''Gryponyx'' (meaning "hooked-claw") is an

extinct Extinction is the termination of an organism by the death of its Endling, last member. A taxon may become Functional extinction, functionally extinct before the death of its last member if it loses the capacity to Reproduction, reproduce and ...

genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...

massopod Sauropodomorpha ( ; from Greek, meaning "lizard-footed forms") is an extinct clade of long-necked, herbivorous, saurischian dinosaurs that includes the sauropods and their ancestral relatives. Sauropods generally grew to very large sizes, had lon ...

sauropodomorph Sauropodomorpha ( ; from Greek, meaning "lizard-footed forms") is an extinct clade of long-necked, herbivorous, saurischian dinosaurs that includes the sauropods and their ancestral relatives. Sauropods generally grew to very large sizes, had lo ...

known from southern Free State, central

South Africa South Africa, officially the Republic of South Africa (RSA), is the Southern Africa, southernmost country in Africa. Its Provinces of South Africa, nine provinces are bounded to the south by of coastline that stretches along the Atlantic O ...

.Broom, R. (1911). On the dinosaurs of the Stormberg, South Africa. ''Annals of the South African Museum'' 7(4):291-308.

Description

''Gryponyx africanus'' is known from the

holotype A holotype (Latin: ''holotypus'') is a single physical example (or illustration) of an organism used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of s ...

SAM 3357-59, a nearly complete

postcrania The postcranium ("behind the cranium"; plural: postcrania) or postcranial skeleton in zoology and vertebrate paleontology is the skeleton apart from the skull. The postcranium encompasses the axial skeleton, which includes the entirety of the verte ...

skeleton A skeleton is the structural frame that supports the body of most animals. There are several types of skeletons, including the exoskeleton, which is a rigid outer shell that holds up an organism's shape; the endoskeleton, a rigid internal fra ...

which includes partial

vertebral column The spinal column, also known as the vertebral column, spine or backbone, is the core part of the axial skeleton in vertebrates. The vertebral column is the defining and eponymous characteristic of the vertebrate. The spinal column is a segmente ...

pelvis The pelvis (: pelves or pelvises) is the lower part of an Anatomy, anatomical Trunk (anatomy), trunk, between the human abdomen, abdomen and the thighs (sometimes also called pelvic region), together with its embedded skeleton (sometimes also c ...

, both

forelimb A forelimb or front limb is one of the paired articulated appendages ( limbs) attached on the cranial (anterior) end of a terrestrial tetrapod vertebrate's torso. With reference to quadrupeds, the term foreleg or front leg is often used inst ...

s and both

hindlimb A hindlimb or back limb is one of the paired articulated appendages ( limbs) attached on the caudal ( posterior) end of a terrestrial tetrapod vertebrate's torso.http://www.merriam-webster.com/medical/hind%20limb, Merriam Webster Dictionary-H ...

s. ''Gryponyx'' has been estimated to have been about in length. It was collected from the

Upper Elliot Formation The Elliot Formation is a geological formation and forms part of the Stormberg Group, the uppermost geological group that comprises the greater Karoo Supergroup. Outcrops of the Elliot Formation have been found in the northern Eastern Cape, south ...

of the

Stormberg Group The Stormberg Group is one of the four geological groups that comprises the Karoo Supergroup in South Africa. It is the uppermost geological group representing the final phase of preserved sedimentation of the Karoo Basin. The Stormberg Group r ...

(

Karoo Basin The Karoo Supergroup is the most widespread stratigraphic unit in Africa south of the Kalahari Desert. The supergroup consists of a sequence of units, mostly of nonmarine origin, deposited between the Late Carboniferous and Early Jurassic, a per ...

), dating to the

Hettangian The Hettangian is the earliest age and lowest stage of the Jurassic Period of the geologic timescale. It spans the time between 201.3 ± 0.2 Ma and 199.3 ± 0.3 Ma (million years ago). The Hettangian follows the Rhaetian (part of the Triass ...

