Mating season occurs between November and February, during which males often fight violently to establish dominance and breeding rights. Eleven-and-a-half months later, a single chulengo is born. Chulengos are able to walk immediately after birth. Male chulengos are chased off from the herd by the dominant male at around one year old.


Conservation

While not considered an endangered species in southern Argentina and Chile, dead guanacos are a common sight throughout this region where they are entangled on fences. Studies have found that annual yearling mortality on fences (5.53%) was higher than adult mortality (0.84%) and was more frequent in ovine (93 cm high) than bovine (113 cm) fences. Most guanacos died entangled by their legs in the highest wire when trying to jump over the fence.


Captivity and domestication

Around 300 guanacos are in U.S. zoos, and around 200 are registered in private herds. Guanacos have long been thought to be the parent species of the domesticated
llama The llama (; or ) (''Lama glama'') is a domesticated South American camelid, widely used as a List of meat animals, meat and pack animal by Inca empire, Andean cultures since the pre-Columbian era. Llamas are social animals and live with ...
, which was confirmed via
molecular phylogenetic Molecular phylogenetics () is the branch of phylogeny that analyzes genetic, hereditary molecular differences, predominantly in DNA sequences, to gain information on an organism's evolutionary relationships. From these analyses, it is possible to ...
analysis in 2001, although the analysis also found that domestic llamas had experienced considerable cross-hybridization with
alpaca The alpaca (''Lama pacos'') is a species of South American camelid mammal. Traditionally, alpacas were kept in herds that grazed on the level heights of the Andes of Southern Peru, Western Bolivia, Ecuador, and Northern Chile. More recentl ...
s, which are descended from the wild
vicuña The vicuña (''Lama vicugna'') or vicuna (both , very rarely spelled ''vicugna'', Vicugna, its former genus name) is one of the two wild South American camelids, which live in the high alpine tundra, alpine areas of the Andes; the other cameli ...
. The guanaco was independently domesticated by the
Mapuche The Mapuche ( , ) also known as Araucanians are a group of Indigenous peoples of the Americas, Indigenous inhabitants of south-central Chile and southwestern Argentina, including parts of Patagonia. The collective term refers to a wide-ranging e ...
of Mocha Island in southern Chile, producing the
chilihueque The chilihueque/chiliweke or hueque/weke (''Lama araucana'') is an extinct hypothetical species of South America, South American Camelidae, camelid. It lived in Central Chile, central and Zona Sur, southern Chile until the Colonial Chile, coloni ...
, which was bred for its wool and to pull the
plough A plough or ( US) plow (both pronounced ) is a farm tool for loosening or turning the soil before sowing seed or planting. Ploughs were traditionally drawn by oxen and horses but modern ploughs are drawn by tractors. A plough may have a wooden ...
. This animal disappeared in the 17th century when it was replaced by
Old World The "Old World" () is a term for Afro-Eurasia coined by Europeans after 1493, when they became aware of the existence of the Americas. It is used to contrast the continents of Africa, Europe, and Asia in the Eastern Hemisphere, previously ...
sheep Sheep (: sheep) or domestic sheep (''Ovis aries'') are a domesticated, ruminant mammal typically kept as livestock. Although the term ''sheep'' can apply to other species in the genus '' Ovis'', in everyday usage it almost always refers to d ...
and draft animals.Westbury, M., Prost, S., Seelenfreund, A., Ramírez, J. M., Matisoo-Smith, E. A., & Knapp, M. (2016). First complete mitochondrial genome data from ancient South American camelids-the mystery of the chilihueques from Isla Mocha (Chile). ''Scientific reports'', 6(1), 1-7.


See also

* Llamanaco


References


Further reading

* *


External links


Puma takes on Guanaco 3 Times Her Weight
, Seven Worlds, One Planet , BBC Earth {{Authority control Mammals of the Andes Mammals of Patagonia Mammals of Chile Mammals of Argentina Mammals of Peru Mammals of Bolivia Camelids Quechua words and phrases Mammals described in 1776 Taxa named by Philipp Ludwig Statius Müller Animal hair products Wool animals