Sinemurian In the geologic timescale, the Sinemurian is an age (geology), age and stage (stratigraphy), stage in the Early Jurassic, Early or Lower Jurassic epoch (geology), Epoch or series (stratigraphy), Series. It spans the time between 199.5 ±0.3 annu ...

stages of the

Lower Jurassic The Early Jurassic Epoch (in chronostratigraphy corresponding to the Lower Jurassic Series) is the earliest of three epochs of the Jurassic Period. The Early Jurassic starts immediately after the Triassic–Jurassic extinction event, 201.3 Ma&nb ...

period. It was originally described by Broom (1911) as a

theropod Theropoda (; from ancient Greek , (''therion'') "wild beast"; , (''pous, podos'') "foot"">wiktionary:ποδός"> (''pous, podos'') "foot" is one of the three major groups (clades) of dinosaurs, alongside Ornithischia and Sauropodom ...

. Huene (1932) named the family Gryponychidae to contain ''Gryponyx'' and ''Aetonyx'' and placed it within

Carnosauria Carnosauria is an extinct group of carnivorous theropod dinosaurs that lived during the Jurassic and Cretaceous periods. While Carnosauria was historically considered largely synonymous with Allosauroidea, some recent studies have revived Ca ...

.Friedrich von Huene (1932) Die fossile Reptil-Ordnung Saurischia, ihre Entwicklung und Geschichte. ''Monographien zur Geologie und Paläontologie, Series 1'' 4: 1–361

Galton Sir Francis Galton (; 16 February 1822 – 17 January 1911) was an English polymath and the originator of eugenics during the Victorian era; his ideas later became the basis of behavioural genetics. Galton produced over 340 papers and b ...

and Cluver synonymized ''G. africanus'' with '' Massospondylus harriesi'' in 1976,Galton, P.M., and Cluver, M.A. (1976). ''Anchisaurus capensis'' (Broom) and a revision of the Anchisauridae (Reptilia, Saurischia). ''Annals of the South African Museum'' 69(6):121-159. which was in turn synonymized by

Michael Cooper Michael Jerome Cooper (born April 15, 1956), nicknamed "Coop", is an American basketball coach and former player. He played for the Los Angeles Lakers during his entire career in the National Basketball Association (NBA), winning five NBA champ ...

in 1981 with ''

Massospondylus carinatus ''Massospondylus'' ( ) is a genus of sauropodomorph dinosaur from the Early Jurassic of southern Africa. It was described by Sir Richard Owen in 1854 from remains discovered in South Africa, and is thus one of the first dinosaurs to have been ...

''Cooper, M.R. (1981). The prosauropod dinosaur ''Massospondylus carinatus'' Owen from Zimbabwe: its biology, mode of life and phylogenetic significance. ''Occasional Papers of the National Museums and Monuments of Rhodesia, Series B, Natural Sciences'' 6(10):689-840. (and today ''M. harriesi'' is considered to be a ''

nomen dubium In binomial nomenclature, a ''nomen dubium'' (Latin for "doubtful name", plural ''nomina dubia'') is a scientific name that is of unknown or doubtful application. Zoology In case of a ''nomen dubium,'' it may be impossible to determine whether a ...

'').Galton, P.M., and Upchurch, P. (2004). Prosauropoda. In: D. B. Weishampel, P. Dodson, & H. Osmólska (eds.), ''The Dinosauria'' (second edition). University of California Press:Berkeley, 232-258. However, Vasconcelos and Yates (2004) found ''Gryponyx'' to be distinctive enough from other basal sauropodomorphs to be placed in its own genus. They found that it differs from other taxa by the following characteristics: total length of metacarpal I exceeds maximum proximal width and a long, narrow pubic apron with straight lateral margins. Although this publication wasn't formal, they conducted a

cladistic Cladistics ( ; from Ancient Greek 'branch') is an approach to biological classification in which organisms are categorized in groups ("clades") based on hypotheses of most recent common ancestry. The evidence for hypothesized relationships is ...

analysis using Yates (2004) sauropodomorph matrix and found ''Gryponyx'' to be the most basal massospondylid.C. C. Vasconcelos, A. M. Yates (2004). Sauropodomorph biodiversity of the upper Elliot Formation (Lower Jurassic) of southern Africa. ''Geoscience Africa, Abstract Volume'' 2:670. The same result was found by

Lü Junchang Lü Junchang (; 1965 – 9 October 2018) was a Chinese palaeontologist and professor at the Institute of Geology, Chinese Academy of Geological Sciences. An expert on Mesozoic reptiles, he described and named dozens of dinosaur and pterosaur taxa ...

''et al.'' (2010). Yates ''et al.'' (2010) recovered ''Gryponyx'' in a

trichotomy A trichotomy can refer to: * Law of trichotomy, a mathematical law that every real number is either positive, negative, or zero ** Trichotomy theorem, in finite group theory * Trichotomy (jazz trio), Australian jazz band, collaborators with Dan ...

with

Massospondylidae Massospondylidae is a family (biology), family of early massopod dinosaurs that existed in Asia, Africa, North America, South America and AntarcticaHellert, Spencer M. "A New Basal Sauropodomorph from The Early Jurassic Hanson Formation of Antarc ...

and

Anchisauria Sauropodiformes is an extinct clade of sauropodomorph dinosaurs that includes the animals more closely-related to Sauropoda than prosauropods like ''Massospondylus'' and '' Plateosaurus''. It was named in 2007 by Paul Sereno. Below is a cladogra ...

. However, ''Gryponyx'' has yet to be formally redescribed. Two additional species of ''Gryponyx'' have been described: ''G. transvaalensis'' was described on the basis of finger bones and the anterior limb metatarsals from the

Late Triassic The Late Triassic is the third and final epoch (geology), epoch of the Triassic geologic time scale, Period in the geologic time scale, spanning the time between annum, Ma and Ma (million years ago). It is preceded by the Middle Triassic Epoch a ...

Bushveld Sandstone Formation,

Transvaal Transvaal is a historical geographic term associated with land north of (''i.e.'', beyond) the Vaal River in South Africa. A number of states and administrative divisions have carried the name ''Transvaal''. * South African Republic (1856–1902; ...

. ''G. taylori'' was described on the basis of sacral and

pelvic The pelvis (: pelves or pelvises) is the lower part of an anatomical trunk, between the abdomen and the thighs (sometimes also called pelvic region), together with its embedded skeleton (sometimes also called bony pelvis or pelvic skeleton). ...

rim from the Upper Elliot Formation, southern Free State. Galton and Cluver (1976) synonymized ''G. taylori'' with ''M. harriesi'' and considered ''G. transvaalensis'' to be a ''nomen dubium''. Both ''G. taylori'' and ''G. transvaalensis'' were synonymized by Michael Cooper (1981) with ''M. carinatus'' and Galton and Upchurch (2004) considered them to be dubious.

Etymology

''Gryponyx'' was first named by

Robert Broom Robert Broom Fellow of the Royal Society, FRS FRSE (30 November 1866 6 April 1951) was a British- South African medical doctor and palaeontologist. He qualified as a medical practitioner in 1895 and received his DSc in 1905 from the University ...

1911 Events January * January 1 – A decade after federation, the Northern Territory and the Australian Capital Territory are added to the Commonwealth of Australia. * January 3 ** 1911 Kebin earthquake: An earthquake of 7.7 m ...

and the

type species In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...

is ''Gryponyx africanus''. The generic name is derived from ''grypos'',

Greek Greek may refer to: Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group *Greek language, a branch of the Indo-European language family **Proto-Greek language, the assumed last common ancestor of all kno ...

for "hooked" and ''onyx'', Greek for "claw". The

specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...

refers to

Africa Africa is the world's second-largest and second-most populous continent after Asia. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 20% of Earth's land area and 6% of its total surfac ...

, where the holotype was discovered.

References

{{Taxonbar, from=Q1989760 Massopoda Dinosaur genera Hettangian dinosaurs Sinemurian dinosaurs Elliot Formation Fossil taxa described in 1911 Taxa named by Robert Broom Dinosaurs of South Africa